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Groundwater: The disregarded component in lake water and nutrient budgets, Part 1: Effects of groundwater on hydrology

Lake eutrophication is a large and growing problem in many parts of the world, commonly due to anthropogenic sources of nutrients. Improved quantification of nutrient inputs is required to address this problem, including better determination of exchanges between groundwater and lakes. This first of a two-part review provides a brief history of the evolution of the study of groundwater exchange with lakes, followed by a listing of the most commonly used methods for quantifying this exchange. Rates of exchange between lakes and groundwater compiled from the literature are statistically summarized for both exfiltration (flow from groundwater to a lake) and infiltration (flow from a lake to groundwater), including per cent contribution of groundwater to lake-water budgets. Reported rates of exchange between groundwater and lakes span more than five orders of magnitude. Median exfiltration is 0.74 cm/day, and median infiltration is 0.60 cm/day. Exfiltration ranges from near 0% to 94% of input terms in lake-water budgets, and infiltration ranges from near 0% to 91% of loss terms. Median values for exfiltration and infiltration as percentages of input and loss terms of lake-water budgets are 25% and 35%, respectively. Quantification of the groundwater term is somewhat method dependent, indicating that calculating the groundwater component with multiple methods can provide a better understanding of the accuracy of estimates. The importance of exfiltration to a lake budget ranges widely for lakes less than about 100 ha in area but generally decreases with increasing lake area, particularly for lakes that exceed 100 ha in area. No such relation is evident for lakes where infiltration occurs, perhaps because of the smaller sample size. Copyright © 2014 John Wiley & Sons, Ltd.

Hydrological Processes↗

Short-term and long-term evapotranspiration rates at ecological restoration sites along a large river receiving rare flow events

Many large rivers around the world no longer flow to their deltas, due to ever greater water withdrawals and diversions for human needs. However, the importance of riparian ecosystems is drawing increasing recognition, leading to the allocation of environmental flows to restore river processes. Accurate estimates of riparian plant evapotranspiration (ET) are needed to understand how the riverine system responds to these rare events and achieve the goals of environmental flows. In 2014, historic environmental flows were released into the Lower Colorado River at Morelos Dam (Mexico); this once perennial but now dry reach is the final stretch to the mighty Colorado River Delta. One of the primary goals was to supply native vegetation restoration sites along the reach with water to help seedlings establish and boost groundwater levels to foster the planted saplings. Patterns in ET before, during, and after the flows are useful for evaluating whether this goal was met and understanding the role that ET plays in this now ephemeral river system. Here, diurnal fluctuations in groundwater levels and MODIS data were used to compare estimates of ET specifically at three native vegetation restoration sites during 2014 planned flow events, while MODIS data was used to evaluate long-term (2002 – 2016) ET responses to restoration efforts at these sites. Overall, ET was generally 0 - 10 mm d -1 across sites and although daily ET values from groundwater data were highly variable, weekly averaged estimates were highly correlated with MODIS-derived estimates at most sites. The influence of the 2014 flow events was not immediately apparent in the results, although the process of clearing vegetation and planting native vegetation at the restoration sites was clearly visible in the results.

Colorado River↗

Distribution of oxygen-18 and deuteriun in river waters across the United States

Reconstruction of continental palaeoclimate and palaeohydrology is currently hampered by limited information about isotopic patterns in the modern hydrologic cycle. To remedy this situation and to provide baseline data for other isotope hydrology studies, more than 4800, depth- and width-integrated, stream samples from 391 selected sites within the USGS National Stream Quality Accounting Network (NASQAN) and Hydrologic Benchmark Network (HBN) were analysed for δ 18 O and δ 2 H ( http://water.usgs.gov/pubs/ofr/ofr00-160/pdf/ofr00-160.pdf ). Each site was sampled bimonthly or quarterly for 2·5 to 3 years between 1984 and 1987. The ability of this dataset to serve as a proxy for the isotopic composition of modern precipitation in the USA is supported by the excellent agreement between the river dataset and the isotopic compositions of adjacent precipitation monitoring sites, the strong spatial coherence of the distributions of δ 18 O and δ 2 H, the good correlations of the isotopic compositions with climatic parameters, and the good agreement between the ‘national’ meteoric water line (MWL) generated from unweighted analyses of samples from the 48 contiguous states of δ 2 H=8·11δ 18 O+8·99 ( r 2 =0·98) and the unweighted global MWL of sites from the Global Network for Isotopes in Precipitation (GNIP) of the International Atomic Energy Agency and the World Meteorological Organization (WMO) of δ 2 H=8·17δ 18 O+10·35. The national MWL is composed of water samples that arise in diverse local conditions where the local meteoric water lines (LMWLs) usually have much lower slopes. Adjacent sites often have similar LMWLs, allowing the datasets to be combined into regional MWLs. The slopes of regional MWLs probably reflect the humidity of the local air mass, which imparts a distinctive evaporative isotopic signature to rainfall and hence to stream samples. Deuterium excess values range from 6 to 15‰ in the eastern half of the USA, along the northwest coast and on the Colorado Plateau. In the rest of the USA, these values range from −2 to 6‰, with strong spatial correlations with regional aridity. The river samples have successfully integrated the spatial variability in the meteorological cycle and provide the best available dataset on the spatial distributions of δ 18 O and δ 2 H values of meteoric waters in the USA.

