Search USGS⌕ Search

SEARCH · Search USGS

Results for “Raven”

Search indexed USGS publications on groundwater, aquifers, geologic maps, mineral resources and earthquakes. Explore source records by subject and place.

Quote a phrase for an exact phrase match. Source license links do not imply unrestricted reuse.

At least 19 recordsLinked to original sources
USGS2002

Reducing predation by common ravens on desert tortoises in the Mojave and Colorado Deserts

Conflicts between humans and natural populations often result from habitat fragmentation and degradation that accompanies human activities. Common raven populations in the Mojave Desert have benefited by human-provided resources; they've expanded precipitously in recent years. Because ravens prey on juveniles of the threatened desert tortoise, they have become the focus of management concerns to help recover dwindling tortoise populations. I have outlined herein a series of management recommendations designed to reduce raven predation on desert tortoises thereby facilitating juvenile tortoise recruitment into the population of reproductive adults. The recommendations are based on the best available scientific information and are intended to provide a basis for a long-term reduction in raven impacts. The recommendations fall into four basic categories. (1) Modify anthropogenic sources of food, water, and nesting substrates to reduce their use by ravens. This includes modifying landfill operations, septage containment practices, livestock management, and other commercial and private practices that help facilitate raven survival and dispersal by providing food and water. Most of these measures are long-term actions deigned to reduce the carrying capacity of the desert for ravens. This action is critical and must be done over very large areas. (2)Lethal removal of ravens by shooting or euthanizing following live trapping. Specific ravens known to prey on tortoises would be targeted as well as all ravens found foraging within specific high-priority desert tortoise management zones (e.g., Desert Tortoise Natural Areas, DTNA). These actions would primarily be deployed on a short-term emergency basis to give specific tortoise populations a necessary boost until other measures become fully implemented and achieve their goals. (3) Conduct research on raven ecology, raven behavior, and methods to reduce raven predation on tortoises. Results of these studies would be used to design future phases of the raven management program. (4) All actions should be approached within an adaptive management framework. As such monitor, actions should be designed as experiments so that monitoring of actions will yield reliable and scientifically sound results. Coordinating and oversight teams should be convened to facilitate cooperation and coordination among agencies and to ensure that the actions are being implemented effectively. Recommendations made herein were developed to help recover tortoise populations by reducing raven predation on juvenile tortoises. If the recommendations made are implemented in concert with actions reducing other causes of mortality, ill health, and lowered reproductive output, they should aid in the long-term recovery of desert tortoise populations. Many important aspects of raven population dynamics, raven predation on tortoises, and how to manage raven populations and behavior are as yet unknown. Because of this, any raven management program must be implemented within an adaptive management framework. Doing so would allow for sufficient flexibility to modify the program as new information is gained.

Report↗
USGS2022

Influence of anthropogenic subsidies on movements of common ravens

Anthropogenic subsidies can benefit populations of generalist predators such as common ravens (ravens; Corvus corax ), which in turn may depress populations of many types of species at lower-trophic levels, including desert tortoises (Gopherus agassizii ) or greater sage-grouse (Centrocercus urophasianus). Management of subsidized ravens often has targeted local breeding populations that are presumed to affect species of concern and ignored “urban” populations of ravens. However, little is known about how ravens move, especially in response to the presence of anthropogenic subsidies. Therefore, subsidized ravens from distant populations that are not managed may influence local prey. To better understand this issue, we deployed global positioning system – global system for mobile communications transmitters to track movements of 19 ravens from September to December 2020 relative to 2 land cover types that provide subsidies: developed areas and cultivated crops. On average, ravens moved 41.5 km (±30.5) per day, although daily movement distances ranged from 0.13– 206.1 km. Raven movement among cover types during the non-breeding season varied widely, with 100% of individuals each using land cover types that provide subsidy and other types at least once in the season. On 100% of days ravens used areas that did not provide subsidy, on 86.7% of days they used developed areas, and on 20.5% of days they used cultivated crops. Although on some days a raven would stay exclusively in areas that did not provide subsidy, there were no days in which a single raven ever stayed exclusively in developed or cultivated crops. Ravens moved shorter distances on days when they used subsidies more frequently. Further, time spent in developed areas and cultivated crops increased when ravens roosted closer to them, although this effect was greater for developed areas than for cultivated crops. Individual ravens were not associated exclusively with either of the subsidy-providing landscapes we considered, but instead all birds used both subsidized and other landscapes. Our research suggests that management of ravens during the non-breeding season and possibly during the breeding season, intended to reduce risk of predation on desert tortoises, will be most effective if conducted on a broad scale because of distances the birds travel and the lack of separation between putative “urban” and “natural” populations of ravens.

