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Development of an aerial population survey method for elk (Cervus elaphus) in Rocky Mountain National Park, Colorado

Executive Summary Since the early 1990s, substantial effort and funding have been expended to conduct research to guide development of a 20-year Elk and Vegetation Management Plan for Rocky Mountain National Park (RMNP) in Colorado. One goal of the plan is to maintain the elk ( Cervus elaphus ) population size at the lower end of the natural range of variation. To implement management actions called for in the plan, accurate and reliable population estimates are needed, as well as a better understanding of the spatial and temporal distribution of elk. The previous aerial survey protocol and population estimation model used by the park had not been updated since the model’s initial calibration more than 15 years ago and the model was developed with an insufficient number (n=44) of observations. Thus we initiated research to reevaluate, update, and improve elk population estimation protocols for RMNP. We considered several alternative survey and analysis methods, and concluded that a hybrid population-estimation model using a simultaneous double-observer technique with sighting covariates was the most appropriate and effective aerial survey methodology for this population and the environmental conditions in RMNP. Instructional protocols for conducting these surveys in the future, along with datasheets, are provided in this report’s appendixes. To develop an improved method for aerial elk surveys, we used elk radio-collar location data from our study and other studies, and applied geographic information system analyses to define the survey area and develop effective, repeatable survey transect lines. We used telemetry data from radio-collared elk to inform our understanding of the temporal and spatial scale of elk movements across the park boundary, elk use of tree cover during potential survey hours, and elk use of different elevations within their range. Determining where elk were during surveys helped to fine-tune a survey design that improved spatial cover-age and decreased costs where possible, while standardizing the method to make it repeatable from year to year. We conducted aerial helicopter surveys to test our methodology in an adaptive, iterative process during three winters: 2007–2008, 2008–2009, and 2009–2010. We gained new information on each survey and used results to refine subsequent surveys. Our results confirm that elk movements were highly dynamic with respect to park boundary crossings; an average of six round trips from the park to Estes Park, Colorado, and back per month were taken by global positioning system-collared bull elk. We observed a strong diurnal temporal pattern of bull elk use of trees; elk were found in dense tree cover from 15:00−22:00 but not as often during morning and early afternoon hours when surveys were conducted. We used information on elk use of different altitudes to refine and establish a more efficient survey area. During the time of our study, a concurrent study in the park deployed 120 very high frequency radio collars on elk cows. We used those collar locations during our flights to increase sample size and to evaluate the level of precision we would gain by using “known fates analysis” (in which elk were known to be in the survey area, out of the survey area, or deceased based on radio-collar locations collected simultaneously during aerial surveys). Using radio-collar locations reduced bias by 1.1–8.8 percent and increased precision (that is, reduced the width of confidence intervals). This report provides final population estimates analyzed with and without radio-collar data to demonstrate what is gained by using marked individuals.

Colorado

Water resources of part of Canyonlands National Park, southeastern Utah

Canyonlands National Park is in about the center of the Canyon Lands section of the Colorado Plateaus physiographic province in southeastern Utah. The part of the park discussed embraces an area of about 400 square miles comprising isolated mesas, precipitous canyons, and dissected broad benches near the confluence of the Green and Colorado Rivers, the only perennial streams in the area. The climate is arid to semiarid; normal annual precipitation ranges from less than 8 to about 10 inches. Potential evapotranspiration is about 41 inches annually. Geology of the park is characterized by nearly horizontal strata that dip gently northward. Exposed rock formations and deposits range in age from Middle Pennsylvanian to Holocene. Owing to the elevated and deeply dissected topography, only parts of the Cedar Mesa and White Rim Sandstone Members of the Cutler Formation of Permian age have potential for development of wells. Strata above and below them support only small springs, are dry, or contain brine. In the northwest part of the park, the Green River at Taylor Canyon is a potential source of surface water for public supplies for the Island In The Sky area and a small part of the northwest White Rim area. It will require filtration and treatment before use. In the same area, two unused wells in Taylor Canyon will supply enough water for present requirements from the White Rim Sandstone Member of the Cutler Formation, about 140 gallons per minute combined, but yield mineralized water that will require treatment before use. Springs yielding good water at the Island In The Sky and White Rim are mostly intermittent and too small for public-water supply. Most of the White Rim area is dry, having no usable ground water. In The Needles area, wells provide water of good quality from the Cedar Mesa Sandstone Member of the Cutler Formation. Springs yielding good water in the same area are available for supplementary supplies. West of The Needles, The Grabens area is without springs or potential aquifers bearing usable water. During 1970 about 510,000 gallons of water was used in Canyonlands National Park. Of this amount, 110,000 gallons was supplied to Island In The Sky by tank truck from a source outside the park, and about 400,000 gallons was withdrawn from the well in use at The Needles. Estimated total annual requirements in 10 years (1980) may be as much as 6 million gallons. Sources of water supplies within the park now in use and potential sources of surface water or ground water outlined by this investigation will meet the estimated requirements. Development of rainfall-collection and cistern-storage systems could furnish small emergency sources of water for waterless areas on the White Rim and in The Grabens.

