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Some limitations on the possible composition of the ore-forming fluid

The activity rations of various important anions (S, CO 3 , SO 4 , OH, F, and Cl) in hydrothermal solutions at the time of deposition are evaluated using a simple thermodynamic technique. The rations are interpreted in the light of the mineralogy of ore deposits and limites are placed on the variability of each ratio in hydrothermal solutions. All of the calculations are made for 25°C and cautious extrapolation to higher temperatures seems justified; however, additional data for elevated temperatures and pressures are needed before more than approximate values may be assigned to these ratios in the ore-forming fluid. The calculated partial pressure of CO 2 in the ore fluid is generally less than one atmosphere, which suggests that a dense CO 2 phase cannot be considered an importatn ore fluid for most deposits. The partial pressure of H 2 S is usually less than 10 -4 atmospheres which makes it extremely difficult to defend the heory that metals (other than the easily complexible mercury, arsenic, antimony, and perhaps fols and silver) are transported in quantity as complex sulfide and hydrosulfides. The sulfate to sulfide ration is such that the oxidation potential at the time of deposition is defined by the following equation: Eh (in volts) = 0.22 ± 0.04 - 0.059 pH.

Trace Elements Investigations

Swatara Creek basin of southeastern Pennsylvania: An evaluation of its hydrologic system

