Coasts: Complex changes affecting the Northwest's diverse shorelines
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Although there have been heated discussions concerning the relative importance of using Mhc diversity as a basis for selecting breeders in conservation projects, most parties agree that the genetic variability residual in an endangered species should be maintained through genetic management, if at all possible. Substantial evidence exists (particularly in birds) documenting the influences of specific Mhc haplotypes on disease outcome and also that those individuals which are heterozygous for Mhc alleles appear to have an advantage for survival over those that are homozygous. Thus, conservation of genetic variability of the Mhc is likely important for the preservation of fitness, especially in small breeding populations. More than half of the world's crane species are listed as endangered. Members of all 15 known species are represented among breeding animals for captive propagation at the International Crane Foundation (Wisconsin) and the USGS Patuxent Wildlife Research Center (Maryland). Collaborative multi-organization efforts and the availability of extensive pedigree records have allowed the study of Mhc variability in several species of cranes. We have found, for example, that Mhc diversity in the captive Florida sandhill crane (Grus canadensis pratensis) population appears high, whereas in the captive whooping crane (Grus americana), which has undergone a severe 'genetic bottleneck,? both the number of alleles and the levels of heterozygosity appear to be substantially reduced.
Several points in the original paper are clarified. (Preceding abstract)-M.S.
The sources of the elements incorporated into pocket-filling clays and pseudomorphous minerals, the conditions of formation, and the paragenesis of these minerals are discussed.-J.A.Z.
Strontium isotopes (expressed as per mill deviation from mean sea water, ??87Sr) reflect interaction between ground water and the aquifer through which it is flowing. In the Cenozoic aquifer of the Yucca Mountain region, ??87Sr values increase from north to south downgradient in the flow system. The largest ??87Sr values occur in the Amargosa Desert where ground water probably encounters alluvial basin fill derived from Precambrian rocks in the Funeral Range. Similarly, large ??87Sr values for ground water in the Paleozoic aquifer at the western end of the Spring Mountains also probably reflect an encounter with Precambrian rocks. In several wells into the volcanic rocks, apparent isotopic disequilibrium between ground water and the producing units suggests that the ground water probably integrates over a substantial part of the saturated section in attaining its strontium isotope signature.
An extremely proximal ejecta ring, with exposures to within 100 m of vent, was deposited during later-stage plinian fall activity during the 1912 Novarupta eruption in Alaska. One bed in the ejecta ring (bed S) contains predominantly andesitic clasts which serve to delineate the striking contrast in thinning rates along dispersal axis of the ejecta ring [Pyle bt values of 70 m (bed S alone) or 190 m (whole ejecta ring)] and the coeval dacitic plinian fall deposits [Pyle bt, values of 4 km (proximal) and 37 km (medial-distal)]. The locally deposited andesitic and dacitic clasts of the ejecta ring are interpreted as products of an irregular 'collar' of low-fountaining ejecta partially sheathing the core of higher-velocity dacitic ejecta that fed the stable, convecting 23-km-high column. The presence of such an extremely proximal accumulation of ejecta appears to be a feature common to several other historic eruptions that generated widespread fall deposits. This feature in part accounts for conflicts between measured and calculated values for thickness maxima in plinian fall deposits and suggests that modifications may be required of existing models for plinian eruption columns.
The Crystal Peak area of the Pikes Peak batholith, near Lake George in central Colorado, is world-renowned for its crystals of amazonite (the blue-green variety of microcline) and smoky quartz. Such crystals, collected from individual miarolitic pegmatites, have a remakably small variation in crystal size within each pegmatite, and the shapes of plots of their crystal size distributions (CSDs) are invariably lognormal or close to lognormal in all cases. These observations are explained by a crystal growth mechanism that was governed initially by surface-controlled kinetics, during which crystals tended to grow larger in proportion to their size, thereby establishing lognormal CSDs. Surface-controlled growth was followed by longer periods of supply controlled growth, during which growth rate was predominantly size-independent, consequently preserving the lognormal shapes of the CSDs and the small size variation. The change from surface- to supply controlled growth kinetics may have resulted from an increasing demand for nutrients that exceeded diffusion limitations of the system. The proposed model for crystal growth in this locality appears to be common in the geologic record, and can be used with other information, such as isotopic data, to deduce physico-chemical conditions during crystal formation.
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In contrast to most local amphibian populations, northeastern populations of the Northern Leopard Frog (Lithobates pipiens) have displayed uncharacteristically high levels of genetic diversity that have been attributed to large, stable populations. However, this widely distributed species also occurs in areas known for great climatic fluctuations that should be reflected in corresponding fluctuations in population sizes and reduced genetic diversity. To test our hypothesis that Northern Leopard Frog genetic diversity would be reduced in areas subjected to significant climate variability, we examined the genetic diversity of L. pipiens collected from 12 sites within the Prairie Pothole Region of North Dakota. Despite the region's fluctuating climate that includes periods of recurring drought and deluge, we found unexpectedly high levels of genetic diversity approaching that of northeastern populations. Further, genetic structure at a landscape scale was strikingly homogeneous; genetic differentiation estimates (D est ) averaged 0.10 (SD = 0.036) across the six microsatellite loci we studied, and two Bayesian assignment tests (STRUCTURE and BAPS) failed to reveal the development of significant population structure across the 68 km breadth of our study area. These results suggest that L. pipiens in the Prairie Pothole Region consists of a large, panmictic population capable of maintaining high genetic diversity in the face of marked climate variability.
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