Utah

Surficial Geologic Map of Mesa Verde National Park, Montezuma County, Colorado

Mesa Verde National Park in southwestern Colorado was established in 1906 to preserve and protect the artifacts and dwelling sites, including the famous cliff dwellings, of the Ancestral Puebloan people who lived in the area from about A.D. 550 to A.D. 1300. In 1978, the United Nations designated the park as a World Heritage Site. The geology of the park played a key role in the lives of these ancient people. For example, the numerous (approximately 600) cliff dwellings are closely associated with the Cliff House Sandstone of Late Cretaceous age, which weathers to form deep alcoves. In addition, the ancient people farmed the thick, red loess (wind-blown dust) deposits on the mesa tops, which because of its particle size distribution has good moisture retention properties. The soil in this loess cover and the seasonal rains allowed these people to grow their crops (corn, beans, and squash) on the broad mesa tops. Today, geology is still an important concern in the Mesa Verde area because the landscape is susceptible to various forms of mass movement (landslides, debris flows, rockfalls), swelling soils, and flash floods that affect the park's archeological sites and its infrastructure (roads, septic systems, utilities, and building sites). The map, which encompasses an area of about 100 mi 2 (260 km 2 ), includes all of Mesa Verde National Park, a small part of the Ute Mountain Indian Reservation that borders the park on its southern and western sides, and some Bureau of Land Management and privately owned land to the north and east. Surficial deposits depicted on the map include: artificial fills, alluvium of small ephemeral streams, alluvium deposited by the Mancos River, residual gravel on high mesas, a combination of alluvial and colluvial deposits, fan deposits, colluvial deposits derived from the Menefee Formation, colluvial deposits derived from the Mancos Shale, rockfall deposits, debris flow deposits, earthflow deposits, translational and rotational landslide deposits, rock rubble deposits, and loess. Bedrock units depicted on the map include the Cliff House Sandstone, Menefee Formation, Point Lookout Sandstone, and Mancos Shale all of Late Cretaceous age. In addition, minette dikes, of Oligocene age, found at several locations in the park are depicted on the map. Descriptions, including associated hazards and resources as used by the Ancestral Puebloans, are given for all map units.

Colorado

Water Quality of the Snake River and Five Eastern Tributaries in the Upper Snake River Basin, Grand Teton National Park, Wyoming, 1998-2002

To address water-resource management objectives of the National Park Service in Grand Teton National Park, the U.S. Geological Survey in cooperation with the National Park Service has conducted water-quality sampling in the upper Snake River Basin. Routine sampling of the Snake River was conducted during water years 1998-2002 to monitor the water quality of the Snake River through time. A synoptic study during 2002 was conducted to supplement the routine Snake River sampling and establish baseline water-quality conditions of five of its eastern tributaries?Pilgrim Creek, Pacific Creek, Buffalo Fork, Spread Creek, and Ditch Creek. Samples from the Snake River and the five tributaries were collected at 12 sites and analyzed for field measurements, major ions and dissolved solids, nutrients, selected trace metals, pesticides, and suspended sediment. In addition, the eastern tributaries were sampled for fecal-indicator bacteria by the National Park Service during the synoptic study. Major-ion chemistry of the Snake River varies between an upstream site above Jackson Lake near the northern boundary of Grand Teton National Park and a downstream site near the southern boundary of the Park, in part owing to the inputs from the eastern tributaries. Water type of the Snake River changes from sodium bicarbonate at the upstream site to calcium bicarbonate at the downstream site. The water type of the five eastern tributaries is calcium bicarbonate. Dissolved solids in samples collected from the Snake River were significantly higher at the upstream site (p-value<0.001), where concentrations in 43 samples ranged from 62 to 240 milligrams per liter, compared to the downstream site where concentrations in 33 samples ranged from 77 to 141 milligrams per liter. Major-ion chemistry of Pilgrim Creek, Pacific Creek, Buffalo Fork, Spread Creek, and Ditch Creek generally did not change substantially between the upstream sites near the National Park Service boundary with the National Forest and the downstream sites near the Snake River; however, variations in the major ions and dissolved solids existed between basins. Variations probably result from differences in geology between the tributary basins. Concentrations of dissolved ammonia, nitrite, and nitrate in all samples collected from the Snake River and the five eastern tributaries were less than water-quality criteria for surface waters in Wyoming. Concentrations of total nitrogen and total phosphorus in samples from the Snake River and the tributaries generally were less than median concentrations determined for undeveloped streams in the United States; however, concentrations in some samples did exceed ambient total-nitrogen and total-phosphorus criteria for forested mountain streams in the Middle Rockies ecoregion recommended by the U.S. Environmental Protection Agency to address cultural eutrophication. Sources for the excess nitrogen and phosphorus probably are natural because these basins have little development and cultivation. Concentrations of trace metals and pesticides were low and less than water-quality criteria for surface waters in Wyoming in samples collected from the Snake River and the five eastern tributaries. Atrazine, dieldrin, EPTC, or tebuthiuron were detected in estimated concentrations of 0.003 microgram per liter or less in 5 of 27 samples collected from the Snake River. An estimated concentration of 0.008 microgram per liter of metolachlor was detected in one sample from the Buffalo Fork. The estimated concentrations were less than the reporting levels for the pesticide analytical method. Suspended-sediment concentrations in 43 samples from the upstream site on the Snake River ranged from 1 to 604 milligrams per liter and were similar to suspended-sediment concentrations in 33 samples from the downstream site, which ranged from 1 to 648 milligrams per liter. Suspended-sediment concentrations in 38 samples collected from the tributary streams ranged from 1 t

Scientific Investigations Report

Streamflow and Water-Quality Characteristics for Wind Cave National Park, South Dakota, 2002-03