Hydrological Processes↗

A targeted approach for mapping groundwater discharge to surface water and fish thermal refuge in four Lake Ontario tributaries

The duration, magnitude, and frequency of heatwaves are predicted to increase in the coming decades, a combination that can reduce the survival of many fish species. Across the world, there is broad interest in identifying thermal refuge for heat-intolerant fish species and exploring opportunities to enhance or protect these areas. Because deeper groundwater maintains a relatively constant temperature, groundwater-influenced areas along streams can provide cool-water refuge for fish during periods of extreme heat. A targeted approach was developed for identifying existing cold-water zones and areas of substantial groundwater discharge in four high priority Lake Ontario tributaries. Our approach included: (1) predicting where groundwater discharge is most likely with a simple geospatial model and (2) using model predictions to select field sites for intensive high-resolution study, including ground-based mapping of groundwater features (springs, seeps, tributaries) as well as drone-based optical and thermal infrared surveys. Results from field sites were used to both verify model performance and map different types and aerial extents of thermal anomalies. Geospatial modelling successfully predicted regions of widespread groundwater upwelling, later verified and mapped by field and drone surveys. Comparison of model and field survey results further highlighted specific geospatial layers, such as soil/bedrock types and topographic wetness index, as being particularly useful for predicting groundwater influence on streams in the study area. In addition, a comparison of geospatial model results with a model of fish abundances along the studied streams showed significant positive correlations for many heat-intolerant fish species over a wide geographic area. The approach developed in this study can be applied to other watersheds to highlight areas of probable groundwater discharge and could be used by fishery and water resource managers to support cold-water fish habitat management decision-making and resource conservation.

New York↗

First-order exchange coefficient coupling for simulating surface water-groundwater interactions: Parameter sensitivity and consistency with a physics-based approach

Distributed hydrologic models capable of simulating fully-coupled surface water and groundwater flow are increasingly used to examine problems in the hydrologic sciences. Several techniques are currently available to couple the surface and subsurface; the two most frequently employed approaches are first-order exchange coefficients (a.k.a., the surface conductance method) and enforced continuity of pressure and flux at the surface-subsurface boundary condition. The effort reported here examines the parameter sensitivity of simulated hydrologic response for the first-order exchange coefficients at a well-characterized field site using the fully coupled Integrated Hydrology Model (InHM). This investigation demonstrates that the first-order exchange coefficients can be selected such that the simulated hydrologic response is insensitive to the parameter choice, while simulation time is considerably reduced. Alternatively, the ability to choose a first-order exchange coefficient that intentionally decouples the surface and subsurface facilitates concept-development simulations to examine real-world situations where the surface-subsurface exchange is impaired. While the parameters comprising the first-order exchange coefficient cannot be directly estimated or measured, the insensitivity of the simulated flow system to these parameters (when chosen appropriately) combined with the ability to mimic actual physical processes suggests that the first-order exchange coefficient approach can be consistent with a physics-based framework. Copyright ?? 2009 John Wiley & Sons, Ltd.

