Scope, setting, and purpose of the Colorado Front Range climate change vulnerability assessment for national forests
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I calculated avian conservation scores, based on published quantitative assessment of bird presence (typically relative abundance) and regional Partners in Flight conservation concern score of each species. I related these avian conservation scores to severity of forest perturbation (i.e., % retention) and number of years since perturbation for forest stands within North America. I characterized avian response to perturbation by comparing avian conservation score on perturbed stands with that of paired unperturbed reference stands. Positive avian response to perturbation, representative of improved avian conservation status, was found when forest retention was >20% of coniferous forests or 25%–75% of hardwood forests. Positive avian response was greatest ~10 years after perturbation that retained 58% of conifer forests or 49% of hardwood forests. Increased avian conservation scores following perturbation may last for up to 23 years after perturbation but occur and diminish sooner with greater forest retention. Average annual avian response, over 23 years post-perturbation, was greatest when 30%–50% of hardwood stands were retained and when 50%–70% of conifer stands were retained.
Wintering birds were captured with mist nets at 12 pairs of forested sites in the New World tropics in 1984 and 1985 to compare populations in small isolated woodlands (generally 5-50 ha) with those in extensive forests (> 1,000 ha). Net-hours of effort were similar in large and small sites, as were total birds captured and banded, but species composition was very different. Members of the Todidae, Dendrocolaptidae, Formicariidae and Thraupinae were significantly more common in extensive forest than in small isolated tracts, indicating that these birds are especially vulnerable to effects of forest fragmentation. However, in winter many species of North American migrants, even species that are restricted to extensive forest during the breeding season, were just as common in small forest fragments as in extensive forest. A high percentage of the North American migrants banded in January 1984 (40 to 50 % for some species) were recaptured in 1985.
Background Dry mixed-conifer forests of the southwestern United States are experiencing rapid, anthropogenically driven fire regime change. Prior to the Euro-American settlement, most of these forests experienced frequent surface fires but are now vulnerable to uncharacteristically large, high-severity fires. Fire directly influences the structure and composition of these forests and, in turn, the wildlife that inhabit them. Changing fire regimes result in a certain decline of some species and uncertain consequences for others. The Mexican spotted owl ( Strix occidentalis lucida ) is a federally listed threatened species of particular note in southwestern mixed-conifer forests. High-severity fire is cited as the owl’s primary threat in the revised species recovery plan, but uncertainties surround the impacts of high-severity fire on the habitat of the threatened owl, particularly across a timeframe longer than a few years. Our objective was to explore the long-term (100-year) effects of fire severity on elements of forest structure vital for Mexican spotted owl nesting. We quantified structural attributes for nest/roost habitat across mixed-conifer forests that burned at varying severity levels and time periods in the last century. We then examined the drivers of structural attributes by detecting statistical differences between severity classes and time periods through permutational multivariate analysis of variance. Results High-severity fire has the strongest deleterious impact on elements of forest structure (total basal area, percent medium tree basal area, percent large tree basal area, large tree density, and canopy cover) vital to Mexican spotted owl nesting, and although the structural differences between severity classes diminish with time, it took ≥ 80–100 years to reach the structural conditions desired for Mexican spotted owl nesting after stand-replacing fires. The most important attribute measured, canopy cover, required 90–100 years after high-severity fires to reach levels most suitable for Mexican spotted owls in the Lincoln National Forest. Conclusions As fires increase in frequency, severity, and size compared to the last century, the Lincoln National Forest is projected to face an overall decrease in the structural conditions needed for Mexican spotted owl nesting habitat in this region. Short intervals between uncharacteristically high-severity fires in particular pose an imminent threat to nesting habitat.