Local concentrations of population in the Swatara Creek basin of Pennsylvania find it necessary to store, transport, and treat water because local supplies are either deficient or have been contaminated by disposal of wastes in upstream areas. Water in the basin is available for the deficient areas and for dilution of the coal-mine drainage in the northern parts and the sewage wastes in the southern parts. Swatara Creek drains 576 square miles just east of Harrisburg, Pa., and is the largest tributary to the Susquehanna River from the north side below Harrisburg. It rises in the southern Pocono Mountains and flows southwestward across the Lebanon Plateau. On an average day Swatara Creek discharges more than 630 million gallons into the Susquehanna River at Middletown, Pa. In a year this amounts to about 23 inches of water over the entire basin and is the residual from an average annual precipitation of 45.5 inches. During an average year the flow in Swatara Creek from the upper third of the basin above Harper Tavern is always greater than 1,300 mgd (million gallons per day) for at least 15 days and is always greater than 25 mgd for at least 350 days. The daily streamflow from the basin averages 1.1 mgd per sq mi, but yields from different areas range from 0.97 to 1.22 mgd per sq mi. These variations are caused chiefly by differences in precipitation and land cover. The area of lowest yield is in the valleys west of Tremont, and the highest yields are in the Upper and Lower Little Swatara Creek subbasins. At high and medium stages the chemical character of the water in the streams is suitable for public and private supplies. At lower stages, defending on the areas and the amounts of contamination by coal-mine drainage and sewage pollution, the natural flow may require some treatment. At low stages the chemical characteristics of the natural flow not affected by man is almost identical with that of the ground water in the area drained by the stream. In general, the total dissolved solids range from about 25 to 400 parts per million and the hardness is as much as about 300 parts per million. The ground-water increment to the base flow of Swatara Creek averages about 240 mgd, or about 8.8 inches annually, for the basin. Generally, ground-water supplies in amounts of less than 0.5 mgd can be developed south of Blue Mountain. Supplies of several million gallons per day have been developed for industrial use from the permeable limestones in the south-central part of the basin. More intensive investigation in other parts of the basin would indicate areas where supplies of more than 0.5 mgd could be developed from properly spaced wells. The chemical character of water from wells depends largely on the host rock. In highly soluble rocks water contains large amount of dissolved solids; in more resistant rocks concentrations are lower. The chemical character of unpolluted ground water generally reflects the composition of the more readily soluble minerals in the local geologic environment. Areas contaminated by septic- tank effluent may have above normal amounts of nitrate and detergent products. Except where polluted, most ground water is suitable for public and industrial uses without extensive treatment. Sites for storage of surface water exist in the part of the basin lying in the valley and ridge area. As much as 30 to 40 percent of the annual flow could be impounded for release as low-flow augmentation for dilution of mine drainage and other wastes in the basin. Low sediment yields of supplying drainage areas would ensure a long life expectancy of reservoirs at these sites. Overbank flooding of the main stem of the Swatara Creek and its tributaries has occurred many times in the past. However, it has not been a hazard because urban development has not encroached on the flood plain. An inundation map of the August 1933 flood provides a basis that urban planners may use to avoid future damage. As water in the Swatara Creek moves downstream to the Susquehanna River, the flow is influenced consecutively by a large annual rainfall on the northern valley and ridge area, the wastes of surface and subsurface coal-mining activities, and less annual rainfall on the part of the basin lying in the Lebanon Plateau area; the flow is supplemented and further influenced by many tributaries and by the industrial and domestic wastes that are carried by these secondary streams. The annual precipitation ranges from 52 inches at the east edge and 49 inches at the west edge of the mountainous part of the basin to about 41 inches at the southwestern part at Middletown. The rainfall generally is adequate during the growing season to mature the crops. The mean annual temperature at Lebanon is about 52°F, and the growing season is about 180 days. In this report the basin has been divided into eight hydrologic zones, leased on runoff, natural use of water, and chemical character of water. Four zones lie in the valley and ridge area, three lie in the Lebanon Plateau area, and one lies in the highland along the southeastern basin boundary. In each of the zones the hydrologic characteristics are virtually the same, but they may be completely different from those in adjacent zones. The boundaries of the zones generally coincide with boundaries between geologic formations, and the areas in each zone include rocks of similar influence on water. Streams in zone 4 at the northeast edge of the plateau have the highest average surface runoff from 1.2 to 1.1 mgd per sq mi whereas those in zone 2 at the northwest edge of the valley and ridge area have the lowest, about 1.0 mgd. Streams in zone 8, along the southeast edge of the basin, have the largest sustained low-flow yield, about 0.26 to 0.19 mgd per sq mi; those in zone 5 overlying the Martinsburg Shale east of Harrisburg have the smallest sustained low-flow yields, 0.03 to 0.01 mgd. Streams in the limestone area of zone 7 have the greatest range in low-flow yields in any one zone from 0.60 to 0 mgd per sq mi. Low-flow yields in zones 1 through 4 range from 0.13 to 0.03 mgd per sq mi. Surface flows from zones 1 and 2 are generally acidic and contain high concentrations of sulfate, iron, and total dissolved solids especially where contaminated with mine wastes. Surface flows from zones 3 and 4 are dilute, slightly alkaline, and suitable for public water supplies. Surface flows from zones 5, 6, and 7 are alkaline and contain moderate concentrations of dissolved solids with waters of highest hardness occurring in zone 7. Surface flows from zone 8 are dilute to moderately mineralized and are relatively high in silica concentration. Nitrate concentrations are high in surf Fee flows below sewage outfalls and in ground water contaminated by septic tank effluent and industrial wastes. Average annual sediment yields of 550 to 650 tons per square mile are characteristic of zones 1 and 2 where strip mining has destroyed the forest cover and coal culm is carried into the streams. From agricultural lands on the Martinsburg Shale in zones 5 and 6, annual sediment yields range from 300 to 350 tons per square mile; but from agricultural lands on the siliceous rocks in zone 8 and zones 3 and 4 in the valley and ridge area, the sediment yield ranges from 200 to 250 tons annually per square mile. Lowest annual sediment yields in the basin are in the forested areas of siliceous rocks in zones 2, 3, 4, and 5, and in the sinkhole topography of the limestones in zone 7 where the yield ranges from 30 to 35 tons and 50 to 60 tons per square mile, respectively. The amount of ground water that can be developed in the basin is dependent on the ability of the underlying rocks to yield water to wells. More than 300 gpm (gallons per minute) can be obtained from wells in alluvial materials in the valley bottoms and in some of the limestones where large solution channels and fractures are penetrated by the wells. From 50 to 300 gpm can be obtained from wells in loosely cemented sandstones and in fractured limestones. From 10 to 50 gpm can be developed from wells in the shales and harder sandstones. The most dense rocks will yield from 1 to 10 gpm from fractures and crevices. Most wells yield water from the upper 350 feet of the formation, for this part contains the most fractures or solution channels. Studies show that the velocity at which a contaminant will move downstream in the basin is related to the discharge of the stream at the time. At a stream discharge of about 400 mgd at Pine Grove, a contaminant in Swatara Creek would require about 40 hours to move from Pine Grove to Middletown. As a result of dispersion and dilution, the maximum concentration of the contaminant at Middletown would be less than 20 percent the concentration at Pine Grove under these conditions. An evaluation of the availability of water in the basin indicates that about I,239 mgd enters as precipitation, 630 mgd leaves as streamflow, 580 mgd is evaporated and transpired, and 56 mgd is diverted for use by man. Not all the diversions for man's use are lost to the basin, as about 27 mgd is returned as sewage for reuse. About one-fourth of the waste water is returned to the ground and the remainder to stream drainageways. Of that diverted by man, 11.6 mgd is used for public supply and 44.4 mgd for industrial and private supplies. Diversions of streamflow furnish 86 percent of the public supply and 27 percent of the industrial supply, and ground-water sources yield the remainder. Municipal and private sewage treatment plants are upgrading the waste water in many places, but no provisions are being made for treatment other than natural dilution and assimilation for the 15 mgd of coal-nine drainage in the northern part of the basin. Technology for economic treatment of mine water is not available at this time, although research in this field is being done. Urbanization eastward from Harrisburg and around Lebanon has increased the population density of the basin. Densities of 500 people per square mile and water use exceeding 2.0 mgd per sq mi can be expected in the future. By the year 2000 the population of the basin may increase 60 percent; and if the per capita rate of use increases 0.5 percent per year the domestic requirements for water will be about two times the present use, or 23 mgd. Similarly, if the present 1:4 ratio of domestic use to industrial use of water continues, at least 89 mgd will be needed for industry in the future. Although an increase to twice the present use of water can be foreseen, or 112 mgd, water for the dilution and assimilation of wastes from treatment systems are not included. Providing water for dilution of wastes from treatment plants has not been a problem, but in the future the amounts needed for this purpose will be greater as the population increases. As water becomes more valuable, treatment of sewage wastes to reduce the biochemical-oxygen-demand load by at least 80 to 90 percent will be necessary to conserve water for more productive uses. As much as 100 mgd may be needed for waste dilution in the basin by year 2000. The present trends in suburban and light industrial development will probably persist in the basin. Problems arising through changes in economic value of water, conflicts in use, and alternatives in development are typical of those confronting the manager of a water-resource system.