California↗
USGS2021

A desert tortoise-common raven viable conflict threshold

Since 1966, common raven ( Corvus corax ; raven) abundance has increased throughout much of this species’ Holarctic distribution, fueled by an ever-expanding supply of anthropogenic resource subsidies (e.g., water, food, shelter, and nesting substrate) to ecoregion specific raven population carrying capacities. Consequently, ravens are implicated in declines of both avian and reptilian species of conservation concern, including the California (USA) endangered and federally threatened Mojave desert tortoise ( Gopherus agassizii ; desert tortoise). While ravens are a natural predator of desert tortoises, the inter-generational stability of desert tortoise populations is expected to be compromised as annual juvenile survival is suppressed below 0.77 through a combination of raven depredation and other sources of mortality. To estimate the extent to which raven depredation suppresses desert tortoise recruitment within the Mojave Desert of California, we collected data from 274 variable-radius point counts, 78 desert tortoise decoy stations, and 8 control stations during the spring of 2020. Additionally, we complied a geodatabase of previously active raven nests, observed between 2013 and 2020. Raven density estimates from 4 monitoring areas ranged between 0.63 (eastern most) and 2.44 (western most) raven km -2 (95% CI: 0.35–1.14 and 1.33–4.48, respectively). We used a Bayesian shared frailty model to estimate the effects of raven density and distance to the nearest previously active raven nest on the annual “survival” of juvenile desert tortoise decoys (75-mm Midline Carapace Length), which we then converted into survival estimates for 0- to 10-year-old desert tortoises by adjusting exposure to reflect natural activity patterns. At the 1.72-km median distance from the nearest previously active raven nest, the estimated annual survival of desert tortoises decreased as raven density increased, ranging among conservation areas from 0.774 (eastern most) to 0.733 (western most). Accordingly, our model predicts that desert tortoise populations exposed to raven densities in excess of 0.89 raven km -2 , at a distance

California↗
USGS2025

The effects of breeding status on common raven movement, home range, and habitat selection

Anthropogenic infrastructure has contributed to increasing common raven ( Corvus corax ) abundance across the Great Basin region of the United States, particularly in sagebrush ecosystems, where high raven densities are correlated with reduced sage-grouse ( Centrocercus urophasianus ) nest survival. Our understanding of how raven reproductive behavior affects sage-grouse nest predation is limited, especially considering their overlapping breeding seasons. Understanding differences in space use and resource selection between breeding and non-breeding ravens could help identify high-use areas and corresponding predation risk for sage-grouse nests. We analyzed space use and resource selection of breeding ( n = 13) and non-breeding ( n = 32) global positioning system (GPS)-marked ravens in Nevada, USA (2017–2022) during the breeding season (1 March–31 June). We compared home-range size, core area size, step lengths, and resource selection within a Bayesian framework with inference made by comparing Bayesian credible intervals (CRI). We generated home range and core area estimates using autocorrelated kernel density methods. We did not find a difference in home range size between breeding (469.33 km 2 , 95% CRI = 228.79–709.45 km 2 ) and non-breeding (525.26 km 2 , 95% CRI = 410.71–654.10 km 2 ) ravens. However, breeding ravens had smaller core areas (10.77 km 2 , 95% CRI = 3.16–35.78 km 2 ) and shorter step lengths (1,160.33 m/hr, 95% CRI = 1,087.78–1,277.17 m/hr) than non-breeding ravens (core area = 279.50 km 2 , 95% CRI = 206.77–363.72 km 2 ; step length = 1,953.74 m/hr, 95% CRI = 1,898.42–2,009.56 m/hr). Ravens in both breeding classes selected high normalized difference vegetation index (NDVI) and low annual grass and shrub cover, but non-breeding ravens showed stronger selection for low annual grass and shrub cover areas. We found strong differences in selection between breeding classes for 6 of our 9 covariates: distance to road, solar radiation, distance to natural water, distance to forest edge, percent annual grass cover, and percent shrub cover. Non-breeding ravens concentrated activity near forest edges, natural water sources, and anthropogenic features, whereas breeding ravens focused activity close to their nests. Our findings suggest that raven management could be more effective if it targeted areas with high NDVI and low annual grass and shrub cover, especially in anthropogenically modified landscapes and near forest edges, and prevented raven nest establishment near prey populations of concern.