Synthesis of results from several Arctic and boreal research programmes provides evidence for the strong role of high-latitude ecosystems in the climate system. Average surface air temperature has increased 0.3??C per decade during the twentieth century in the western North American Arctic and boreal forest zones. Precipitation has also increased, but changes in soil moisture are uncertain. Disturbance rates have increased in the boreal forest; for example, there has been a doubling of the area burned in North America in the past 20 years. The disturbance regime in tundra may not have changed. Tundra has a 3-6-fold higher winter albedo than boreal forest, but summer albedo and energy partitioning differ more strongly among ecosystems within either tundra or boreal forest than between these two biomes. This indicates a need to improve our understanding of vegetation dynamics within, as well as between, biomes. If regional surface warming were to continue, changes in albedo and energy absorption would likely act as a positive feedback to regional warming due to earlier melting of snow and, over the long term, the northward movement of treeline. Surface drying and a change in dominance from mosses to vascular plants would also enhance sensible heat flux and regional warming in tundra. In the boreal forest of western North America, deciduous forests have twice the albedo of conifer forests in both winter and summer, 50-80% higher evapotranspiration, and therefore only 30-50% of the sensible heat flux of conifers in summer. Therefore, a warming-induced increase in fire frequency that increased the proportion of deciduous forests in the landscape, would act as a negative feedback to regional warming. Changes in thermokarst and the aerial extent of wetlands, lakes, and ponds would alter high-latitude methane flux. There is currently a wide discrepancy among estimates of the size and direction of CO2 flux between high-latitude ecosystems and the atmosphere. These discrepancies relate more strongly to the approach and assumptions for extrapolation than to inconsistencies in the underlying data. Inverse modelling from atmospheric CO2 concentrations suggests that high latitudes are neutral or net sinks for atmospheric CO2, whereas field measurements suggest that high latitudes are neutral or a net CO2 source. Both approaches rely on assumptions that are difficult to verify. The most parsimonious explanation of the available data is that drying in tundra and disturbance in boreal forest enhance CO2 efflux. Nevertheless, many areas of both tundra and boreal forests remain net sinks due to regional variation in climate and local variation in topographically determined soil moisture. Improved understanding of the role of high-latitude ecosystems in the climate system requires a concerted research effort that focuses on geographical variation in the processes controlling land-atmosphere exchange, species composition, and ecosystem structure. Future studies must be conducted over a long enough time-period to detect and quantify ecosystem feedbacks.
Restoring forest ecosystems has become an increasingly high priority for land managers across the American West. Millions of hectares of forest are in need of drastic yet strategic reductions in density (e.g., basal area). Meeting the restoration and management goals requires quantifying metrics of vertical and horizontal forest structure, which has relied upon field‐based measurements, manned airborne or satellite remote sensing datasets. We used unmanned aerial vehicle (UAV ) image‐derived Structure‐from‐Motion (SfM) models and high‐resolution multispectral orthoimagery in this study to quantify vertical and horizontal forest structure at both the fine‐ (<4 ha) and mid‐scales (4–400 ha) across a forest density gradient. We then used these forest structure estimates to assess specific objectives of a forest restoration treatment. At the fine‐scale, we found that estimates of individual tree height and canopy diameter were most accurate in low‐density conditions, with accuracies degrading significantly in high‐density conditions. Mid‐scale estimates of canopy cover and forest density followed a similar pattern across the density gradient, demonstrating the effectiveness of UAV image‐derived estimates in low‐ to medium‐density conditions as well as the challenges associated with high‐density conditions. We found that post‐treatment conditions met a majority of the prescription objectives and demonstrate the UAV image application in quantifying changes from a mechanical thinning treatment. We provide a novel approach to forest restoration monitoring using UAV ‐derived data, one that considers varying density conditions and spatial scales. Future research should consider a more spatially extensive sampling design, including different restoration treatments, as well as experimenting with different combinations of equipment, flight parameters, and data processing workflows.