A 2-year study of streamflow and water-quality characteristics in Wind Cave National Park was performed by the U.S. Geological Survey in cooperation with the National Park Service. During this study, streamflow and water-quality data were collected for three of the park's perennial streams (Cold Spring, Beaver, and Highland Creeks) from January 2002 through November 2003. The potential influence of parking lot runoff on cave drip within Wind Cave also was investigated by collecting and analyzing several time-dependent samples from a drainage culvert downstream from the parking lot and from Upper Minnehaha Falls inside the cave following a series of simulated runoff events. The primary focus of the report is on data collected during the 2-year study from January 2002 to November 2003; however, data collected previously also are summarized. Losing reaches occur on both Beaver and Highland Creeks as these streams flow across outcrops of bedrock aquifers within the park. No streamflow losses occur along Cold Spring Creek because its confluence with Beaver Creek is located upstream from the outcrop of the Madison aquifer, where most streamflow losses occur. Physical properties, major ions, trace elements, nutrients, bacteria, benthic macroinvertebrates, organic (wastewater) compounds, bottom sediment, and suspended sediment are summarized for samples collected from 2 sites on Cold Spring Creek, 2 sites on Beaver Creek, and 1 site on Highland Creek. None of the constituent concentrations for any of the samples collected during 2002-03 exceeded any of the U.S. Environmental Protection Agency drinking-water standards, with the exception of the Secondary Maximum Contaminant Level for pH, which was exceeded in numerous samples from Beaver Creek and Highland Creek. Additionally, the pH values in several of these same samples also exceeded beneficial-use criteria for coldwater permanent fisheries and coldwater marginal fisheries. Water temperature exceeded the coldwater permanent fisheries criterion in numerous samples from all three streams. Two samples from Highland Creek also exceeded the coldwater marginal fisheries criterion for water temperature. Mean concentrations of ammonia, orthophosphate, and phosphorous were higher for the upstream site on Beaver Creek than for other water-quality sampling sites. Concentrations of E. coli, fecal coliform, and total coliform bacteria also were higher at the upstream site on Beaver Creek than for any other site. Samples for the analysis of benthic macroinvertebrates were collected from one site on each of the three streams during July 2002 and May 2003. The benthic macroinvertebrate data showed that Beaver Creek had lower species diversity and a higher percentage of tolerant species than the other two streams during 2002, but just the opposite was found during 2003. However, examination of the complete data set indicates that the quality of water at the upstream site was generally poorer than the quality of water at the downstream site. Furthermore, the quality of water at the upstream site on Beaver Creek is somewhat degraded when compared to the quality of water from Highland and Cold Spring Creeks, indicating that anthropogenic activities outside the park probably are affecting the quality of water in Beaver Creek. Samples for the analysis of wastewater compounds were collected at least twice from four of the five water-quality sampling sites. Bromoform, phenol, caffeine, and cholesterol were detected in samples from Cold Spring Creek, but only phenol was detected at concentrations greater than the minimum reporting level. Concentrations of several wastewater compounds were estimated in samples collected from sites on Beaver Creek, including phenol, para-cresol, and para-nonylphenol-total. Phenol was detected at both sites on Beaver Creek at concentrations greater than the minimum reporting level. Bromoform; para-cresol; ethanol,2-butoxy-phosphate; and cholesterol were detected

Scientific Investigations Report

Synthesis of the Hydrogeologic Framework of the Floridan Aquifer System and Delineation of a Major Avon Park Permeable Zone in Central and Southern Florida

The carbonate Floridan aquifer system of central and southern Florida (south of a latitude of about 29 degrees north) is an invaluable resource with a complex framework that has previously been mapped and managed primarily in a subregional context according to geopolitical boundaries. As interest and use of the Floridan aquifer system in this area increase, a consistent regional hydrogeologic framework is needed for effective management across these boundaries. This study synthesizes previous studies on the Floridan aquifer system and introduces a new regional hydrogeologic conceptual framework, linking physical relations between central and southern Florida and between the west and east coastal areas. The differences in hydrogeologic nomenclature and interpretation across the study area from previous studies were identified and resolved. The Floridan aquifer system consists of the Upper Floridan aquifer, middle confining unit, and Lower Floridan aquifer. This study introduces and delineates a new major, regional productive zone or subaquifer, referred to as the Avon Park permeable zone. This zone is contained within the middle confining unit and synthesizes an extensive zone that has been referred to differently in different parts of the study area in previous studies. The name of this zone derives from the description of this zone as the ?Avon Park highly permeable zone? in west-central Florida in a previous study. Additionally, this zone has been identified previously in southeastern Florida as the ?middle Floridan aquifer.? An approximately correlative or approximate time-stratigraphic framework was developed and was used to provide guidance in the identification and determination of aquifers, subaquifers, and confining units within the Floridan aquifer system and to determine their structural relations. Two stratigraphic marker horizons within the Floridan aquifer system and a marker unit near the top of the aquifer system were delineated or mapped. The marker horizons are correlative points in the stratigraphic section rather than a unit with upper and lower boundaries. The two marker horizons and the marker unit originated from previous studies, wherein they were based on lithology and correlation of geophysical log signatures observed in boreholes. The depths of these marker horizons and the marker unit were extended throughout the study area by correlation of natural gamma-ray logs between wells. The Floridan aquifer system includes, in ascending order, the upper part of the Cedar Keys Formation, Oldsmar Formation, Avon Park Formation, Ocala Limestone, Suwannee Limestone, and in some areas the lower part of the Hawthorn Group. The first marker horizon is in the lower part of the aquifer system near the top of the Oldsmar Formation and is associated with the top of distinctive glauconitic limestone beds that are present in some regions; the second marker horizon is near the middle of the aquifer system in the middle part of the Avon Park Formation. The marker unit lies at the top of a basal unit in the Hawthorn Group and provides a stratigraphic constraint for the top of the Floridan aquifer system. The marker horizons do not have distinguishing lithologic characteristics or a characteristic gamma-ray log pattern in all areas but are still thought to be valid because of correlation of the entire section and correlation of all sufficiently deep wells with gamma-ray logs. The Avon Park permeable zone is contained entirely within the Avon Park Formation; its position within the section is either near the middle Avon Park marker horizon or within a thick part of the section that extends several hundred feet above the marker horizon. This subaquifer is present over most of the study area and characteristically consists of thick units of dolostone and interbedded limestone, and limestone in its upper part. Permeability is primarily associated with fracturing. This subaquifer is well developed in west-cen