Nevada↗
USGS2025

Evaluation of a rapid assessment function to aid monitoring and management of common ravens (Corvus corax) in Washington state

Expanding human enterprise leading to resource subsidies for generalist species has resulted in widespread increases in common raven ( Corvus corax ) populations across the Western U.S. Ravens are an efficient predator and increased population abundance has led to adverse effects to multiple sensitive prey species. In regions where problematic interactions between ravens and their prey exist, managers seek efficient and effective tools for monitoring and controlling expanding raven populations. We previously developed a Rapid Assessment Function (RAF) for managers to quickly estimate raven population density and assess the need for management actions. We developed the RAF for the Great Basin (GB RAF) by first estimating raven density using robust distance sampling protocols with >30,000 raven point count surveys from sagebrush ecosystems in California, Nevada, Idaho, and Oregon across 131 field sites and years. We then used the relationship between raven density estimates from distance sampling and n ravens observed site-year / n surveys site-year (that is, raven index) at each site-year combination to develop a function that accounts for detection probability and adjusts simple counts to provide a prediction of ‘true’ density. Our function produced reliable density estimates given approximately 50–100 surveys, thereby reducing the field-based and analytical efforts typically needed to estimate raven density, facilitating more efficient raven management in open sagebrush habitats. In this study, we sought to test our original GB RAF using data from sagebrush ecosystems outside of the Great Basin. Using raven point count data from two field site units in Washington state collected from 2016 to 2023, we calculated density estimates from distance sampling methods, comparable to what was done for previous analyses. We then used the GB RAF to generate predictions of density and compared those values to the more robust estimates from distance sampling. Additionally, we developed modified RAFs specifically for Washington data (WA RAFs) to assess how well they predicted raven density compared to the GB RAF. We found the detection curves estimated for the Washington sites largely aligned with those used to generate the original GB RAF. Furthermore, the estimates from the GB RAF exhibited similar or higher correlation with densities calculated from distance models ( Pearson’s r = 0.73) than the modified WA RAFs with 1.33 km and 1.25 km truncation distances ( Pearson’s r = 0.63 and 0.73, respectively). Producing an equivalently performing modified WA RAF would likely necessitate more data to reduce estimation error and produce more reliable estimates. These results provide evidence for the applicability of our GB RAF for more widespread use within sagebrush ecosystems, possibly negating the need for locally developed RAFs. Continued assessments of the GB RAF outside of the Great Basin would further verify its applicability across the sagebrush biome.

BioRxiv↗
USGS2021

Efficacy of manipulating reproduction of common ravens to conserve sensitive prey species: Three case studies