American black bears (Ursus americanus) are endemic to North America, having speciated from other ursids some 1.2 to 1.8 million years ago (Kurtn & Anderson 1994). During that time, black bears came to occupy nearly all of the forested areas of the North American continent. Historically, black bears were one of the most important mammals to indigenous peoples of North America by providing food, fat, hides, and tools (Raybourne 1987). Bears also played important roles in indigenous culture as symbols of strength, hard work, and love (see Rockwell 1991). In the 18th and 19th centuries in the U.S., Canada, and Mexico, European settlers with firearms reduced black bear numbers (Williams 1930), but the axe and plow had equally negative effects on bear populations as forests gave way to agriculture and livestock. Black bears also faced intense pressure from unregulated market hunting, poaching, and predator control programs. Black bears disappeared from large portions of their North American range, being relegated to swamps, thickets, rugged mountains, and other areas deemed too inaccessible, unsuitable, or undesirable to be occupied by humans (Pelton & van Manen 1994). In the early 20th century, Canadian and U.S. governments established national parks, forests, and refuges, and state and provincial legislatures created wildlife management agencies and enacted regulations to manage wildlife populations. By the late 20th century, the combination of public land infrastructure, habitat improvements, harvest regulations, law enforcement, and information-education programs resulted in significant increases in black bear numbers and range expansions in the U.S. and Canada (Pelton & van Manen 1994), aided in no small part by the remarkable adaptability of this animal (Pelton 2003; Scheick & McCown 2014). Today, the American black bear is one of the most iconic, abundant, and economically important bear species in the world. Unfortunately, black bear populations in Mexico have not recovered similarly to those in the U.S and Canada, and the status of the species there is uncertain.
North American Breeding Bird Survey data show that wood thrush (Hylocichla mustelina) populations in eastern U.S. forests have declined 1.8% per year during 1966-95. The declining quality of breeding forest tracts in North America is one possible cause for the apparent decline of some neotropical migratory birds, such as the wood thrush. In Georgia, however, wood thrush populations have declined during a period of increasing pine forest area and larger patch sizes. We hypothesized that forest management practices such as thinning and prescribed burning might create unsuitable habitat for wood thrushes. We conducted a four-year before/after, treatment/control experiment at the Piedmont National Wildlife Refuge in central Georgia to study to the effects of a treatment of thinning and prescribed burning on wood thrush demographic parameters. We simultaneously monitored wood thrush adults and juveniles with mark-recapture, radio-telemetry, nest searches, and plot-map surveys. Our analyses showed that wood thrushes were less likely to emigrate from the study compartments after the treatment, and wood thrushes exhibited some tendency to increase preference for hardwood habitats and decrease preference for pine habitats following the treatment. However, we observed no effects of treatment on nest success, adult survival, and adult and juvenile dispersal distances. We also found that female wood thrushes had lower survival rates than males during the breeding season, and we documented large-scale, within-year dispersal movements of adult (up to 17 km) and juvenile (up to 7 km) wood thrushes. We conclude that landscape level habitat quantity and quality must be considered during songbird management decisions. The documentation of sex- and age-specific wood thrush survival and movement rates was critical for construction of a set of population models. We used three stochastic models to learn more about wood thrush population dynamics and make predictions about population growth rates, reproductive success, and the effect of habitat changes on wood thrush populations. The simplest source/sink population model suggests that the Piedmont National Wildlife Refuge's wood thrush population is probably stable or increasing, and wood thrush populations in treated areas had higher growth rates than birds in untreated areas. We were able to use the individual-based model of wood thrush productivity to predict fecundity, a parameter that we could not measure directly in the field. Again, females on treated areas had higher fecundity than birds on untreated areas. Our spatially-based model predicted that wood thrush populations should respond positively to predicted changes in the age/size class structure of the Refuge's pine forests. Our model also showed that most wood thrushes leave the Refuge's forest compartments during the breeding season, and these dispersal movements are extremely important to understanding and managing wood thrush populations. The use of prescribed burning and retention shelterwood silviculture at the Piedmont National Wildlife Refuge does not appear to negatively affect the local wood thrush population. Continued use of the current management regime should result in adequate nesting, foraging, and escape habitats for wood thrushes. However, landscape-level habitat availability and quality, including lands outside the Refuge, must be considered when making management decisions that may affect wood thrushes.
