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The North American Breeding Bird Survey 1966–2011: Summary analysis and species accounts

The North American Breeding Bird Survey is a roadside, count-based survey conducted by volunteer observers. Begun in 1966, it now is a primary source of information on spatial and temporal patterns of population change for North American birds. We analyze population change for states, provinces, Bird Conservation Regions, and the entire survey within the contiguous United States and southern Canada for 426 species using a hierarchical log-linear model that controls for observer effects in counting. We also map relative abundance and population change for each species using a spatial smoothing of data at the scale of survey routes. We present results in accounts that describe major breeding habitats, migratory status, conservation status, and population trends for each species at several geographic scales. We also present composite results for groups of species categorized by habitats and migratory status. The survey varies greatly among species in percentage of species' range covered and precision of results, but consistent patterns of decline occur among eastern forest, grassland, and aridland obligate birds while generalist bird species are increasing.

North American Fauna

A hierarchical model for estimating change in American Woodcock populations

The Singing-Ground Survey (SGS) is a primary source of information on population change for American woodcock (Scolopax minor). We analyzed the SGS using a hierarchical log-linear model and compared the estimates of change and annual indices of abundance to a route regression analysis of SGS data. We also grouped SGS routes into Bird Conservation Regions (BCRs) and estimated population change and annual indices using BCRs within states and provinces as strata. Based on the hierarchical model?based estimates, we concluded that woodcock populations were declining in North America between 1968 and 2006 (trend = -0.9%/yr, 95% credible interval: -1.2, -0.5). Singing-Ground Survey results are generally similar between analytical approaches, but the hierarchical model has several important advantages over the route regression. Hierarchical models better accommodate changes in survey efficiency over time and space by treating strata, years, and observers as random effects in the context of a log-linear model, providing trend estimates that are derived directly from the annual indices. We also conducted a hierarchical model analysis of woodcock data from the Christmas Bird Count and the North American Breeding Bird Survey. All surveys showed general consistency in patterns of population change, but the SGS had the shortest credible intervals. We suggest that population management and conservation planning for woodcock involving interpretation of the SGS use estimates provided by the hierarchical model.

Journal of Wildlife Management

Modeling wetland blackbird populations as a function of waterfowl abundance in the prairie pothole region of the United States and Canada

Blackbirds share wetland habitat with many waterfowl species in Bird Conservation Region 11 (BCR 11), the prairie potholes. Because of similar habitat preferences, there may be associations between blackbird populations and populations of one or more species of waterfowl in BCR11. This study models populations of red-winged blackbirds and yellow-headed blackbirds as a function of multiple waterfowl species using data from the North American Breeding Bird Survey within BCR11. For each blackbird species, we created a global model with blackbird abundance modeled as a function of 11 waterfowl species; nuisance effects (year, route, and observer) also were included in the model. Hierarchical Poisson regression models were fit using Markov chain Monte Carlo methods in WinBUGS 1.4.1. Waterfowl abundances were weakly associated with blackbird numbers, and no single waterfowl species showed a strong correlation with any blackbird species. These findings suggest waterfowl abundance from a single species is not likely a good bioindicator of blackbird abundance; however, a global model provided good fit for predicting red-winged blackbird abundance. Increased model complexity may be required for accurate predictions of blackbird abundance; the amount of data required to construct appropriate models may limit this approach for predicting blackbird abundance in the prairie potholes. Copyright ?? Taylor & Francis Group, LLC.