Hydrological Processes↗

Estimating basin scale evapotranspiration (ET) by water balance and remote sensing methods

Evapotranspiration (ET) is an important hydrological process that can be studied and estimated at multiple spatial scales ranging from a leaf to a river basin. We present a review of methods in estimating basin scale ET and its applications in understanding basin water balance dynamics. The review focuses on two aspects of ET: (i) how the basin scale water balance approach is used to estimate ET; and (ii) how ‘direct’ measurement and modelling approaches are used to estimate basin scale ET. Obviously, the basin water balance-based ET requires the availability of good precipitation and discharge data to calculate ET as a residual on longer time scales (annual) where net storage changes are assumed to be negligible. ET estimated from such a basin water balance principle is generally used for validating the performance of ET models. On the other hand, many of the direct estimation methods involve the use of remotely sensed data to estimate spatially explicit ET and use basin-wide averaging to estimate basin scale ET. The direct methods can be grouped into soil moisture balance modelling, satellite-based vegetation index methods, and methods based on satellite land surface temperature measurements that convert potential ET into actual ET using a proportionality relationship. The review also includes the use of complementary ET estimation principles for large area applications. The review identifies the need to compare and evaluate the different ET approaches using standard data sets in basins covering different hydro-climatic regions of the world.

Hydrological Processes↗

Rethinking infiltration in wildfire-affected soils

Wildfires frequently result in natural hazards such as flash floods (Yates et al ., 2001) and debris flows (Cannon et a l., 2001a,b; Gabet and Sternberg, 2008). One of the principal causes of the increased risk of post-wildfire hydrologically driven hazards is reduced in filtration rates (e.g. Scott and van Wyk, 1990; Cerdà, 1998; Robichaud, 2000; Martin and Moody, 2001). Beyond the reduction in peak infiltration rate, there is mounting evidence that the fundamental physics of infiltration in wild fire-affected soils is different from unburned soils (e.g. Imeson et al. , 1992; Moody et al. , 2009; Moody and Ebel, 2012). Understanding post-wildfire hydrology is critical given the increasing wildfire incidence in the western USA (Westerling et al. , 2006) and elsewhere in the world (Kasischke and Turetsky, 2006; Holz and Veblen, 2011; Pausas and Fernández-Muñoz, 2012). Wildfire is a disturbance event with global distribution (Bowman et al. , 2009; Krawchuk et al. , 2009; Pechony and Shindell, 2010; Moritz et al. , 2012), and with increasing populations moving into fire-prone areas, understanding post-wildfire infiltration is of increasing importance for predicting post-wildfire consequences. Runoff is generally controlled by the infiltration-excess mechanism in fire-affected soils (e.g. Mayor et al ., 2007; Onda et al. , 2008; Kinner and Moody, 2010). It is essential that the fire community have conceptual models, physical equations and tools (i.e. numerical models) to predict infiltration and thus excess rainfall (after Horton, 1933), which can provide estimates of peak discharge, start of runoff, time to peak and total runoff for hydroclimatic scenarios after wildfires. Reductions in saturated hydraulic conductivity K sat [LT -1 ] are common for fire-affected soils, and the relatively low values observed explain the elevated flash flood hazards (e.g. K sat of 1–100 mm h -1 , Robichaud, 2000; Yates et al. , 2000; Martin and Moody, 2001; Robichaud et al. , 2007; Moody et al. , 2009; Neary, 2011; Nyman et al. , 2011).

Hydrological Processes↗

Power source, data retrieval method, and attachment type affect success of dorsally mounted tracking tag deployments in 37 species of shorebirds

Animal-borne trackers are commonly used to study bird movements, including in long-distance migrants such as shorebirds. Selecting a tracker and attachment method can be daunting, and methodological advancements often have been made by trial and error and conveyed by word of mouth. We synthesized tracking outcomes across 2745 dorsally mounted trackers on 37 shorebird species around the world. We evaluated how attachment method, power source, data retrieval method, relative tracker mass, and biological traits affected success, where success was defined as whether or not each tag deployment reached its expected tracking duration (i.e. all aspects succeeded for the intended duration of the study: attachment, tracking, data acquisition, and bird survival). We conducted separate analyses for tag deployments with remote data retrieval (‘remote-upload tag deployments') and those that archived data and had to be recovered (‘archival tag deployments'). Among remote-upload tag deployments, those that were a lighter mass relative to the bird, were beyond their first year of production, transmitted data via satellite, or were attached with a leg-loop harness were most often successful at reaching their expected tracking duration. Archival tag deployments were most successful when applied at breeding areas, or when applied to males in any season. Remote-upload tag deployments with solar power, satellite data retrieval, or leg-loop harnesses continued tracking for longer than those with battery power, other types of data retrieval, or glue attachments. However, the majority of tag deployments failed to reach their expected tracking duration (71% of remote-upload, 83% of archival), which could have been due to tracker failure, attachment failure, or bird mortality. Our findings highlight that many tag deployments may fail to meet the goals of a study if tracking duration is crucial. Using our results, we provide guidelines for selecting a tracker and attachment to improve success at meeting study goals.