Ungulate browsing in predator depleted North American landscapes is believed to be causing widespread tree recruitment failures. However, canopy disturbances and variations in ungulate densities are sources of heterogeneity that can buffer ecosystems against herbivory. Relatively little is known about the functional response (the rate of consumption in relation to food availability) of ungulates in eastern temperate forests, and therefore how “top down” control of vegetation may vary with disturbance type, intensity, and timing. This knowledge gap is relevant in the Northeastern United States today with the recent arrival of hemlock woolly adelgid (HWA; Adelges tsugae ) that is killing eastern hemlocks ( Tsuga canadensis ) and initiating salvage logging as a management response. We used an existing experiment in central New England begun in 2005, which simulated severe adelgid infestation and intensive logging of intact hemlock forest, to examine the functional response of combined moose ( Alces americanus ) and white-tailed deer ( Odocoileus virginianus ) foraging in two different time periods after disturbance (3 and 7 years). We predicted that browsing impacts would be linear or accelerating (Type I or Type III response) in year 3 when regenerating stem densities were relatively low and decelerating (Type II response) in year 7 when stem densities increased. We sampled and compared woody regeneration and browsing among logged and simulated insect attack treatments and two intact controls (hemlock and hardwood forest) in 2008 and again in 2012. We then used AIC model selection to compare the three major functional response models (Types I, II, and III) of ungulate browsing in relation to forage density. We also examined relative use of the different stand types by comparing pellet group density and remote camera images. In 2008, total and proportional browse consumption increased with stem density, and peaked in logged plots, revealing a Type I response. In 2012, stem densities were greatest in girdled plots, but proportional browse consumption was highest at intermediate stem densities in logged plots, exhibiting a Type III (rather than a Type II) functional response. Our results revealed shifting top–down control by herbivores at different stages of stand recovery after disturbance and in different understory conditions resulting from logging vs. simulated adelgid attack. If forest managers wish to promote tree regeneration in hemlock stands that is more resistant to ungulate browsers, leaving HWA-infested stands unmanaged may be a better option than preemptively logging them.
Kelp forests are phyletically diverse, structurally complex and highly productive components of coldwater rocky marine coastlines. This paper reviews the conditions in which kelp forests develop globally and where, why and at what rate they become deforested. The ecology and long archaeological history of kelp forests are examined through case studies from southern California, the Aleutian Islands and the western North Atlantic, well-studied locations that represent the widest possible range in kelp forest biodiversity. Global distribution of kelp forests is physiologically constrained by light at high latitudes and by nutrients, warm temperatures and other macrophytes at low latitudes. Within mid-latitude belts (roughly 40–60° latitude in both hemispheres) well-developed kelp forests are most threatened by herbivory, usually from sea urchins. Overfishing and extirpation of highly valued vertebrate apex predators often triggered herbivore population increases, leading to widespread kelp deforestation. Such deforestations have the most profound and lasting impacts on species-depauperate systems, such as those in Alaska and the western North Atlantic. Globally urchin-induced deforestation has been increasing over the past 2–3 decades. Continued fishing down of coastal food webs has resulted in shifting harvesting targets from apex predators to their invertebrate prey, including kelp-grazing herbivores. The recent global expansion of sea urchin harvesting has led to the widespread extirpation of this herbivore, and kelp forests have returned in some locations but, for the first time, these forests are devoid of vertebrate apex predators. In the western North Atlantic, large predatory crabs have recently filled this void and they have become the new apex predator in this system. Similar shifts from fish- to crab-dominance may have occurred in coastal zones of the United Kingdom and Japan, where large predatory finfish were extirpated long ago. Three North American case studies of kelp forests were examined to determine their long history with humans and project the status of future kelp forests to the year 2025. Fishing impacts on kelp forest systems have been both profound and much longer in duration than previously thought. Archaeological data suggest that coastal peoples exploited kelp forest organisms for thousands of years, occasionally resulting in localized losses of apex predators, outbreaks of sea urchin populations and probably small-scale deforestation. Over the past two centuries, commercial exploitation for export led to the extirpation of sea urchin predators, such as the sea otter in the North Pacific and predatory fishes like the cod in the North Atlantic. The large-scale removal of predators for export markets increased sea urchin abundances and promoted the decline of kelp forests over vast areas. Despite southern California having one of the longest known associations with coastal kelp forests, widespread deforestation is rare. It is possible that functional redundancies among predators and herbivores make this most diverse system most stable. Such biodiverse kelp forests may also resist invasion from non-native species. In the species-depauperate western North Atlantic, introduced algal competitors carpet the benthos and threaten future kelp dominance. There, other non-native herbivores and predators have become established and dominant components of this system. Climate changes have had measurable impacts on kelp forest ecosystems and efforts to control the emission of greenhouse gasses should be a global priority. However, overfishing appears to be the greatest manageable threat to kelp forest ecosystems over the 2025 time horizon. Management should focus on minimizing fishing impacts and restoring populations of functionally important species in these systems.