Environmental Bioindicators

A review of the population estimation approach of the North American landbird conservation plan

As part of their development of a continental plan for monitoring landbirds (Rich et al. 2004), Partners in Flight (PIF) applied a new method to make preliminary estimates of population size for all 448 species of landbirds present in the continental United States and Canada (Table 1). Estimation of the global population size of North American landbirds was intended to (1) identify the degree of vulnerability of each species, (2) provide estimates of the current population size for each species, and (3) provide a starting point for estimating population sizes in states, provinces, territories, and Bird Conservation Regions (Rich et al. 2004). A method proposed by Rosenberg and Blancher (2005) was used to derive population estimates from available survey data. To enhance the credibility of these estimates, PIF organized a review of the methodology used to estimate North American landbird population sizes. A planning committee selected members from the ornithological and biometrical communities (hereafter “the panel”), with the aim of selecting individuals from academia, state natural-resource agencies, and the U.S. and Canadian federal governments, including the Canadian Wildlife Service, the U.S. Geological Survey, and the U.S. Department of Agriculture Forest Service. The panel addressed three questions: (1) Were the methods of population estimation proposed by PIF reasonable? (2) What actions could be taken to improve the data or analyses on which the PIF population estimates were based? and (3) How should the PIF population estimates be interpreted?

The Auk

Quantifying avian resilience to habitat change to support conservation decision-making

Habitat loss and degradation are two of the main drivers of contemporary avian population declines. Wildlife managers are increasingly advocating for tools that provide decision support to set priorities for restoration or conservation efficiently. We demonstrate how to derive two species-specific management and resilience metrics: the greatest management impact point (GMIP) and the ecological resilience threshold (RT). The GMIP indicates the amount of environmental change in a landscape where habitat improvements are expected to have the greatest impact on species’ occurrence. The RT represents the amount of environmental change in a landscape that a species can tolerate before the steepest change in occupancy is expected to begin. We estimate species-specific metrics using the amount of uncharacteristic exotic vegetation as an index of environmental change, and demonstrate how multispecies patterns may suggest potential management strategies. We estimated occupancy models using 5 years of multispecies avian detection data from the Integrated Monitoring in Bird Conservation Regions program in the State of Utah, USA (hereafter, Utah). Based on the estimated relationship between species’ occupancy and amounts of uncharacteristic exotic vegetation, we derive RT and GMIP scores for 61 species breeding in Utah. We found wide interspecific variation in resilience to amounts of exotic vegetation, with species generally clustering at extreme values. Our results demonstrate that birds in Utah appear more resilient to amounts of uncharacteristic exotic vegetation at coarser spatial resolution, showing greater variance and lower average RTs at finer spatial resolution. Species that are not fully resilient to the range of uncharacteristic exotic vegetation observed in this study are expected to respond most strongly, on average, to management actions in landscapes with high levels of exotic vegetation; however, early detection and rapid response is likely the most effective strategy. Quantified across many species, these metrics can be used to identify and prioritize landscapes where current environmental conditions could be maintained to avoid the greatest species’ declines, or which maximize expected biodiversity returns on investment in environmental restoration. Managers can consider either focal species’ resilience for tailored conservation planning or summarize species resilience to create efficient management plans that maximize outcomes for multiple species.

Utah

Evaluating outcomes of management targeting the recovery of a migratory songbird of conservation concern

Assessing outcomes of habitat management is critical for informing and adapting conservation plans. From 2013 – 2019, a multi-stage management initiative aims to create >25,000 ha of shrubland and early-successional vegetation to benefit Golden-winged Warblers (Vermivora chrysoptera) in managed forested landscapes of the western Great Lakes region. We studied a dense breeding population of Golden-winged Warblers at Rice Lake National Wildlife Refuge (NWR) in Minnesota, USA, where shrubs and young trees were sheared during the winter of 2014-2015 in a single treatment supported in part by the American Bird Conservancy (ABC) and in part by other funding source(s) to benefit Golden-winged Warblers and other species associated with young forest [e.g., American Woodcock (Scalopax minor)] and as part of maintenance of early successional forest cover on the refuge.