Journal of Avian Biology↗

The LTAR Grazing Land Common Experiment at the Jornada Experimental Range: Old genetics, new precision technologies, and adaptive value chains

Rangelands and the supply chains connected to them are central to the agrifood systems of the Southwestern United States. Local ranchers are simultaneously arid lands managers, herd managers, and marketing managers. To stay in business, they must constantly adapt to unpredictable forage resources and markets while conserving soils and vegetation resources for the long term. As climate warming and drying exacerbate the complexity and difficulty of day-to-day production, producers and policy-makers are seeking alternatives to “business as usual”. To meet this need, the LTAR-Jornada team has developed a package of strategies to help producers adapt to the local and inter-regional challenges. The package includes heritage cattle, precision ranching systems, and adaptive value chains. Five ranches across the Southwest have adopted different combinations of the strategies and are partnering to measure their benefits and drawbacks in real-world conditions. Opportunities for controlled experimentation differ among the ranches, so we use LTAR’s indicator system to assess and compare results. Even as we invest in co-producing knowledge about these three strategies, we recognize that progressive aridification and urbanization of Southwestern rangelands create challenges for which a single “silver bullet” agricultural innovation is unlikely to provide durable solutions. We are learning from our customers and stakeholders about ways to adjust the development of new options.

New Mexico↗

Assessing spring direct mortality to avifauna from wind energy facilities in the Dakotas

T he Northern Great Plains (NGP) contains much of the remaining temperate grasslands, an ecosystem that is one of the most converted and least protected in the world. Within the NGP, the Prairie Pothole Region (PPR) provides important habitat for >50% of North America's breeding waterfowl and many species of shorebirds, waterbirds, and grassland songbirds. This region also has high wind energy potential, but the effects of wind energy developments on migratory and resident bird and bat populations in the NGP remains understudied. This is troubling considering >2,200 wind turbines are actively generating power in the region and numerous wind energy projects have been proposed for development in the future. Our objectives were to estimate avian and bat fatality rates for wind turbines situated in cropland- and grassland-dominated landscapes, document species at high risk to direct mortality, and assess the influence of habitat variables on waterfowl mortality at 2 wind farms in the NGP. From 10 March to 7 June 2013–2014, we completed 2,398 searches around turbines for carcasses at the Tatanka Wind Farm (TAWF) and the Edgeley-Kulm Wind Farm (EKWF) in South Dakota and North Dakota. During spring, we found 92 turbine-related mortalities comprising 33 species and documented a greater diversity of species ( n  = 30) killed at TAWF (predominately grassland) than at EKWF ( n  = 9; predominately agricultural fields). After accounting for detection rates, we estimated spring mortality of 1.86 (SE = 0.22) deaths/megawatt (MW) at TAWF and 2.55 (SE = 0.51) deaths/MW at EKWF. Waterfowl spring (Mar–Jun) fatality rates were 0.79 (SE = 0.11) and 0.91 (SE = 0.10) deaths/MW at TAWF and EKWF, respectively. Our results suggest that future wind facility siting decisions consider avoiding grassland habitats and locate turbines in pre-existing fragmented and converted habitat outside of high densities of breeding waterfowl and major migration corridors.