Expansion of human enterprise across western North America has resulted in an increase in availability of anthropogenic resource subsidies for generalist species. This has led to increases in generalists’ population numbers across landscapes that were previously less suitable for their current demographic rates. Of particular concern are growing populations of common ravens ( Corvus corax ; ravens), because predation by ravens is linked to population declines of sensitive species. Ecosystem managers seek management options for mitigating the adverse effects of raven predation where unsustainable predator–prey conflicts exist. We present 3 case studies examining how manipulating reproductive success of ravens influences demographic rates of 2 sensitive prey species. Two case studies examine impacts of removing raven nests or oiling raven eggs on nest survival of greater sage-grouse ( Centrocercus urophasianus ; sage-grouse) within Wyoming and the Great Basin of California and Nevada, USA, respectively. The third case study uses Mojave desert tortoise ( Gopherus agassizii ; tortoise) decoys to examine effects of oiling raven eggs on depredation rates of juvenile tortoises in the Mojave Desert in California. Initial trial years from all 3 case studies were consistent in finding improved vital rates associated with the application of strategies for reducing reproductive success of ravens. Specifically, removal of raven nests resulted in increased nest survival of sage-grouse within treatment areas where predation by ravens was the primary cause of nest failure. In addition, nest survival of sage-grouse and survival of juvenile tortoise decoys was higher following a treatment of oiling the eggs of ravens in their nests at 2 sites within the Great Basin and 4 tortoise conservation areas in the Mojave Desert in California. Along with specialized technologies that can make techniques such as egg-oiling more feasible, these findings support these management practices as important tools for managing ravens, especially in areas where breeding ravens have negative impacts on sensitive prey species.

California, Nevada, Wyoming↗
USGS2021

Common ravens disrupt greater sage-grouse lekking behavior in the Great Basin, USA

Expansion of human enterprise has contributed to increased abundance and distribution of common ravens ( Corvus corax ; ravens) across sagebrush ( Artemisia spp.) ecosystems within western North America. Ravens are highly effective nest predators of greater sage-grouse ( Centrocercus urophasianus ; sage-grouse), a species of high conservation concern. Sage-grouse population trends are estimated using count survey data of males attending traditional breeding grounds, known as leks. We sought to investigate associations of ravens to sage-grouse lek sites and document interactions between the sage-grouse and ravens as well as those between sage-grouse and other animals observed around leks. First, we used extensive raven point counts and sage-grouse lek observation data collected across Nevada and California, USA, from 2009–2019 to evaluate spatial associations between sage-grouse and ravens while accounting for other environmental covariates. We found that ravens were more likely to be observed closer to lek sites, especially as leks increased in size. Second, we used a subset of the lek dataset from 2006–2019 to describe behavioral changes of male sage-grouse in the presence of ravens and other predators. Our analysis indicated that ravens are attracted to lek sites and were associated with disrupting lekking sage-grouse by causing flushes or ceasing displaying behaviors. These results suggest that adult and yearling sage-grouse perceive ravens as a reason to alter breeding activity, and ravens may adversely influence their reproduction during the lekking stage. Additionally, standardized techniques to count sage-grouse on leks for population trend analyses could be biased low if raven presence during surveys is not accounted for.

California, Nevada↗
USGS2021

Estimating trends of common raven populations in North America, 1966—2018

Over the last half century, common raven ( Corvus corax; raven) populations have increased in abundance across much of North America. Ravens are generalist predators known to depredate the eggs and young of several sensitive species. Quantifying raven population increases at multiple spatial scales across North America will help wildlife resource managers identify areas where population increases present the greatest risk to species conservation. We used a hierarchical Bayesian modeling approach to analyze trends of standardized raven counts from 1966 to 2018 using Breeding Bird Survey data within each Level I and II ecoregion of the United States and Canada. We also compared raven abundance within and outside the distributions of 9 sensitive or endangered species. Although we found substantial evidence that raven populations have increased across North America, populations varied in growth rates and relative abundances among regions. We found 73% of Level I (11/15) and II (25/34) ecoregions demonstrated positive annual population growth rates ranging from 0.2–9.4%. We found higher raven abundance inside versus outside the distributions of 7 of the 9 sensitive species included in our analysis. Gunnison sage-grouse ( Centrocercus minimus ) had the highest discrepancy, with 293% more ravens within compared to outside of their range, followed by greater sandhill crane ( Antigone canadensis tabida ; 280%), and greater sage-grouse ( C. urophasianus ; 204%). Only 2 species, least tern ( Sternula antillarum ) and piping plover ( Charadrius melodus ), indicated lower raven abundance within relative to outside their distributions. Our findings will help wildlife resource managers identify regional trends in abundance of ravens and anticipate which sensitive species are at greatest risk from elevated raven populations. Future research directed at identifying the underlying regional drivers of these trends could help elucidate the most appropriate and responsive management actions and, thereby, guide the development of raven population management plans to mitigate impacts to sensitive species.