Minnesota

Use of NEXRAD to study shorebird migration in the Prairie Pothole region: A feasibility study

An essential component of shorebird conservation is identifying, protecting, and managing high-priority stopover sites and migration habitats crucial to the long-term persistence of migrating shorebirds. Because of the tremendous variability in migrant shorebird occurrence patterns in the Prairie Pothole Region of the U.S. (Skagen 1997), it is labor- and cost-intensive to locate the majority of sites used heavily by shorebirds in any one migration period. Because WSR-88D (Weather Surveillance Radar – 1988 Doppler) or NEXRAD (NEXt generation weather RADar) has been useful for locating migrating birds and revealing migration patterns and important roosting sites of some species (e.g., Diehl and others 2003, Gauthreaux and Belser 2003), we undertook a pilot field study to determine wheTHER it also might be feasible to use NEXRAD for locating important stopover sites used by migrating shorebirds in the prairie potholes landscape. Coordinated efforts to advance the applicability of radar technology to bird conservation are underway (Ruth and others 2005).

Open-File Report

Regional conservation planning tool: A spreadsheet model to support spatial prioritization and resource allocation decisions

Prioritization is a central component of natural resource management because conservation needs routinely exceed available resources. Waterfowl and wetland conservation programs in North America are at the forefront of landscape-scale prioritization and transboundary management decisions due to the migratory nature of ducks, geese, and swans. The growing availability of geographic information systems (GIS) and geospatial technologies has accelerated the development of multi-objective landscape prioritization models, including applications of structured decision making and multi-criteria decision analysis to spatial planning for waterfowl and wetlands at the continental scale. However, regional managers and conservationists could benefit from flexibility in downscaling continental tools, selecting objectives, and assigning weights for rapid production of spatial prioritization models at smaller spatial scales without extensive computer coding or GIS analysis. We developed a spatial value model that prioritizes landscapes at sub-continental scales (e.g., states and provinces, bird conservation regions, etc.) and provides flexibility for users to select waterfowl conservation objectives of interest and weights. Our model can be used for direct downscaling of an existing continental geospatial model or further customized with region-specific geospatial data. We illustrate how regional prioritization can vary with the spatial scale selected by the user. The spatial value modeling framework and the downscaling tool presented here could increase the use of multi-criteria decision analysis and linear value modeling in spatial landscape prioritization, while also providing flexibility for selecting scales, objectives, and weights. Our spreadsheet tool was developed specifically for use by regional biologists, conservationists, and managers and does not require knowledge of GIS software (although results can be exported from the spreadsheet for spatial analysis using GIS). Together, the model outputs and the accompanying spreadsheet tool provide a bridge between continental waterfowl conservation and regional implementation, enabling rapid, stakeholder-driven, value-explicit prioritization.

Wildlife Society Bulletin

Population trends for North American winter birds based on hierarchical models

Managing widespread and persistent threats to birds requires knowledge of population dynamics at large spatial and temporal scales. For over 100 yrs, the Audubon Christmas Bird Count (CBC) has enlisted volunteers in bird monitoring efforts that span the Americas, especially southern Canada and the United States. We employed a Bayesian hierarchical model to control for variation in survey effort among CBC circles and, using CBC data from 1966 to 2013, generated early-winter population trend estimates for 551 species of birds. Selecting a subset of species that do not frequent bird feeders and have ≥25% range overlap with the distribution of CBC circles (228 species) we further estimated aggregate (i.e., across species) trends for the entire study region and at the level of states/provinces, Bird Conservation Regions, and Landscape Conservation Cooperatives. Moreover, we examined the relationship between ten biological traits—range size, population size, migratory strategy, habitat affiliation, body size, diet, number of eggs per clutch, age at sexual maturity, lifespan, and tolerance of urban/suburban settings—and CBC trend estimates. Our results indicate that 68% of the 551 species had increasing trends within the study area over the interval 1966–2013. When trends were examined across the subset of 228 species, the median population trend for the group was 0.9% per year at the continental level. At the regional level, aggregate trends were positive in all but a few areas. Negative population trends were evident in lower latitudes, whereas the largest increases were at higher latitudes, a pattern consistent with range shifts due to climate change. Nine of 10 biological traits were significantly associated with median population trend; however, none of the traits explained >34% of the deviance in the data, reflecting the indirect relationships between population trend estimates and species traits. Trend estimates based on the CBC are broadly congruent with estimates based on the North American Breeding Bird Survey, another large-scale monitoring program. Both of these efforts, conducted by citizen scientists, will be required going forward to ensure robust inference about population dynamics in the face of climate and land cover changes.