North Dakota, South Dakota↗

Increased scientific rigor will improve reliability of research and effectiveness of management

Rigorous science that produces reliable knowledge is critical to wildlife management because it increases accurate understanding of the natural world and informs management decisions effectively. Application of a rigorous scientific method based on hypothesis testing minimizes unreliable knowledge produced by research. To evaluate the prevalence of scientific rigor in wildlife research, we examined 24 issues of the Journal of Wildlife Management from August 2013 through July 2016. We found 43.9% of studies did not state or imply a priori hypotheses, which are necessary to produce reliable knowledge. We posit that this is due, at least in part, to a lack of common understanding of what rigorous science entails, how it produces more reliable knowledge than other forms of interpreting observations, and how research should be designed to maximize inferential strength and usefulness of application. Current primary literature does not provide succinct explanations of the logic behind a rigorous scientific method or readily applicable guidance for employing it, particularly in wildlife biology; we therefore synthesized an overview of the history, philosophy, and logic that define scientific rigor for biological studies. A rigorous scientific method includes 1) generating a research question from theory and prior observations, 2) developing hypotheses (i.e., plausible biological answers to the question), 3) formulating predictions (i.e., facts that must be true if the hypothesis is true), 4) designing and implementing research to collect data potentially consistent with predictions, 5) evaluating whether predictions are consistent with collected data, and 6) drawing inferences based on the evaluation. Explicitly testing a priori hypotheses reduces overall uncertainty by reducing the number of plausible biological explanations to only those that are logically well supported. Such research also draws inferences that are robust to idiosyncratic observations and unavoidable human biases. Offering only post hoc interpretations of statistical patterns (i.e., a posteriori hypotheses) adds to uncertainty because it increases the number of plausible biological explanations without determining which have the greatest support. Further, post hoc interpretations are strongly subject to human biases. Testing hypotheses maximizes the credibility of research findings, makes the strongest contributions to theory and management, and improves reproducibility of research. Management decisions based on rigorous research are most likely to result in effective conservation of wildlife resources.

Journal of Wildlife Management↗

Identifying holes in the greater sage-grouse conservation umbrella

The umbrella species concept, wherein multiple species are indirectly protected under the umbrella of a reserve created for one, is intended to enhance conservation efficiency. Although appealing in theory and common in practice, empirical tests of the concept have been scarce. We used a real-world, semi-protected reserve established to protect a high-profile umbrella species (greater sage-grouse [ Centrocercus urophasianus ]) to investigate 2 potential mechanisms underlying the concept's successful application: reserve size and species similarity. We estimated how much habitat protection the established reserve provided to 52 species of conservation concern associated with vegetation communities where greater sage-grouse occur. To illustrate the importance of reserve size, we compared the effectiveness of the established reserve to alternative greater sage-grouse reserves of various sizes and to simulated reserves of equal size but sited with no regard for greater sage-grouse. We further assessed whether key species’ traits were associated with different levels of protection under the umbrella reserve. The established umbrella reserve protected 82% of the state's greater sage-grouse population and 0–63% of the habitat of the background species examined. The reserve outperformed equally sized, simulated reserves for only 12 of 52 background species. As expected, larger alternative reserves served as better umbrellas, but regardless of reserve size, not all species received equal protection. The established reserve was most effective at protecting the habitat of species that were most similar to the umbrella species (i.e., avian species, those highly associated with sagebrush plant communities, and those with widespread habitat). In contrast, the habitat of species with restricted distributions, particularly when combined with vegetation associations not closely matching the umbrella species, was not protected as well by the umbrella reserve. Such species require additional, targeted attention to achieve conservation objectives. Successful application of the umbrella species concept requires careful consideration of the characteristics of the umbrella species, the reserve delineated on its behalf, and the similarity of the umbrella species to its purported background species.

Wyoming↗

Demographic response of brown treesnakes to extended population suppression

From a management perspective, reptiles are relatively novel invasive taxa. Few methods for reptile control have been developed and very little is known about their effectiveness for reducing reptile populations, particularly when the goal is eradication. Many reptiles, and especially snakes, are cryptic, secretive, and undergo extended periods of inactivity, traits that decrease detection probabilities and create challenges in estimating population size or evaluating management effects. The brown treesnake ( Boiga irregularis ) is a notorious invasive species that continues to cause major ecological and economic harm following their introduction to the island of Guam after World War II. They have been the subject of intensive research on the effectiveness of various techniques to control snakes, including the first ever aerial system for the distribution of toxic acetaminophen baits for reptile control. We provide a cohort-based life table for a cryptic and invasive reptile undergoing extended population control using toxic baits from March 2017–2020. We also evaluated the effects of single (toxic bait) versus multi-tool (toxic bait and live trapping) management efforts on population trajectories, and estimated which population vital rates are most important for influencing population growth or decline in a treated landscape. Treatment of the population with acetaminophen-laced baits resulted in an immediate reduction followed by a gradual population decline that suggested that eradication was the probable outcome given sufficient treatment time but that the period of treatment was decades in magnitude. Inclusion of live trapping reduced the predicted time required to achieve eradication by more than half. Preventing the transition of 1,000-mm snout-vent length (SVL) females to larger sizes was predicted to have the greatest effect on population reduction based on integral projection modeling. Our results suggest that toxic baits are capable of eradicating brown treesnakes in an enclosure, although inclusion of trapping reduced overall treatment time required. Tools that effectively target females >1,000 mm SVL may have the greatest effect on reducing overall treatment timelines.