Pennsylvania

Bald eagle predation on common loon egg

The Common Loon ( Gavia immer ) must defend against many potential egg predators during incubation, including corvids, Herring Gulls ( Larus argentatus ), raccoons ( Procyon lotor ), striped skunk ( Mephitis mephitis ), fisher ( Martes pennanti ), and mink ( Neovison vison ) ( McIntyre 1988 , Evers 2004 , McCann et al. 2005 ). Bald Eagles ( Haliaeetus leucocephalus ) have been documented as predators of both adult Common Loons and their chicks ( Vliestra and Paruk 1997 , Paruk et al. 1999 , Erlandson et al. 2007 , Piper et al. 2008 ). In Wisconsin, where nesting Bald Eagles are abundant (>1200 nesting pairs, >1 young/pair/year), field biologists observed four instances of eagle predation of eggs in loon nests during the period 2002&ndash;2004 (M. Meyer pers. comm.). In addition, four cases of eagle predation of incubating adult loons were inferred from evidence found at the loon nest (dozens of plucked adult loon feathers, no carcass remains) and/or loon leg, neck, and skull bones beneath two active eagle nests, including leg bones containing the bands of the nearby (<25 m) incubating adult loon. However, although loon egg predation has been associated with Bald Eagles, predation events have yet to be described in peer-reviewed literature. Here we describe a photographic observation of predation on a Common Loon egg by an immature Bald Eagle as captured by a nest surveillance video camera on Lake Umbagog, a large lake (32 km 2 ) at Umbagog National Wildlife Refuge (UNWR) in Maine.

Maine

The spring migration of adult North American Ospreys

Most North American Ospreys ( Pandion haliaetus ) are migratory, breeding in northern latitudes and migrating long distances to and from their wintering grounds in the tropics. Although fall migration patterns of North American Ospreys have been described and studied, very little has been published about the spring migration of these birds. We used satellite telemetry to: (1) determine the characteristics (timing, duration, migratory routes) of spring migrations of Ospreys; (2) determine if differences in spring migration patterns existed between sexes and among three breeding populations (east coast, midwestern, and western); and (3) compare consecutive fall and spring migrations of individual Ospreys. The median dates for departure from the wintering grounds and arrival on the breeding grounds did not differ significantly between adult male and female Ospreys. Compared to their fall migrations, all male and all east coast Ospreys spent fewer days on migration, fewer days in stopover periods along the migration route, traveled shorter distances overall, and traveled farther (on average) each day during spring. In contrast, fall and spring migration characteristics of all female and western Ospreys were similar. Our findings suggest that, although sex and breeding location might influence the spring migration strategy used by individual Ospreys, both males and females minimize the time spent on migration to ensure a timely arrival on the breeding grounds to establish or defend a nesting territory.