Soils in Alaska, and in high latitude terrestrial ecosystems in general, contain significant amounts of organic carbon, most of which is believed to have accumulated since the start of the Holocene about 10 ky before present. High latitude soils are estimated to contain 30-40% of terrestrial soil carbon (Melillo et al., 1995; McGuire and Hobbie, 1997), or ~ 300-400 Gt C (Gt = 1015 g), which equals about half of the current atmospheric burden of carbon. Boreal forests in particular are estimated to have more soil carbon than any other terrestrial biome (Post et al., 1982; Chapin and Matthews, 1993). The relations among net primary production, soil carbon storage, recurrent fire disturbance, nutrients, the hydrologic cycle, permafrost and geomorphology are poorly understood in boreal forest. Fire disturbance has been suggested to play a key role in the interactions among the complex biogeochemical processes influencing carbon storage in boreal forest soils (Harden et al., 2000; Zhuang et al., 2002). There has been an observed increase in fire disturbance in North American boreal black spruce (Picea mariana) forests in recent decades (Murphy et al., 1999; Kasichke et al., 2000), concurrent with increases in Alaskan boreal and arctic surface temperatures and warming of permafrost (Osterkamp and Romanofsky, 1999). Understanding the role of fire in long term carbon storage and how recent changes in fire frequency and severity may influence future high latitude soil carbon pools is necessary for those working to understand or mitigate the effects of global climate change.
The information on the mangrove forests for the Pacific region is scarce or outdated. A regional assessment based on a consistent methodology and data sources was needed to understand their true extent. Our investigation offers a regionally consistent, high resolution (30 m), and the most comprehensive mapping of mangrove forests on the islands of American Samoa, Fiji, French Polynesia, Guam, Hawaii, Kiribati, Marshall Islands, Micronesia, Nauru, New Caledonia, Northern Mariana Islands, Palau, Papua New Guinea, Samoa, Solomon Islands, Tonga, Tuvalu, Vanuatu, and Wallis and Futuna Islands for the year 2000. We employed a hybrid supervised and unsupervised image classification technique on a total of 128 Landsat scenes gathered between 1999 and 2004, and validated the results using existing geographic information science (GIS) datasets, high resolution imagery, and published literature. We also draw a comparative analysis with the mangrove forests inventory published by the Food and Agriculture Association (FAO) of the United Nations. Our estimate shows a total of 623755 hectares of mangrove forests in the Pacific region; an increase of 18% from FAO's estimates. Although mangrove forests are disproportionately distributed toward a few larger islands on the western Pacific, they are also significant in many smaller islands.
Assessing outcomes of habitat management is critical for informing and adapting conservation plans. From 2013 – 2019, a multi-stage management initiative aims to create >25,000 ha of shrubland and early-successional vegetation to benefit Golden-winged Warblers (Vermivora chrysoptera) in managed forested landscapes of the western Great Lakes region. We studied a dense breeding population of Golden-winged Warblers at Rice Lake National Wildlife Refuge (NWR) in Minnesota, USA, where shrubs and young trees were sheared during the winter of 2014-2015 in a single treatment supported in part by the American Bird Conservancy (ABC) and in part by other funding source(s) to benefit Golden-winged Warblers and other species associated with young forest [e.g., American Woodcock (Scalopax minor)] and as part of maintenance of early successional forest cover on the refuge.