Scientific Investigations Report

Synthesis and interpretation of surface-water quality and aquatic biota data collected in Shenandoah National Park, Virginia, 1979-2009

Shenandoah National Park in northern and central Virginia protects 777 square kilometers of mountain terrain in the Blue Ridge physiographic province and more than 90 streams containing diverse aquatic biota. Park managers and visitors are interested in the water quality of park streams and its ability to support healthy coldwater communities and species, such as the native brook trout ( Salvelinus fontinalis ), that are at risk in the eastern United States. Despite protection from local stressors, however, the water quality of streams in the park is at risk from many regional stressors, including atmospheric pollution, decline in the health of the surrounding forests because of invasive forest pests, and global climate change. In 2010, the U.S. Geological Survey, in cooperation with the National Park Service, undertook a study to compile, analyze, and synthesize available data on water quality, aquatic macroinvertebrates, and fish within Shenandoah National Park. Specifically, the effort focused on creating a comprehensive water-resources database for the park that can be used to evaluate temporal trends and spatial patterns in the available data, and characterizing those data to better understand interrelations among water quality, aquatic macroinvertebrates, fish, and the landscape.

Virginia

Spring types and contributing aquifers from water-chemistry and multivariate statistical analyses for seeps and springs in Theodore Roosevelt National Park, North Dakota, 2018

Water resources in Theodore Roosevelt National Park, North Dakota, support wildlife, visitors, and staff, and play a vital role in supporting the native ecology of the park. The U.S. Geological Survey, in cooperation with the National Park Service, completed field work in 2018 for a study to address concerns about water availability and possible sources of groundwater contamination for seeps and springs in Theodore Roosevelt National Park. The objective of the study was to improve hydrologic knowledge and determine the water composition of 11 seeps and springs in the park by collecting water-chemistry data at springs, streams, wells, and rain collectors. Water samples were collected at 26 sites at springs, streams, wells, and rain collectors in the North and South Units of Theodore Roosevelt National Park. Samples in the North Unit were collected at 5 springs, 1 stream, 2 wells, and 1 rain collector. Samples in the South Unit were collected at 6 springs, 2 streams, 8 wells, and 1 rain collector. Samples from springs, streams, and wells were collected in May, July, and September 2018. Samples from rain collectors were collected when enough daily precipitation accumulated in the collectors. Sampled precipitation events during the study period were in May, June, July, August, and September 2018. Physical properties of sampled water—temperature, pH, and specific conductance—were measured in the field. Water samples were analyzed for stable isotopes of oxygen and hydrogen and for chloride concentration. Recharge rates for aquifers supplying springs were determined using precipitation volume and chloride concentrations for a 12-day period before the sample-collection date. Multivariate statistical analysis methods used on water-chemistry data included principal component analysis, cluster analysis, and end-member mixing analysis. Water composition was used to determine the spring type and contributing aquifers for 11 springs in the North and South Units of Theodore Roosevelt National Park from analyses of water-chemistry data between May and September 2018. In the North Unit, Achenbach Spring was classified as a filtration spring with water from an unconfined part of the upper Fort Union aquifer and infiltration of precipitation. Hagen Spring, Mandal Spring, and Stevens Spring were classified as contact springs supplied by semiconfined parts of the upper Fort Union aquifer. Overlook Spring at one time may have been a natural spring or seep but now is a developed spring that behaves like a flowing artesian well completed in a confined part of the upper Fort Union aquifer. In the South Unit, six springs were classified into two spring types: filtration and contact springs. Boicourt Spring and Sheep Butte Spring were classified as filtration springs that have water supplied by unconfined parts of the upper Fort Union aquifer and infiltrated precipitation. Big Plateau Spring, Lone Tree Spring, Sheep Pasture Spring, and Southeast Corner Spring were classified as contact springs that receive waters from a semiconfined part of the upper Fort Union aquifer.

North Dakota

Correlation analysis of groundwater and hydrologic data, Kaloko-Honokōhau National Historical Park, Hawai‘i