Ecosphere

An integrated population model for bird monitoring in North America

Integrated population models (IPMs) provide a unified framework for simultaneously analyzing data sets of different types to estimate vital rates, population size, and dynamics; assess contributions of demographic parameters to population changes; and assess population viability. Strengths of an IPM include the ability to estimate latent parameters and improve the precision of parameter estimates. We present a hierarchical IPM that combines two broad‐scale avian monitoring data sets: count data from the North American Breeding Bird Survey (BBS) and capture–recapture data from the Monitoring Avian Productivity and Survivorship (MAPS) program. These data sets are characterized by large numbers of sample sites and observers, factors capable of inducing error in the sampling and observation processes. The IPM integrates the data sets by modeling the population abundance as a first‐order autoregressive function of the previous year's population abundance and vital rates. BBS counts were modeled as a log‐linear function of the annual index of population abundance, observation effects (observer identity and first survey year), and overdispersion. Vital rates modeled included adult apparent survival, estimated from a transient Cormack‐Jolly‐Seber model using MAPS data, and recruitment (surviving hatched birds from the previous season + dispersing adults) estimated as a latent parameter. An assessment of the IPM demonstrated it could recover true parameter values from 200 simulated data sets. The IPM was applied to data sets (1992–2008) of two bird species, Gray Catbird ( Dumetella carolinensis ) and Wood Thrush ( Hylocichla mustelina ) in the New England/Mid‐Atlantic coastal Bird Conservation Region of the United States. The Gray Catbird population was relatively stable (trend +0.4% per yr), while the Wood Thrush population nearly halved (trend −4.5% per yr) over the 17‐yr study period. IPM estimates of population growth rates, adult survival, and detection and residency probabilities were similar and as precise as estimates from the stand‐alone BBS and CJS models. A benefit of using the IPM was its ability to estimate the latent recruitment parameter. Annual growth rates for both species correlated more with recruitment than survival, and the relationship for Wood Thrush was stronger than for Gray Catbird. The IPM's unified modeling framework facilitates integration of these important data sets.

Ecological Applications

American woodcock migratory connectivity as indicated by hydrogen isotopes

To identify factors contributing to the long-term decline of American woodcock, a holistic understanding of range-wide population connectivity throughout the annual cycle is needed. We used band recovery data and isotopic composition of primary (P1) and secondary (S13) feathers to estimate population sources and connectivity among natal, early fall, and winter ranges of hunter-harvested juvenile American woodcock. We used P1 feathers from known-origin pre-fledged woodcock ( n  = 43) to create a hydrogen δ 2 H f isoscape by regressing δ 2 H f against expected growing-season precipitation ( δ 2 H p ). Modeled δ 2 H p values explained 79% of the variance in P1 δ 2 H f values, indicating good model fit for estimating woodcock natal origins. However, a poor relationship ( r 2  = 0.23) between known-origin, S13 δ 2 H f values, and expected δ 2 H p values precluded assignment of early fall origins. We applied the δ 2 H f isoscape to assign natal origins using P1 feathers from 494 hunter-harvested juvenile woodcock in the United States and Canada during 2010–2011 and 2011–2012 hunting seasons. Overall, 64% of all woodcock origins were assigned to the northernmost (>44°N) portion of both the Central and Eastern Management Regions. In the Eastern Region, assignments were more uniformly distributed along the Atlantic coast, whereas in the Central Region, most woodcock were assigned to origins within and north of the Great Lakes region. We compared our origin assignments to spatial coverage of the annual American woodcock Singing Ground Survey (SGS) and evaluated whether the survey effectively encompasses the entire breeding range. When we removed the inadequately surveyed Softwood shield Bird Conservation Region (BCR) from the northern portion of the SGS area, only 48% of juvenile woodcock originated in areas currently surveyed by the SGS. Of the individuals assigned to the northernmost portions of the breeding range, several were harvested in the southern extent of the wintering range. Based upon this latitudinal winter stratification, we examined whether woodcock employed a leapfrog migration strategy. Using δ 2 H f values and band-recovery data, we found some support for this migration strategy hypothesis but not as a singular explanation. The large harvest derivation of individuals from the northernmost portions of the breeding range, and the difference in breeding distributions within each Management Region should be considered in future range-wide conservation and harvest management planning for American woodcock.