Human-Wildlife Interactions↗
USGS2021

A rapid assessment function to estimate common raven population densities: Implications for targeted management

Common raven ( Corvus corax ; raven) populations have increased over the past 5 decades within the western United States. Raven population increases have been largely attributed to growing resource subsidies from expansion of human enterprise. Concomitantly, managers are becoming increasingly concerned about elevated adverse effects on multiple sensitive prey species, damage to livestock and agriculture, and human safety. Managers could benefit from a rapid but reliable method to estimate raven densities across spatiotemporal scales to monitor raven populations more efficiently and inform targeted and adaptive management frameworks. However, obtaining estimates of raven density is data- and resource-intensive, which renders monitoring within an adaptive framework unrealistic. To address this need, we developed a rapid survey protocol for resource managers to estimate site-level density based on the average number of ravens per survey. Specifically, we first estimated raven densities at numerous field sites with robust distance sampling procedures and then used regression to investigate the relationship between those density estimates and the number of ravens per survey, which revealed a strong correlation ( R 2 = 0.86). For management application, we provide access to R function software through a web-based interface to estimate density using number of ravens per survey, which we refer to as a Rapid Assessment Function (RAF). Then, using a simulation analysis of data from sites with abundant surveys and the RAF, we estimated raven density based on different numbers of surveys to help inform how many surveys are needed to achieve reliable estimates within this rapid assessment. While more robust procedures of distance sampling are the preferred methods for estimating raven densities from count surveys, the RAF tool presented herein provides a reliable approximation for informing management decisions when managers are faced with resource and small sample size constraints.

California, Idaho, Nevada, Oregon↗
USGS2016

Landscape characteristics and livestock presence influence common ravens: Relevance to greater sage-grouse conservation

Common raven ( Corvus corax ; hereafter, raven) population abundance in the sagebrush steppe of the American West has increased threefold during the previous four decades, largely as a result of unintended resource subsidies from human land-use practices. This is concerning because ravens frequently depredate nests of species of conservation concern, such as greater sage-grouse ( Centrocercus urophasianus ; hereafter, sage-grouse). Grazing by livestock in sagebrush ecosystems is common practice on most public lands, but associations between livestock and ravens are poorly understood. The primary objective of this study was to identify the effects of livestock on raven occurrence while accounting for landscape characteristics within human-altered sagebrush steppe habitat, particularly in areas occupied by breeding sage-grouse. Using data from southeastern Idaho collected during spring and summer across 3 yr, we modeled raven occurrence as a function of the presence of livestock while accounting for multiple landscape covariates, including land cover features, topographical features, and proximity to sage-grouse lek sites (breeding grounds), as well as site-level anthropogenic features. While accounting for landscape characteristics, we found that the odds of raven occurrence increased 45.8% in areas where livestock were present. In addition, ravens selected areas near sage-grouse leks, with the odds of occurrence decreasing 8.9% for every 1-km distance, increase away from the lek. We did not find an association between livestock use and distance to lek. We also found that ravens selected sites with relatively lower elevation containing increased amounts of cropland, wet meadow, and urbanization. Limiting raven access to key anthropogenic subsidies and spatially segregating livestock from sage-grouse breeding areas would likely reduce exposure of predatory ravens to sage-grouse nests and chicks.

Idaho↗
USGS2004

Diet composition of common ravens across the urban-wildland interface of the West Mojave Desert