Designated in 1978, Kaloko-Honokōhau National Historical Park is located on the west coast of the Island of Hawaiʻi. The Kaloko-Honokōhau National Historical Park encompasses about 1,200 acres of coastal land and nearshore ecosystems, which include wetlands, anchialine pools (landlocked bodies of brackish water with hydrologic connections to the ocean), fishponds, a fishtrap, and coral reefs. These nearshore ecosystems are dependent on groundwater discharge with a freshwater component and provide habitat for threatened and endangered, endemic species, such as the orangeblack Hawaiian damselfly ( Megalagrion xanthomelas ) and the Hawaiian coot (ʻAlae keʻokeʻo, Fulica alai ). The populations of these native species, however, are threatened because of habitat loss related to urban development and environmental changes. Kaloko-Honokōhau National Historical Park is within the Keauhou aquifer system and the North Kona District, which experienced a 52 percent resident-population increase between 2000 and 2020 and a 41 percent visitor increase between 2008 and 2019. To support the current water demand associated with this growing population, groundwater is the primary source of freshwater used in the North Kona District, with about 15 million gallons of groundwater withdrawn from the Keauhou aquifer system per day since 2009. With anticipated development, future (2015–35) groundwater withdrawal from the Keauhou aquifer system is projected to be about 55 percent greater than recent (2012–14) withdrawal. Because Kaloko-Honokōhau National Historical Park is located within a coastal aquifer, natural and human-induced changes can affect the quality and quantity of groundwater, which can threaten groundwater-dependent ecosystems. To improve understanding of recent groundwater conditions, the U.S. Geological Survey, in cooperation with the National Park Service, undertook this study to document correlations between hydrologic time-series datasets from sites in and near Kaloko-Honokōhau National Historical Park using the nonparametric (distribution-free) Kendall’s tau statistical test. For the statistical analyses, dependent variables representing the groundwater system include groundwater level, the groundwater-level difference between pairs of sites, and specific conductance, and independent variables include datasets of sea level, rainfall, and groundwater withdrawal. About 34 percent of the 140 non-time-lagged Kendall’s tau statistical tests evaluated in this report are statistically significant ( p -value ≤ 0.050) with generally weak (0.1 ≤ tau ≤ 0.2) to moderate (0.2 ≤ tau ≤ 0.3) correlations. Groundwater levels measured at monitoring sites have the strongest correlation with the multivariate El Niño–Southern Oscillation index and withdrawal from production wells at the nearby Kohanaiki Private Club Community. Specific conductance is not consistently and significantly correlated with the independent hydrologic variables investigated in this report. Because the relations between hydrologic variables are commonly not instantaneous, a second set of correlations was evaluated after applying a range of time lags to the independent variable datasets. Relative to the non-time-lagged case (the set of correlations that did not use time-lagged independent variables), some of the time-lagged independent variables improved correlations with some of the dependent variables. For a particular independent variable, similar time lags were expected between the independent variable and dependent variable at all four monitoring sites. However, different time lags among the four sites sometimes produced the strongest correlations. This study identified several correlations that are statistically significant and hydrologically plausible, but the correlations could indicate that multiple concurrent factors are controlling the observed groundwater-system response, which might be better addressed using multivariate analyses. This study only investigates bivariate correlations, which may not explain all the variance in the data. The correlations analyzed in this report are limited by the quantity of available hydrologic data in the area near Kaloko-Honokōhau National Historical Park and are based on 14 years of time-series data, which were aggregated to a relatively coarse monthly temporal resolution that represents the minimum resolution common to all datasets.

Hawaii

Vegetation community monitoring: Species composition and biophysical gradients in Klamath Network parks

The Klamath Network of the National Park Service consists of six park units located in northern California and southern Oregon. The Network began implementing a vegetation monitoring protocol in 2011 to identify ecologically significant vegetation trends in the parks. The premise of the protocol is that multivariate analyses of species composition data is the most robust early detection means for identifying vegetation change over time. Here we present these community metrics, based on our initial sampling efforts. We use these metrics to establish a baseline for comparison in future trend analysis, and to evaluate the adequacy of the protocol for meeting the Network’s objectives of detecting temporal changes across contrasting vegetation types. The park landscapes were subdivided into three strata: Matrix (low- to mid-elevation upland habitats), Riparian (within 10 meters of a perennial stream), and High-Elevation (above a predefined elevation, park specific). Across the three strata, we established a total of 241 permanent plots at random locations to measure complete species composition and cover. We describe baseline biophysical conditions and relate them to the data obtained from all 241 plots using ordination analyses. The unconstrained gradient analyses were moderately robust at illustrating the relationships among plots and correlating them to environmental gradients. We also prepared species accumulation curves representing gamma diversity, which showed overall species richness, and also illustrated how well the observed vs. expected richness values of each stratum were captured by the sampling. Most park/strata were well sampled; for others, we found that additional samples would improve how well the protocol captures the vegetation composition within park/strata. Specifically, all sample frames at Whiskeytown and the High-Elevation sample frames at Lassen were not well sampled. Comparisons of alpha diversity values showed High-Elevations had the lowest diversity, while Riparian areas were by far the most diverse across all parks. The Matrix stratum at Oregon Caves National Monument was also especially diverse and had the highest Matrix alpha diversity we observed in all parks We suggest that after three rounds of sampling, the Network perform analyses to identify possible ways to improve statistical power. These options include adding sites or lengthening the sampling interval. Results of these analyses could support protocol modifications. This report on vegetation composition is the first in a series of analysis and synthesis reports. Future analysis and synthesis reports will analyze structure and function.

California, Oregon

Environmental contaminant hazards to wildlife at National Capital region and Mid-Atlantic National Park Service units

Pollutant data for air, water, soil and biota were compiled from databases and internet sources and by staff interviews at 23 National Park Service (NPS) units in 2005. A metric was derived describing the quality and quantity of data for each park, and in combination with known contaminant threats, the need for ecotoxicological study was identified and ranked. Over half of NP units were near Toxic Release Inventory sites discharging persistent pollutants, and fish consumption advisories were in effect at or near 22 of the units. Pesticide and herbicide use was found to be minimal, with the exception of those units with agricultural leases. Only 70 reports were found that describe terrestrial vertebrate environmental contaminant data at or near the units. Of the >75,000 compounds in commerce, empirical exposure data were limited to merely 58 halogenated compounds, insecticides, rodenticides, metals, and some contemporary compounds. Further ecotoxicological monitoring and research is warranted at several units including Shenandoah National Park, Richmond National Battlefield Park, Chesapeake & Ohio Canal National Historical Park, Valley Forge National Historical Park, Hopewell Furnace National Historic Site, Monocacy National Battlefield, and Harpers Ferry National Historical Park. The types of investigations vary according to the wildlife species present and potential contaminant threats, but should focus on contemporary use pesticides and herbicides, polychlorinated biphenyls, mercury, lead, and perhaps antibiotics, flame retardants, pharmaceuticals, and surfactants. Other management recommendations include inclusion of screening level contaminant risk assessments into the NPS Vital Signs Program, development of protocols for toxicological analysis of seemingly affected wildlife, alternative methods and compounds for pest management, and use of non-toxic fishing tackle by visitors.