Journal of Wildlife Management

Allowable take of black vultures in the eastern United States

Black vultures ( Coragyps atratus ) have been increasing in density and expanding their range in the eastern United States since at least the 1960s. In many areas, their densities have increased to the level where they are causing damage to property and livestock and the number of requests for allowable take permits has increased throughout these areas. The United States Fish and Wildlife Service (USFWS) requires updated information to help inform the number of take permits that could reduce conflicts while meeting obligations under the Migratory Bird Treaty Act. We expanded analyses used to estimate allowable take in Virginia to cover the range of black vultures in the eastern United States. We used the prescribed take level approach, which integrates demographic rates, population size estimates, and management objectives into an estimate of allowable take. We provide estimates of allowable take at 4 different scales: individual states, Bird Conservation Regions, USFWS administrative regions, and flyways. Our updated population time series provides evidence of rapidly increasing black vulture populations in many regions of the eastern United States, with an overall population estimate of approximately 4.26 million in 2015 in the Atlantic and Mississippi Flyways. Estimated allowable take ranged from a few hundred individuals per year in states at the northern end of the species range to approximately 287,000/year over the entire eastern United States. The USFWS has no legal mandate regarding the spatial scale at which take should be managed and we found little biological evidence of subpopulation structure for black vultures in the eastern United States. We suggest that allowable take for the species be implemented at a scale that meets stakeholder objectives (e.g., reducing conflict, and ensuring that black vultures are not extirpated from local areas) and is efficient for administrative and monitoring purposes.

Journal of Wildlife Management

Combining waterfowl and breeding bird survey data to estimate wood duck breeding population size in the Atlantic Flyway

We combined data from the Atlantic Flyway Breeding Waterfowl Survey (AFBWS) and the North American Breeding Bird Survey (BBS) to estimate the number of wood ducks ( Aix sponsa ) in the United States portion of the Atlantic Flyway from 1993 to 2013. The AFBWS is a plot-based survey that covers most of the northern and central portions of the Flyway; when analyzed with adjustments for survey time of day effects, these data can be used to estimate population size. The BBS provides an index of wood duck abundance along roadside routes. Although factors influencing change in BBS counts over time can be controlled in BBS analysis, BBS indices alone cannot be used to derive population size estimates. We used AFBWS data to scale BBS indices for Bird Conservation Regions (BCR), basing the scaling factors on the ratio of estimated AFBWS population sizes to regional BBS indices for portions of BCRs that were common to both surveys. We summed scaled BBS results for portions of the Flyway not covered by the AFBWS with AFBWS population estimates to estimate a mean yearly total of 1,295,875 (mean 95% CI: 1,013,940–1,727,922) wood ducks. Scaling factors varied among BCRs from 16.7 to 148.0; the mean scaling factor was 68.9 (mean 95% CI: 53.5–90.9). Flyway-wide, population estimates from the combined analysis were consistent with alternative estimates derived from harvest data, and also provide population estimates within states and BCRs. We recommend their use in harvest and habitat management within the Atlantic Flyway.