Journal of Raptor Research

Use of noninvasive genetics to assess nest and space use by white-tailed eagles

Movement and space use are important components of animal interactions with the environment. However, for hard-to-monitor raptor species, there are substantial gaps in our understanding of these key determinants. We used noninvasive genetic tools to evaluate the details of space use over a 3-yr period by White-tailed Eagles ( Haliaeetus albicilla ) at the Naurzum Zapovednik in northern Kazakhstan. We genotyped, at 10 microsatellite markers and one mitochondrial marker, 859 eagle feathers and assigned naturally shed feathers to individuals. We identified 124 White-tailed Eagles, including both members of 5–10 pairs per year, and were able to monitor birds across years. Distances between eagle nests and hunting perches were always greater than nearest neighbor distances, eagles never used the closest available hunting perch, and hunting perches were always shared with other eagles. When eagles switched nests between years, the nests they chose were almost always well outside the space that theory predicted they defended the prior year. Our data are inconsistent with classical territorial and colonial models of resource use; they more closely resemble semi-colonial behavior. It is unlikely that standard methods of animal tracking (e.g., marking and telemetry), would have provided a similarly cost-effective mechanism to gain these insights into spatial and temporal aspects of eagle behavior. When combined with existing information on space use of other local species, these data suggest that partitioning of spatial resources among White-tailed Eagles and other eagles at the Zapovednik may be facilitated by the alternative strategies of space use they employ.

Naurzum Zapovednik

Comparison of preservation and extraction methods on five taxonomically disparate coral microbiomes

All animals are host to a multitude of microorganisms that are essential to the animal’s health. Host-associated microbes have been shown to defend against potential pathogens, provide essential nutrients, interact with the host’s immune system, and even regulate mood. However, it can be difficult to preserve and obtain nucleic acids from some host-associated microbiomes, making studying their microbial communities challenging. Corals are an example of this, in part due to their potentially remote, underwater locations, their thick surface mucopolysaccharide layer, and various inherent molecular inhibitors. This study examined three different preservatives (RNAlater, DNA/RNA Shield, and liquid nitrogen) and two extraction methods (the Qiagen PowerBiofilm kit and the Promega Maxwell RBC kit with modifications) to determine if there was an optimum combination for examining the coral microbiome. These methods were employed across taxonomically diverse coral species, including deep-sea/shallow, stony/soft, and zooxanthellate/azooxanthellate: Lophelia pertusa , Paragorgia johnsoni , Montastraea cavernosa , Porites astreoides , and Stephanocoenia intersepta . Although significant differences were found between preservative types and extraction methods, these differences were subtle, and varied in nature from coral species to coral species. Significant differences between coral species were far more profound than those detected between preservative or extraction method. We suggest that the preservative types presented here and extraction methods using a bead-beating step provide enough consistency to compare coral microbiomes across various studies, as long as subtle differences in microbial communities are attributed to dissimilar methodologies. Additionally, the inclusion of internal controls such as a mock community and extraction blanks can help provide context regarding data quality, improving downstream analyses.

Frontiers in Marine Science

Usurpation and brooding of Least Tern (Sternula antillarum) chicks by Common Terns (Sterna hirundo)

While nest usurpation and subsequent incubation of eggs and even brooding of chicks from other species has been reported for Common Terns ( Sterna hirundo ), such behavior is considered rare. We report an observation of a Common Tern pair usurping the Least Tern ( Sternula antillarum ) nest and brooding the Least Tern chicks. While the Least Tern pair attempted to provide care for the chicks, the Common Terns displayed aggressive behavior and defended the nest. Though both species attempted to feed the chicks, no feeding events were observed due to harassment from the other species. Neither pair was observed nesting prior to or following this event, and all chicks are believed to have been lost to predation. We discuss the possible scenarios leading to the observed usurpation event, the possibility that usurpations are more common than previously believed, and the need for different monitoring methods to elucidate the causes of usurpations.