Article for outlet: Plant Ecology. Abstract: Big sagebrush (Artemisia tridentata Nutt.) plant communities are widespread non-forested drylands in western North American and similar to all shrub steppe ecosystems world-wide are composed of a shrub overstory layer and a forb and graminoid understory layer. Forbs account for the majority of plant species diversity in big sagebrush plant communities and are important for ecosystem function. Few studies have explored the geographic patterns of forb species richness and composition and their relationships with environmental variables in these communities. Our objectives were to examine the small and large-scale spatial patterns in forb species richness and composition and the influence of environmental variables. We sampled forb species richness and composition along transects at 15 field sites in Colorado, Idaho, Montana, Nevada, Oregon, Utah, and Wyoming, built species-area relationships to quantify differences in forb species richness at sites, and used Principal Components Analysis and nonmetric multidimensional scaling to identify relationships among environmental variables and forb species richness and composition. We found that species richness was most strongly correlated with soil texture, while species composition was most related to climate. The combination of climate and soil texture influences water availability, with important consequences for forb species richness and composition, which suggests climate-change induced modification of soil water availability may have important implications for plant species diversity in the future. Our paper is the first to our knowledge to examine forb biodiversity patterns in big sagebrush ecosystems in relation to environmental factors across the big sagebrush region.
The diversity and distribution of mammals in the American tropics remain incompletely known. We describe a new species of small-eared shrew (Soricidae, Cryptotis ) from the Lacandona rain forest, Chiapas, southern Mexico. The new species is distinguished from other species of Cryptotis on the basis of a unique combination of pelage coloration, size, dental, cranial, postcranial, and external characters, and genetic distances. It appears most closely related to species in the Cryptotis nigrescens species group, which occurs from southern Mexico to montane regions of Colombia. This discovery is particularly remarkable because the new species is from a low-elevation habitat (approximately 90 m), whereas most shrews in the region are restricted to higher elevations, typically > 1,000 m. The only known locality for the new shrew is in one of the last areas in southern Mexico where relatively undisturbed tropical vegetation is still found. The type locality is protected by the Mexican government as part of the Yaxchilán Archaeological Site on the border between Mexico and Guatemala.
Many studies and production inventory systems have shown the utility of coupling covariates derived from Light Detection and Ranging (LiDAR) data with forest variables measured on georeferenced inventory plots through regression models. The objective of this study was to propose and assess the use of a Bayesian hierarchical modeling framework that accommodates both residual spatial dependence and non-stationarity of model covariates through the introduction of spatial random effects. We explored this objective using four forest inventory datasets that are part of the North American Carbon Program, each comprising point-referenced measures of above-ground forest biomass and discrete LiDAR. For each dataset, we considered at least five regression model specifications of varying complexity. Models were assessed based on goodness of fit criteria and predictive performance using a 10-fold cross-validation procedure. Results showed that the addition of spatial random effects to the regression model intercept improved fit and predictive performance in the presence of substantial residual spatial dependence. Additionally, in some cases, allowing either some or all regression slope parameters to vary spatially, via the addition of spatial random effects, further improved model fit and predictive performance. In other instances, models showed improved fit but decreased predictive performance—indicating over-fitting and underscoring the need for cross-validation to assess predictive ability. The proposed Bayesian modeling framework provided access to pixel-level posterior predictive distributions that were useful for uncertainty mapping, diagnosing spatial extrapolation issues, revealing missing model covariates, and discovering locally significant parameters.
Within the forests of Pictured Rocks National Lakeshore, biologists are trying to understand the effects beech bark disease has on wildlife species, especially species that need forest connectivity to thrive. This project used aerial imagery collected in 2005, shortly after beech bark disease infestation, and satellite imagery from 2018. The 2018 imagery represents present day conditions and was used to locate forest canopy gaps through object-based image analysis. Forest canopy gaps were identified using the multiresolution segmentation algorithm within Trimble’s eCognition software. A time change analysis was completed to understand how the forest canopy had changed from 2005 to 2018. The analysis showed areas that had maintained forest canopy, maintained a forest canopy gap, created a new canopy gap (closed forest canopy in 2005 but open canopy gap in 2018), or created new forest canopy (open canopy gap in 2005 but closed forest canopy in 2018). There were 9,127 acres of forest canopy lost, and 72.8 percent of that lost canopy occurred in a forest type where Fagus grandifolia Ehrh. (American beech) is a common tree species. The datasets developed through this project can enhance knowledge of where canopy gaps exist and help place focus on certain areas for wildlife studies. In addition, these datasets can be used in future studies to monitor the health of the forest and conduct additional change analyses.