Common ravens ( Corvus corax ) are human-subsidized scavengers and predators in the Mojave Desert. They have increased dramatically in number and have been implicated as contributors to the decline in desert tortoise ( Gopherus agassizii ) populations. Known patterns of increased fledging success near human developments suggested that food was the most likely resource subsidy received by ravens. Because ravens are opportunistic foragers with a generalist diet, we predicted that the types of resource subsidy provided by different kinds of human developments should be reflected in measures of diet composition of breeding ravens. We estimated diet composition from contents of raven pellets collected at nests and related diet composition to distance of the nests from roads and point sources of resource subsidies, such as towns or landfills. Ravens that nested close to point subsidies far from major roads had the greatest incidence of trash in their diets. Ravens that nested close to roads but far from point subsidies had a low incidence of trash and a higher incidence of presumably road-killed mammals and reptiles. Ravens far from both roads and point subsidies had more plant material and arthropods, and ravens close to both roads and point subsidies had more birds and amphibians. Diet diversity was not related to distance from roads or developments. Fledging success was correlated with diet composition, such that birds with diets consistent with trash or road-kill subsidies fledged the greatest number of chicks. Our results suggest that ravens forage opportunistically on foods available near their nests, and different kinds of human developments contribute different foods. Improved management of landfills and highway fencing to reduce road-kills may help slow the growth of raven populations in the Mojave.

Wildlife Society Bulletin↗
USGS2014

Common raven occurrence in relation to energy transmission line corridors transiting human-altered sagebrush steppe

Energy-related infrastructure and other human enterprises within sagebrush steppe of the American West often results in changes that promote common raven ( Corvus corax ; hereafter, raven) populations. Ravens, a generalist predator capable of behavioral innovation, present a threat to many species of conservation concern. We evaluate the effects of detailed features of an altered landscape on the probability of raven occurrence using extensive raven survey ( n = 1045) and mapping data from southern Idaho, USA. We found nonlinear relationships between raven occurrence and distances to transmission lines, roads, and facilities. Most importantly, raven occurrence was greater with presence of transmission lines up to 2.2 km from the corridor.We further explain variation in raven occurrence along anthropogenic features based on the amount of non-native vegetation and cover type edge, such that ravens select fragmented sagebrush stands with patchy, exotic vegetative introgression. Raven occurrence also increased with greater length of edge formed by the contact of big sagebrush ( Artemisia tridentate spp.) with non-native vegetation cover types. In consideration of increasing alteration of sagebrush steppe, these findings will be useful for planning energy transmission corridor placement and other management activities where conservation of sagebrush obligate species is a priority.

Idaho↗
USGS2021

Synthesis of nest predation impacts of common ravens on sensitive avian species

Decades of mounting scientific evidence have revealed that common raven ( Corvus corax ; raven) population numbers have been increasing across nearly all regions of their geographic range in North America. Concomitantly, numerous native wildlife species have experienced elevated predation rates from ravens as populations have increased and expanded their range. Managers are concerned that increased raven predation of many threatened and endangered avian species in the U.S. and Canada during nesting periods may be hampering species recovery. We explored the literature to aggregate existing knowledge and evaluate the impacts of raven predation on nests and young of sensitive avian species. We used this information to develop a simple relative index for each species, the “Raven Impact Index” (RII). The RII incorporated the species demographic rates, abundance of ravens in relation to each sensitive species’ breeding range, and the degree of overlap between raven and sensitive prey distributions. We also developed a second relative descriptor describing our confidence in each RII, termed a “Impact Credibility Index (ICI).” The species ICI was based on the number of published studies and the type of evidence presented (e.g., circumstantial vs. direct). We found evidence of nest predation on 8 sensitive avian species and suspected nest predation on 1 additional species. All species shared aspects of nesting biology that suggested they would likely be susceptible to raven nest predation. The RII varied among prey species, with greater sage-grouse ( Centrocercus urophasianus ) having the highest relative impact values, followed by snowy plover ( Charadrius nivosus nivosus ), marbled murrelet ( Brachyramphus marmoratus ), and Gunnison sage-grouse ( Centrocercus minimus ). Our species RII is intended to inform management decisions regarding actions that mitigate the negative effects of raven predation of sensitive avian species. Although elevated nest predation may be of high conservation concern, it is important to recognize that all of the sensitive native prey species we established an RII for also face multiple conservation threats.