Book chapter

A review of the relationships between visitors and ungulates in national parks

The growth in ungulate populations and the enhanced viewing opportunities they provide in many national parks, a mixed blessing to park managers, has been viewed positively by most park visitors. Visitors, long the major constituency of parks, have played a major role in the way ungulates are managed. I trace the history of the relationships between visitors and ungulates in parks and provide examples of how visitors have influenced ungulate management policies. Today, although public influence on ungulate management seems to be greater than ever, the public has a poor understanding of park management actions. Park interpretive programs can be used to address this problem.

Wildlife Society Bulletin

Birds in Hawai'i Volcanoes National Park: Summary of the 2010 inventory and monitoring program survey

The National Park Service (NPS) created the Inventory and Monitoring (I&M) Program in 1998 to establish baseline information and assess long-term trends in "vital signs" or key abiotic and biotic elements of National Parks (Fancy et al. 2009). The Pacific Island Network of the I&M Program developed a Landbirds Monitoring Protocol (LMP; Camp et al. 2011) to estimate species-specific status and monitor longterm trends in landbird distribution and abundance. Parks included in the LMP that harbor habitat critically important to native forest birds are Haleakala National Park (Maui Island), Hawai'i Volcanoes National Park (HAVO; Hawai'i Island), and the National Park of American Samoa (American Samoa). In 2010, the LMP was implemented in HAVO to survey landbird density and abundance. This implementation was the first anywhere in the Pacific Islands by the I&M Program, and continued monitoring is planned every five years in all three parks.

Hawai'i

Elk monitoring in Lewis and Clark National Historical Park: 2008-2012 synthesis report