Journal of Wildlife Management↗

Assessing the sustainability of Pacific walrus harvest in a changing environment

Harvest sustainability is a primary goal of wildlife management and conservation, and in a changing world, it is increasingly important to consider environmental drivers of population dynamics alongside harvest in cohesive management plans. This is particularly pertinent for harvested species that acutely experience effects of climate change. The Pacific walrus ( Odobenus rosmarus divergens ), a crucial subsistence resource for Indigenous communities, is simultaneously subject to rapid habitat loss associated with diminishing sea ice and an increasing anthropogenic footprint in the Arctic. We developed a theta-logistic population modeling-management framework to evaluate various harvest scenarios combined with 4 potential climate and disturbance scenarios (ranging from optimistic to pessimistic, based largely on sea ice projections from general circulation models) to simulate Pacific walrus population dynamics to the end of the twenty-first century, focusing on the independent-aged female subset of the population. We considered 2 types of harvest strategies: 1) state-dependent harvest scenarios wherein we calculated harvest as a percentage of the population and updated annual harvests at set intervals as the population was reassessed, and 2) annually consistent harvest scenarios wherein annual harvest levels remain consistent into the future. All climate and disturbance scenarios indicated declines of varying severity in Pacific walrus abundance to the end of the twenty-first century, even in the absence of harvest. However, we found that a state-dependent annual harvest of 1.23% of the independent-aged female subset of the population (e.g., 1,280 independent-aged females harvested in 2020, similar to contemporary harvest levels) met our criterion for sustainability under all climate and disturbance scenarios, considering a medium risk tolerance level of 25%. This indicates that the present rate of Pacific walrus harvest is sustainable and will continue to be—provided the population is assessed at regular intervals and harvest is adapted to match changes in population dynamics. Our simulations indicate that a sustainable annually-consistent harvest is also possible but only at low levels if the population declines as expected. Applying a constant annual harvest of 1,280 independent-aged females failed to meet our criterion for sustainability under 3 of the 4 climate and disturbance scenarios we evaluated and had a higher probability of quasi-extinction than an equivalent state-dependent harvest scenario (1.23%). We highlight the importance of state-dependent management strategies and suggest our modeling framework is useful for managing harvest sustainability in a changing climate.

Journal of Wildlife Management↗

Intercolony variation in growth of black brant goslings on the Yukon-Kuskokwim Delta, Alaska

Recent declines in black brant ( Branta bernicla nigricans ) are likely the result of low recruitment. In geese, recruitment is strongly affected by habitat conditions experienced by broods because gosling growth rates are indicative of forage conditions during brood rearing and strongly influence future survival and productivity. In 2006–2008, we studied gosling growth at 3 of the 4 major colonies on the Yukon‐Kuskokwim Delta, Alaska. Estimates of age‐adjusted gosling mass at the 2 southern colonies (approx. 30% of the world population of breeding black brant) was low (gosling mass at 30.5 days ranged 346.7 ± 42.5 g to 627.1 ± 15.9 g) in comparison to a third colony (gosling mass at 30.5 days ranged 640.0 ± 8.3 g to 821.6 ± 13.6 g) and to most previous estimates of age‐adjusted mass of brant goslings. Thus, our results are consistent with the hypothesis that poor gosling growth is negatively influencing the brant population. There are 2 non‐mutually exclusive explanations for the apparent growth rates we observed. First, the population decline may have been caused by density‐independent factors and habitat capacity has declined along with the population as a consequence of the unique foraging feedback between brant and their grazing habitats. Alternatively, a reduction in habitat capacity, as a result of changes to the grazing system, may have negatively influenced gosling growth, which is contributing to the overall long‐term population decline. We found support for both explanations. For colonies over habitat capacity we recommend management to enhance foraging habitat, whereas for colonies below habitat capacity we recommend management to increase nesting productivity.