Connecticut, Delaware, Maryland, Massachusetts, Ne

The future demographic niche of a declining grassland bird fails to shift poleward in response to climate change

Context Temperate grasslands and their dependent species are exposed to high variability in weather and climate due to the lack of natural buffers such as forests. Grassland birds are particularly vulnerable to this variability, yet have failed to shift poleward in response to recent climate change like other bird species in North America. However, there have been few studies examining the effect of weather on grassland bird demography and consequent influence of climate change on population persistence and distributional shifts. Objectives The goal of this study was to estimate the vulnerability of Henslow’s Sparrow ( Ammodramus henslowii ), an obligate grassland bird that has been declining throughout much of its range, to past and future climatic variability. Methods We conducted a demographic meta-analysis from published studies and quantified the relationship between nest success rates and variability in breeding season climate. We projected the climate-demography relationships spatially, throughout the breeding range, and temporally, from 1981 to 2050. These projections were used to evaluate population dynamics by implementing a spatially explicit population model. Results We uncovered a climate-demography linkage for Henslow’s Sparrow with summer precipitation, and to a lesser degree, temperature positively affecting nest success. We found that future climatic conditions—primarily changes in precipitation—will likely contribute to reduced population persistence and a southwestward range contraction. Conclusions Future distributional shifts in response to climate change may not always be poleward and assessing projected changes in precipitation is critical for grassland bird conservation and climate change adaptation.

Landscape Ecology

A hidden cost of single species management: Habitat-relationships reveal potential negative effects of conifer removal on a non-target species

Land management priorities and decisions may result in population declines for non-target wildlife species. In the western United States, large-scale removal of conifer from sagebrush ecosystems ( Artemisia spp.) is occurring to recover greater sage-grouse ( Centrocercus urophasianus ) populations and may result in pinyon jay ( Gymnorhinus cyanocephalus ) habitat loss. Jay populations have experienced long-term declines, due to unknown causes, resulting in a recent petition for listing under the Endangered Species Act of 1973. We developed a Bayesian hierarchical model of jay abundance, using 13 years of point count data (2008–2020) collected across the western United States, to estimate regional population trends, model habitat requirements, assess conifer removal effects on jays, and generate hypotheses regarding jay population declines. Our model included climate and landcover covariates and regional trends in pinyon jay density. We applied our modeled habitat relationships to map predicted pinyon jay density, given 2008 and 2020 resource conditions, and map density changes from 2008 to 2020. Our results indicate pinyon jay populations are declining within Bird Conservation Region 16. Jay density was positively associated with sagebrush cover, Palmer Drought Severity Index, and pinyon-juniper cover. Conversely, jay populations were negatively associated with Normalized Difference Vegetation Index (NDVI). We found higher pinyon jay densities within locations possessing both sagebrush and pinyon-juniper cover; conditions characteristic of phase I and II conifer encroachment which are preferentially targeted for conifer removal to restore sagebrush communities. Conifer removal, if conducted at locations with high pinyon jay densities, is therefore likely to negatively affect jay abundance.

Arizona, California, Colorado, Idaho, Kansas, Mont

Canada Warbler use of harvested stands following timber management in the southern portion of their range

We evaluated Canada Warbler ( Cardellina canadensis ) responses to changes in habitat characteristics (landscape metrics, landcover, and microhabitat features) at multiple spatial scales resulting from timber harvests (clear-cut, heavy partial, and light partial) between 1996 and 2009. Relative abundance of Canada Warblers decreased over time on our West Virginia study area (Wildlife and Ecosystem Research Forest) and within the Appalachian Bird Conservation Region. Initially, relative abundance was greater closer to roads, but as timber harvests became more common, relative abundance became positively associated with amount of light partial harvests at the local scale. Nest survival was 45.6 ± 18.3% during 1996–1998 and 24.9 ± 14.6% during 2007–2009, but did not differ ( P = 0.38) between these periods. Areas around nests in 2007–2009 ( n = 17) had less intermediate canopy cover and fewer residual trees but more green cover, woody debris, and pole trees than areas around nests in 1996–1998 ( n = 10). Successful nests had more low cover, less vertical diversity, more woody debris, more saplings, and greater edge density than failed nests. We found a positive association between relative abundance and all three types of timber harvests and an improvement in habitat through understory development and retention of residual trees. Our research finds preliminary support for use of timber harvests, particularly light partial harvests, as a management tool for Canada Warblers in the southern portion of their range with the need for extended research using treatments and controls to confirm successful management.