Pinyon-juniper woodlands are one of the largest ecosystems in the Southwest and in the Middle Rio Grande Basin (Fig. 1). The woodlands have been important to the region's inhabitants since prehistoric times for a variety of natural resources and amenities. The ecosystems have not been static; their distributions, stand characteristics, and site conditions have been altered by changes in climatic patterns and human use and, often, abuse. Management of these lands since European settlement has varied from light exploitation and benign neglect, to attempts to remove the trees in favor of forage for livestock, and then to a realization that these lands contain useful resources and should be managed accordingly. Land management agencies are committed to ecosystem management. While there are several definitions of ecosystem management, the goal is to use ecological approaches to create and maintain diverse, productive, and healthy ecosystems (Kaufmann et al. 1994). Ecosystem management recognizes that people are an integral part of the system and that their needs must be considered. Ecological approaches are central to the concept, but our understanding of basic woodland ecology is incomplete, and there are different opinions and interpretations of existing information (Gottfried and Severson 1993). There are many questions concerning proper ecosystem management of the pinyon-juniper woodlands and how managers can achieve these goals (Gottfried and Severson 1993). While the broad concept of ecosystem management generally is accepted, the USDA Forest Service, other public land management agencies, American Indian tribes, and private landowners may have differing definitions of what constitutes desired conditions. Key questions about the pinyon-juniper ecosystems remain unanswered. Some concern the basic dynamics of biological and physical components of the pinyon-juniper ecosystems. Others concern the distribution of woodlands prior to European settlement and changes since the introduction of livestock and fire control. This relates to whether tree densities have been increasing or whether trees are invading grasslands and, to a lesser extent, drier ponderosa pine (Pinus ponderosa) forests. In areas where woodlands were heavily used by American Indians for fuelwood prior to European contact, the advance of pinyon and juniper could represent the slow recovery from intensive use (Samuels and Betancourt 1982). There are numerous questions regarding declines in watershed condition related to changes in pinyon-juniper tree stand densities and to the density and composition of understory vegetation. There are different opinions about proper management of woodland ecosystems. Should these lands be managed for a single resource, such as forage for livestock production, or managed for sustained production of multiple resource products and amenities? Depending on site and stand conditions, the woodlands can produce variable quantities of fuelwood, pinyon nuts, wildlife habitat, forage for livestock, and cover for watershed protection. Management must also consider increasing recreational demands, threatened and endangered species, and protection of archeological sites. Many pinyon-juniper woodland watersheds in New Mexico have unsatisfactory soil and watershed conditions (USDA Forest Service 1993); managers must develop restoration procedures that recognize the value of woodland ecosystems. The concerns, questions, and conflicts surrounding management of pinyon-juniper lands, as well as the ecological foundations of ecosystem management, require that all interested parties reevaluate attitudes toward the woodlands. Ecosystem management goals and concepts recognize diversity. Pinyonjuniper woodlands are diverse, and stand characteristics and site productivities vary. Management objectives and prescriptions must evaluate the potential of each site, and decisions must be based on sound scientific information. This information is often unavailable. Therefore, this paper describes what we do know about the characteristics, distribution, and ecology of pinyon-juniper woodlands, including the effects of natural and human factors, within the southwestern United States and particularly the Middle Rio Grande Basin. It also reviews some past and present management options in this widespread and important vegetation type. The review draws on research and management information from the Rio Grande Basin and from similar areas in the Southwest and adjacent regions. It does not attempt to review all of the relevant literature; additional sources can be found within the articles cited in the References.