Human–Wildlife Interactions↗
USGS2021

SMaRT: A science-based tiered framework for common ravens

Large-scale increases and expansion of common raven ( Corvus corax ; raven) populations are occurring across much of North America, leading to increased negative consequences for livestock and agriculture, human health and safety, and sensitive species conservation. We describe a science-based adaptive management framework that incorporates recent quantitative analyses and mapping products for addressing areas with elevated raven numbers and minimizing potential adverse impacts to sensitive species, agricultural damage, and human safety. The framework comprises 5 steps: (1) desktop analysis; (2) field assessments; (3) comparison of raven density estimates to an ecological threshold (in terms of either density or density plus distance to nearest active or previous nest); (4) prescribing management options using a 3-tiered process (i.e., habitat improvements, subsidy reductions, and direct actions using StallPOPd.V4 software); and (5) post-management monitoring. The framework is integrated within the Science-based Management of Ravens Tool (SMaRT), a web-based application outfitted with a user-friendly interface that guides managers through each step to develop a fully customized adaptive plan for raven management. In the SMaRT interface, users can: (1) interact with pre-loaded maps of raven occurrence and density and define their own areas of interest within the Great Basin to delineate proposed survey or treatment sites; (2) enter site-level density estimates from distance sampling methods or perform estimation of raven densities using the rapid assessment protocol that we provide; (3) compare site-level density estimates to an identified ecological threshold; and (4) produce a list of potential management options for their consideration. The SMaRT supports decision-making by operationalizing scientific products for raven management and facilitates realization of diverse management goals including sensitive species conservation, protection of livestock and agriculture, safeguarding human health, and addressing raven overabundance and expansion. We illustrate the use of the framework through SMaRT using an example of greater sage-grouse ( Centrocercus urophasianus ) conservation efforts within the Great Basin, USA.

California, Idaho, Nevada, Oregon, Utah↗
USGS2021

Spatial modeling of common raven density and occurrence helps guide landscape management within Great Basin sagebrush ecosystems

Common ravens ( Corvus corax ; ravens) are a behaviorally flexible nest predator of several avian species, including species of conservation concern. Movement patterns based on life history phases, particularly territoriality of breeding birds and transiency of nonbreeding birds, are thought to influence the frequency and efficacy of nest predation. As such, predicting where on the landscape territorial resident and non-territorial transient birds may be found in relation to the distribution of sensitive prey is of increasing importance to managers and conservationists. From 2007 to 2019, we conducted raven point count surveys between mid-March and mid-September across 43 different field sites representing typical sagebrush ( Artemisia spp.) ecosystems of the Great Basin, USA. The surveys conducted during 2007–2016 were used in previously published maps of raven occurrence and density. Here, we examined the relationship between occurrence and density of ravens using spatially explicit predictions from 2 previously published studies and differentiate areas occupied by higher concentrations of resident ravens as opposed to transients. Surveys conducted during 2017–2019 were subsequently used to evaluate the predicted trends from our analytical approach. Specifically, we used residuals from a generalized linear regression to establish the relationship between occurrence and density, which ultimately resulted in a spatially explicit categorical map that identifies areas of resident versus transient ravens. We evaluated mapped categories using independently collected observed raven group sizes from the 2017–2019 survey data, as well as an independent dataset of global positioning system locations of resident and transient individuals monitored during 2019–2020. We observed moderate agreement between the mapped categories and independent datasets for both evaluation approaches. Our map provides broad inference about spatial variation in potential predation risk from ravens for species such as greater sage-grouse ( Centrocercus urophasianus ) and can be used as a valuable spatial layer for decision support tools aimed at guiding raven management decisions and, ultimately, improving survival and reproduction of sensitive prey within the Great Basin.

Arizona, California, Idaho, Montana, Nevada, Orego↗
USGS2000

Cryptic genetic variation and paraphyly in ravens

Widespread species that are morphologically uniform may be likely to harbour cryptic genetic variation. Common ravens ( Corvus corax ) have an extensive range covering nearly the entire Northern Hemisphere, but show little discrete phenotypic variation. We obtained tissue samples from throughout much of this range and collected mitochondrial sequence and nuclear microsatellite data. Our study revealed a deep genetic break between ravens from the western United States and ravens from throughout the rest of the world. These two groups, the ‘California clade’ and the ‘Holarctic clade’ are well supported and over 4% divergent in mitochondrial coding sequence. Microsatellites also reveal significant differentiation between these two groups. Ravens from Minnesota, Maine and Alaska are more similar to ravens from Asia and Europe than they are to ravens from California. The two clades come in contact over a huge area of the western United States, with mixtures of the two mitochondrial groups present in Washington, Idaho and California. In addition, the restricted range Chihuahuan raven ( Corvus cryptoleucus ) of the south–west United States and Mexico is genetically nested within the paraphyletic common raven. Our findings suggest that the common raven may have formerly consisted of two allopatric groups that may be in the process of remerging.