Maintaining elk (Cervus elaphus roosevelti) herds that frequent Lewis and Clark National Historical Park (NHP) is central to the park’s purpose of preserving the historic, cultural, scenic, and natural resources associated with the winter encampment of the Lewis and Clark expedition. Elk were critically important to the Lewis and Clark expedition in providing food and hides that sustained the expedition during the winter of 1805-06 and supplied them for their return east during 1806. Today, elk remain a key component of interpreting the Lewis and Clark story to over 200,000 park visitors each year at the Fort Clatsop visitor center. In 2008, the US Geological Survey (USGS) began collaborating with Lewis and Clark NHP and the NPS North Coast and Cascades Network to develop a protocol for monitoring long-term changes in the magnitude and spatial patterns of elk use within and adjacent to Lewis and Clark NHP (Griffin et al. 2011). Specific objectives of the monitoring program were to measure trends in (1) relative use of the Fort Clatsop unit by elk during winter; (2) the proportion of areas where elk sign is present in the Fort Clatsop unit in winter; and (3) the frequency of elk sightings from roads in and around the Fort Clatsop unit. This report synthesizes the results of the first four years of monitoring elk distribution and use in Lewis and Clark NHP from 2008-2012. We also present data from FY2012 (Appendix 1), in lieu of an annual report for that year. We used fecal pellet group surveys as the cornerstone for monitoring trends in both relative use of the Fort Clatsop Unit by elk and the proportion of areas where elk sign was present at the end of winter. We estimated pellet group density based on data collected from a network of fecal pellet plots distributed systematically throughout the unit. We developed a double observer sampling scheme that enabled us to estimate detection biases and improve the accuracy of pellet group density estimates. We computed the estimated detection probability for any pellet group observed; this probability was a function of the pellet group size and stage of decay, as well as lighting and vegetation conditions, and the number of observers (one or two) searching for pellets in that subplot. We then used these estimated detection probabilities to adjust the raw counts of the detected pellet groups to account for groups that likely went undetected under similar pellet and environmental conditions (each observed pellet group was weighted by the inverse of its estimated detection probability). We also used results from the late winter fecal pellet surveys to quantify the proportion of areas where elk pellets occurred (PAO), which was based on the presence of fecal pellet groups and estimation of detection biases (i.e., accounting for pellet groups that likely went undetected by both observers). In this synthesis, we report temporal trends in both pellet group density and PAO from 2008-2012, based on weighted linear regression analyses as well as spatial variation of pellet group densities over time. We completed late winter fecal pellet surveys at 61-66 plots annually, depending on yearly variation in access. We cleared fecal pellets at survey points in late October / early November each year and returned in late February / early March to count pellet groups left by elk over the winter. The estimated probability that a pellet group was detected by any one observer during late winter was affected most by the pellet group size and was less affected by decay class and lighting conditions. Per-observer detection probabilities ranged from as low as ~10-15% for single pellets to ~85-90% for pellet groups with 50 pellets. Average pellet group density in the Fort Clatsop unit ranged annually from 0.58 (+/- 1.43 standard error [SE]) to 0.93 (+/- 2.25 SE) pellet groups per 3-m radius subplot. Pellet group density declined over time, at approximately 8.8% per year (+/- 2.5% SE), but that slope was not statistically distinguishable from zero (2- tailed P=0.16). Following correction for detection biases, the proportion of surveyed points used by elk (i.e., PAO) ranged from 0.44 (+/- 0.07 SE) to 0.53 (+/- 0.07 SE) during the 4 winters. The estimated proportion of areas where elk pellets occurred (PAO) declined at a rate of 2.6% per year (+/- 1.2% per year SE), but that trend also was not statistically distinguishable from zero (2- tailed P=0.17). Statistical significance of a measure’s trend depends on both the magnitude (i.e., slope) of the observed trend and the number of years the trend continues in the same increasing or decreasing direction. Through simulation modeling we determined how many additional years of surveys would be required to reveal a statistically significant trend, based on the same trends in pellet group density and PAO, and associated variation, observed from 2009-2012. Assuming the same trends persist in the future, simulations indicated that there is a 70% probability that a statistically significant trend would be detected after two more years of conducting pellet group surveys. Relative use by elk during winter, as indexed by elk pellet group density, was generally greatest in the southeast region of the Fort Clatsop unit in or near the large freshwater marsh at the mouth of Colewort Creek and adjacent upland areas. Pellet group density was also higher than average in the north-central forested area, not far from a privately-owned pasture north of the park boundary. This spatial pattern in pellet group densities across the Fort Clatsop unit was consistent across all four years, although specific pellet group densities varied from year to year. Pellet group density declined significantly over time at two points in the southeast of the Fort Clatsop unit, even though pellet group density at those points remained higher than the unit average. Pellet group density increased significantly over time at one point in the north-central region, and at one point in the south-central region of the unit, indicating a slight shift in the distribution of elk use within the Fort Clatsop Unit over the four years. As an index of visitors’ opportunities to see elk in and around the Fort Clatsop Unit, we conducted replicated roadside elk surveys 3-5 times monthly during February, April, June, August, October and December 2008-2012. During each morning of survey, we searched for elk along four routes that totaled 32 km. We examined bimonthly trends in the numbers of elk groups seen, the total number of elk seen, and the observed composition ratios for those six months of the year. The average number of elk groups seen per survey ranged from 0.75 (+/- 0.32 SE) during February to a peak of 1.95 (+/- 0.36 SE) during June. Despite this seasonal variation in numbers of elk groups seen, the average total number of elk seen per morning was less variable. The average ratios of antlered elk to antlerless adult elk (i.e., bulls:cows) and calves to antlerless adult elk (i.e. calves:cows) varied seasonally, with the highest of both average ratios observed in August. We detected no significant trends in the average number of elk groups and total numbers of elk seen per survey from 2008-2012. Similarly, ratios of calves and antlered elk per antlerless elk did not differ over time. Elk groups were frequently seen from January to August in the southeast region of the Fort Clatsop unit, in the vicinity of Colewort Creek. Outside of NPS lands, we observed elk most frequently in open areas near the Astoria regional airport, in the pastures and forests immediately north of the Fort Clatsop unit and, prior to the construction of a residential development, in a pasture northwest of the Fort Clatsop unit. Elk monitoring at Lewis and Clark NHP is still in its initial years and additional monitoring will be required to verify trends that appear to be emerging. For example, the initial monitoring suggested incipient declining trends in both pellet group density and proportion of plots with pellets present, as well as, potentially, a small shift in elk distribution away from a new trail that was recently constructed in the southeast portion of the Fort Clatsop unit. Continued monitoring will aid in determining whether this local change in distribution persists (or, alternatively, resulted from short-term random variation), and whether there will be any positive or negative effect in the northern portion of the unit where a new trail has been constructed. High variability in road counts prevented our ability to find any clear trend in numbers or composition of elk observed in and near Fort Clatsop, but changes in the patterns of observations of elk from roadways suggest that residential development outside the park has reduced the available habitat for elk in some of the areas surveyed, and may have affected spatial use patterns of elk adjacent to some areas of the park. In addition to monitoring future effects of land use changes outside the park, continued monitoring may also prove useful for assessing elk responses to natural succession in forests disturbed by windthrow in December 2007 and to NPS vegetation management activities such as variable density thinning in the forest, trail development, and restoration at Otter Point tidal area and Colewort Creek Slough.

Oregon;Washington

Fire history of southeastern Glacier National Park: Missouri River Drainage

In 1982, Glacier National Park (GNP) initiated long-term studies to document the fire history of all forested lands in the 410,000 ha. park. To date, studies have been conducted for GNP west of the Continental Divide (Barrett et al. 1991), roughly half of the total park area. These and other fire history studies in the Northern Rockies (Arno 1976, Sneck 1977, Arno 1980, Romme 1982, Romme and Despain 1989, Barrett and Arno 1991, Barrett 1993a, Barrett 1993b) have shown that fire history data can be an integral element of fire management planning, particularly wen natiral fire plans are being developed for parks and wilderness. The value of site specific fire history data is apparent when considering study results for lodgepole pin ( Pinus contorta var. latifolia ) forests. Lodgepole pine is a major subalpine type in the Northern Rockies and such stands experiences a wide range of presettlement fire patterns. On relatively warm-dry sites at lower elevations, such as in GNP's North Fork drainage (Barrett et al. 1991), short to moderately long interval (25-150 yr) fires occurred in a mixed severity pattern ranging from non-lethal underburns to total stand replacement (Arno 1976, Sneck 1977, Barrett and Arno 1991). Markedly different fire history occurred at high elevation lodgepole pine stands on highly unproductive sites, such as on Yellowstone National Park's (YNP) subalpine plateau. Romme (1982) found that, on some sites, stand replacing fires recurred after very long intervals (300-400 yr), and that non-lethal surface fires were rare. For somewhat more productive sites in the Absaroka Mountains in YNP, Barrett (1993a) estimated a 200 year mean replacement interval, in a pattern similar to that found in steep mountain terrain elsewhere, such as in the Middle Fork Flathead River drainage (Barrett et al. 1991, Sneck 1977). Aside from post-1900 written records (ayres 1900; fire atlas data on file, GNP Archives Div. and GNP Resources Mgt. Div.), little fire history information existed for GNP's east-side forests, which are dominated primarily by lodgepole pine. In fall 1992, the park initiated a study to determine the fire history of the Missouri River drainage portion of southeastern GNP. Given the known variation in pre-1900 fire patterns for lodgepole pine, this study was seen as a potentially important contribution to GNP's Fire Management Plan, and to the expanding data base of fire history studies in the region. Resource managers sought this information to assist their development of appropriate fire management strategies for the east-side forests, and the fire history data also would be a useful interactive component of the park's Geographic Information System (GIS). Primary objectives were to: 1) determine pre-1900 fire periodicities, severities, burning patterns, and post-fire succession for major forest types, and 2) document and map the forest age class mosaic, reflecting the history of stand replacing fires at the landscape level of analysis. Secondary objectives were to interpret the possible effects of modern fire suppression on area forests, and to determine fire regime patterns relative to other lodgepole pine ecosystems in the Northern Rockies.