Forest Ecology and Management

At the end of the road: Lessons learned from comparing model- and design-based approaches to estimate population sizes of boreal birds in Alberta, Canada

Estimating population abundance is a challenging task complicated by the amount, type, and quality of available data. Conservationists have relied on design-based estimates from Partners in Flight (PIF), which primarily uses roadside data from the North American Breeding Bird Survey (BBS) to estimate populations sizes. However, the BBS was not designed to estimate population sizes. We developed models incorporating land cover and climate variables based on roadside and off-road point-count surveys. We calculated spatially explicit, model-based population estimates for 81 landbird species in Bird Conservation Region 6 in Alberta, Canada, and compared these to PIF estimates. We also developed a framework to evaluate how the differences between the detection distance, time-of-day, roadside count, and habitat representation adjustments explain discrepancies between the two estimators. We showed that the key assumptions of the PIF population size estimator were commonly violated in this region, and the two approaches provided very different population size estimates for most species. The average differences between estimators were explained by differences in the detection distance and time-of-day components, but these adjustments left much unexplained variation among species. Differences in the roadside count and habitat representation components explained most of the among-species variation. The variation caused by these factors was large enough to change the population size ranking of the species. The roadside count bias needs serious attention when roadside surveys are used to extrapolate over off-road areas. Habitat representation bias is likely prevalent in regions sparsely and non-representatively sampled by roadside surveys, such as the boreal region of North America, and thus population size estimates for these regions need to be treated with caution for certain species. Model-based integration of available data sources and additional sampling can contribute towards more accurate population size estimates for conservation in remote areas of North America.

Alberta

Despite regional variation, Gymnorhinus cyanocephalus (Pinyon Jay) densities generally increase with local pinyon–juniper cover and heterogeneous ground cover

Traditionally, local-scale habitat-relationship models are developed over small spatial extents, limiting model transferability and inference outside the study area. Thus, habitat managers frequently lack fine-scale information regarding the influence of vegetation composition and structure on site suitability or species abundance. Gymnorhinus cyanocephalus (Pinyon Jay) represents one declining species for which managers have limited information regarding the influence that vegetation composition and structure have on abundance at broad scales. To address this need, we developed a hierarchical Bayesian abundance model using summertime bird and vegetation data collected under the Integrated Monitoring in Bird Conservation Regions program to explain jay abundance as a function of local conditions. Our G. cyanocephalus abundance model allowed abundance relationships with pinyon pine ( Pinus edulis and P. monophylla ) and juniper ( Juniperus spp.) to vary by ecoregion, thereby accounting for potential regional differences in habitat associations. We found G. cyanocephalus abundance was generally positively associated with pinyon pine and juniper cover; however, habitat relationships varied by ecoregion. Additionally, we found positive associations between jay abundance and grass cover, sagebrush cover, and percent bare ground. Our results agree with prior research suggesting mechanical removal of pinyon pine and juniper trees for sagebrush restoration or fuel treatments may negatively affect G. cyanocephalus . Managers wishing to reduce pinyon and juniper tree cover without negatively affecting G. cyanocephalus may benefit from targeting sites where both large-scale distribution models and our local habitat relationships suggest G. cyanocephalus are likely to occur in low numbers. Additionally, our modeled relationships indicate restoration that increases grass cover, sagebrush cover, and bare ground, while maintaining pinyon and (or) juniper cover, may lead to increased local densities of G. cyanocephalus .

Ornithological Applications