Proceedings of the Royal Society B: Biological Sci↗
USGS2021

Occurrence, resource use, and demography of the common raven in North America: A research synthesis

We reviewed the scientific literature to inventory existing studies of common raven ( Corvus corax ; raven) ecology in western North America. We conducted an intial literature review between June 2015 and March 2018. Prior to completing our review, we revisited the published literature for any additional relevant studies in July 2021. Our goal was to identify knowledge gaps and to synthesize the current understanding of environmental features that may support raven populations that pose general threats to biodiversity and sensitive species in particular. We focused our review on studies with direct conservation applications related to 3 processes of raven ecology: occurrence, resource use, and demography. We identified covariates that researchers associated with these processes of raven ecology, and we also quantified the geographic distribution of studies. Our review identified 54 studies, with an increasing number of studies published per decade and a geographic bias characterized by more studies conducted in the Mojave and Columbia Plateau ecoregions than elsewhere. Most studies (44) reported on a single ecological process, but 10 studies reported on multiple ecological processes. Results related to raven occurrence appeared 31 times; demographic results appeared 21 times; and resource use was reported 17 times. We also identified 13 explanatory covariates regularly invoked to explain variation in raven ecological processes. Greater attention was given to covariates including vegetation land cover, human settlement, recreation, and linear rights-of-ways than were used to explain variation in ecological processes. Most demographic studies investigated raven reproduction exclusively, but a small number of studies considered raven survival exclusively or in combination with reproduction. Along with a detailed summary of individual studies provided as an appendix, we intend for our findings to serve as a reference and to help identify future research priorities.

Human–Wildlife Interactions↗
USGS2021

Inter- and intra-annual effects of lethal removal on common raven abundance in Nevada and California, USA

Populations of common ravens ( Corvus corax ; ravens) have increased rapidly within sagebrush ( Artemisia spp.) ecosystems between 1960 and 2020. Although ravens are native to North America, their population densities have expanded to levels that negatively influence the population dynamics of other wildlife species of conservation concern, such as greater sage-grouse ( Centrocercus urophasianus ) and desert tortoises ( Gopherus agassizii ). For this reason, lethal removal, such as the application of the avicide DRC-1339, has been used to manage raven numbers at local scales and under certain circumstances. Because the relative effectiveness of DRC-1339 in reducing raven populations densities is not thoroughly understood, we completed 2 case studies using a before-after-control-impact experimental design of density estimates generated from point count data within a Bayesian hierarchical distance sampling framework. Specifically, we analyzed >16,000 point count surveys collected during 2009–2019 and split into 2 study designs covering multiple field sites within the Great Basin region. The first experiment evaluated intra-annual changes in density by comparing before and after treatment time periods within a single breeding season for multiple treatment regions compared to 2 control regions. The other experiment focused on inter-annual differences by comparing time periods across years before and after the onset of annual avicide application for a single treatment region compared to multiple control regions. Our models estimated a 100% probability of decline in density relative to control sites for both the intra- and inter-annual model designs. At treatment sites, expected densities of ravens varied but were reduced by 43% (95% CRI: 33–49%) and 54% (95% CRI: 24–71%) according to intra- and inter-annual analyses, respectively, whereas densities increased by 42% (95% CRI: 27–60%) and 15% (95% CRI: -17 to 58%) at control sites. Although population densities were reduced with treatments, trends indicated that sustained effort would likely be needed to maintain densities at acceptable levels within regions of interest. Effectively reducing the adverse effects of raven populations on other native species likely will depend on a variety of targeted management actions such as improving habitat quality for prey species, possibly reducing ravens’ population density, and treating the cause of increased raven abundance to reduce future carrying capacity and prevent rebounds.

California, Nevada↗