Journal of Wildlife Management↗

Incorporating harvest rates into the sex-age-kill model for white-tailed deer

Although monitoring population trends is an essential component of game species management, wildlife managers rarely have complete counts of abundance. Often, they rely on population models to monitor population trends. As imperfect representations of real-world populations, models must be rigorously evaluated to be applied appropriately. Previous research has evaluated population models for white-tailed deer ( Odocoileus virginianus ); however, the precision and reliability of these models when tested against empirical measures of variability and bias largely is untested. We were able to statistically evaluate the Pennsylvania sex-age-kill (PASAK) population model using realistic error measured using data from 1,131 radiocollared white-tailed deer in Pennsylvania from 2002 to 2008. We used these data and harvest data (number killed, age-sex structure, etc.) to estimate precision of abundance estimates, identify the most efficient harvest data collection with respect to precision of parameter estimates, and evaluate PASAK model robustness to violation of assumptions. Median coefficient of variation (CV) estimates by Wildlife Management Unit, 13.2% in the most recent year, were slightly above benchmarks recommended for managing game species populations. Doubling reporting rates by hunters or doubling the number of deer checked by personnel in the field reduced median CVs to recommended levels. The PASAK model was robust to errors in estimates for adult male harvest rates but was sensitive to errors in subadult male harvest rates, especially in populations with lower harvest rates. In particular, an error in subadult (1.5-yr-old) male harvest rates resulted in the opposite error in subadult male, adult female, and juvenile population estimates. Also, evidence of a greater harvest probability for subadult female deer when compared with adult (≥2.5-yr-old) female deer resulted in a 9.5% underestimate of the population using the PASAK model. Because obtaining appropriate sample sizes, by management unit, to estimate harvest rate parameters each year may be too expensive, assumptions of constant annual harvest rates may be necessary. However, if changes in harvest regulations or hunter behavior influence subadult male harvest rates, the PASAK model could provide an unreliable index to population changes.

Journal of Wildlife Management↗

Trap style influences wild pig behavior and trapping success

Despite the efforts of many natural resource professionals, wild pig ( Sus scrofa ) populations are expanding in many areas of the world. Although many creative techniques for controlling pig populations are being explored, trapping has been and still is the most commonly used method of population control for many public and private land managers. We conducted an observational study to examine the efficiency of 2 frequently used trap styles: a small, portable box‐style trap and a larger, semi‐permanent, corral‐style trap. We used game cameras to examine patterns of trap entry by wild pigs around each style of trap, and we conducted a trapping session to compare trapping success between trap styles. Adult female and juvenile wild pigs entered both styles of trap more readily than did adult males, and adult males seemed particularly averse to entering box traps. Less than 10% of adult male visits to box traps resulted in entries, easily the least percentage of any class at any style of trap. Adult females entered corral traps approximately 2.2 times more often per visit than box traps and re‐entered corral traps >2 times more frequently. Juveniles entered and re‐entered both box and corral traps at similar rates. Overall (all‐class) entry‐per‐visit rates at corral traps (0.71) were nearly double that of box traps (0.37). Subsequent trapping data supported these preliminary entry data; the capture rate for corral traps was >4 times that of box traps. Our data suggest that corral traps are temporally and economically superior to box traps with respect to efficiency; that is, corral traps effectively trap more pigs per trap night at a lower cost per pig than do box traps

Georgia↗

Increased flexibility for modeling telemetry and nest-survival data using the multistate framework

Although telemetry is one of the most common tools used in the study of wildlife, advances in the analysis of telemetry data have lagged compared to progress in the development of telemetry devices. We demonstrate how standard known-fate telemetry and related nest-survival data analysis models are special cases of the more general multistate framework. We present a short theoretical development, and 2 case examples regarding the American black duck and the mallard. We also present a more complex lynx data analysis. Although not necessary in all situations, the multistate framework provides additional flexibility to analyze telemetry data, which may help analysts and biologists better deal with the vagaries of real-world data collection.

Journal of Wildlife Management↗