Montana

2013 National Park visitor spending effects: economic contributions to local communities, states, and the nation

The National Park Service (NPS) manages the nation's most iconic destinations that attract millions of visitors form across the nation and around the world. Trip-related spending by NPS visitors generates and supports a considerable amount of economic activity within park gateway communities. This economic effects analysis measures how NPS visitor spending cycles through local economies, generating business sales and supporting jobs and income. In 2013, the National Park System received over 273 million recreation visits. NPS visitors spent $14.6 billion in local gateway regions (defined as communities within 60 miles of a park). The contribution of this spending to the national economy was 238 thousand jobs, $9.2 billion in labor income, $15.6 billion in value added, and $26.5 billion in output. The lodging sector saw the highest direct contributions with 38 thousand jobs and $4.4 billion in output directly contributed to local gateway economies nationally. The sector with the next greatest direct contributions was restaurants and bats, with 50 thousand jobs and $2.0 billion in output directly contributed to local gateway economies nationally. While it is typical for visitation levels to fluctuate across the park units each year, system-wide visitation estimates in 2013 declines by 3.2% (or 9.1 million visits) compared ro 2012 (Street, 2014). Although many factors can influence park visitsation, two events signficiantly contrubuterd to this decline: the Government shutdown in October 2013, and lonf-term park closures related to the lasting effects of Hurrican Sandt from October 2012 through July 2013.

Natural Resource Report

2014 National Park visitor spending effects: economic contributions to local communities, states, and the nation

The National Park System covers more than 84 million acres and is comprised of more than 401 sites across the Nation. These lands managed by the National Park Service (NPS) serve as recreational destinations for visitors from across the Nation and around the world. On vacations or on day trips, NPS visitors spend time and money in the gateway communities surrounding NPS sites. Spending by NPS visitors generates and supports a considerable amount of economic activity within park gateway economies. The NPS has been measuring and reporting visitor spending and economic effects for the past 25 years. The 2012 analysis marked a major revision to the NPS visitor spending effects analyses, with the development of the Visitor Spending Effects model (VSE model) which replaced the previous Money Generation Model (see Cullinane Thomas et al. (2014) for a description of how the VSE model differs from the previous model). This report provides updated VSE estimates associated with 2014 NPS visitation. System-wide visitation estimates in 2014 increased by 7% (or 19.2 million visits) compared to 2013 (Ziesler, 2015). Visitation in 2014 rebounded from a 2013 decline that included a 16-day government shutdown and many park closures for repairs after Superstorm Sandy hit the Northeast in late 2012. The re-opening of the Washington Monument, some 21 months after it was rocked by an earthquake and repaired, also added to 2014 visitation numbers. Additionally, several national parks saw record-breaking visitation in 2014, including Joshua Tree, Rocky Mountain, Grand Teton and Glacier national parks. This report begins by presenting an overview of economic effects analyses, followed by details about the data and methods used for this analysis and 2014 model updates. Estimates of NPS visitor spending in 2014 and resulting economic effects at the local, state, regional, and national levels are then presented. The report concludes with a description of current data limitations. Park-level spending and economic effects estimates are included in the appendix. New this year, results from the Visitor Spending Effects report series are available online via an interactive tool. Users can explore current year visitor spending, jobs, labor income, value added, and output effects by sector for national, state, and local economies. This interactive tool is available via the NPS Social Science Program webpage at http://www.nature.nps.gov/socialscience/economics.cfm.

Natural Resource Report

Evaluating genetic viability of pronghorn in Wind Cave National Park

The pronghorn (Antilocapra americana) was reintroduced into Wind Cave National Park, South Dakota, in 1914 and thus, has inhabited the Park for almost a century. A decline in the population has raised concern for the continued existence of pronghorn inside Wind Cave National Park. Historically, pronghorn numbers reached greater than 300 individuals in the 1960&rsquo;s but declined to about 30 individuals by 2002. The primary objective of our study was to evaluate genetic characteristics of pronghorn to determine if reduced heterozygosity contributed to the decline of pronghorn in Wind Cave National Park. Microsatellite DNA was collected from 75 pronghorn inhabiting Wind Cave National Park in western South Dakota (n = 11), northwestern South Dakota (n = 33), and southwestern South Dakota (n = 31). Pronghorn in Wind Cave National Park had similar levels of observed heterozygosity (0.473 to 0.594) and low inbreeding coefficients (-0.168 to 0.037) when compared with other populations in western South Dakota. Furthermore, indices of population structure indicated no differentiation occurred among pronghorn populations. Results indicated that genetic variability was not a primary factor in the decline of pronghorn in Wind Cave National Park.

Prairie Naturalist