Diversity

Observations of recruitment and colonization by tunicates and associated invertebrates using giant one-meter 2 recruitment plates at Woods Hole, Massachusetts

Large recruitment plates measuring 1 &times; 1 m were deployed over an 18-month period from September 2013 to March 2015 for the purpose of documenting recruitment and colonization processes of marine invertebrate species at Woods Hole, Massachusetts. Each side of two plates was subdivided into 16 subareas (25 &times; 25 cm), and an observational strategy was developed whereby, at approximately two-week intervals, a different subarea was cleaned. Using this approach, we were able to photographically document species recruitment and growth interactions. Water temperature records from the site show that steady warming and cooling between 3 and 20&deg; C changed at a mean rate of 0.2 &deg; C d -1 . However, temperature changes during the coolest and warmest parts of the temperature cycle were highly variable. In 2014, between the first and last occurrence of 0&deg; C, temperatures were &le;0&deg; C 15 percent of the time, but in 2015 temperatures were &le;0&deg; C 93 percent of the time. In 2014, between the first and last occurrence of 21&deg; C, temperatures were &ge;21&deg; C 88 percent of the time, and this warm period correlated with the disappearance of the hydroid Ectopleura crocea , the solitary tunicates Ascidiella aspersa and Ciona intestinalis , and the 2013 generation of Botrylloides violaceus . In Woods Hole, large plates provided enough space to accommodate both fast- and slow-colonizing species, resulting in the establishment of a diverse assemblage that was observed over a long time period. The most successful colonizing species had relatively long reproductive and recruitment periods, grew rapidly, repelled settlement onto their surfaces by larvae of any species, defended themselves against overgrowth by any species, overwintered, and lived a long time. Of the three dominant species observed in this study, the colonial tunicates Didemnum vexillum and Botrylloides violaceus had these qualities; the encrusting colonial bryozoan Schizoporella unicornis had all but one, it grew more slowly than the others. Barnacles constituted the only biological substrate that was effectively colonized by other species, both by larval recruitment and overgrowth. In Woods Hole, after a substrate had become fully colonized, there was very little opportunity for new recruitment or colony growth until new substrate opened after the death of colonies and individuals and the disappearance of biogenic structures such as amphipod tubes. An understanding of colonization processes utilized by invasive species allows prediction of their potential effects on ecosystems in areas where they are not yet present.

Massachusetts

Dynamics of the double-crested cormorant population on Lake Ontario

After nearly 30 years of recolonization and expansion across North America, the double-crested cormorant (Phalacrocorax auritus) occupies the role of a perceived and, in some situations, realized threat to fish stocks and other resources. However, population data necessary to plan, defend, and implement management of this species are few. Our purpose was to gain insight into the relative contribution of various population parameters to the overall rate of population growth and identify data needs critical to improving our understanding of the dynamics of double-crested cormorant populations. We demonstrated the construction of a biologically reasonable representation of cormorant population growth on Lake Ontario (1979-2000) by referencing literature values for fertility, age at first breeding, and survival. These parameters were incorporated into a deterministic stage-classified matrix model. By calculating the elasticity of matrix elements (i.e., statgspecific fertility and survival), we found that cormorant population growth on Lake Ontario was most sensitive to survival of birds about to turn age 3 and older. Finally, we demonstrated how this information could be used to evaluate management scenarios and direct future research by simulating potential environmental effects on fertility and survival, as well as a 5-year egg-oiling program. We also demonstrated that survival of older birds exerts more effective population control than changes in fertility.

Wildlife Society Bulletin

The role of nutrient reserves in mallard reproduction

Mallard ( Anas platyrhynchos ) populations breeding in temperate North America obtain a significant part of the energy and lipid requirements of reproduction at sites occupied prior to arrival on the breeding grounds. Protein for egg formation, however, is obtained principally from the diet during the nesting period. Both sexes arrive heavy and fat in North Dakota but experience substantial weight loss and lipid depletion during the nesting cycle. Weight loss is most pronounced among females and averages 25% from prelaying to late incubation. Body weights of both sexes are positively correlated with carcass lipid content. The paired male draws upon lipids early in the nesting season when an activity center is being established and defended and when females are preparing to nest. The female's lipid reserves are utilized primarily during laying and early incubation. The significance of lipid reserves diminishes as the nesting season progresses, and females do not acquire substantial lipid stores prior to renesting when the initial clutch is destroyed. The magnitude of lipid reserves carried in female carcasses is positively correlated with clutch size from mid-April to early June. Protein transfer for egg formation from flight and leg muscle and body organs can account for only a small part of the protein requirement for the clutch. By utilizing lipid reserves to meet energy requirements, the female can acquire sufficient protein from the diet to produce a large initial clutch even when foods are relatively scarce, whereas the renesting female must rely entirely upon food resources available at the breeding site for its nutrient and energy requirements.

The Auk

Parental care in Tundra Swans during the pre-fledgling period

Among studies that have quantified the care of precocial young, few have investigated forms of parental care other than vigilance. During the pre-fledging period, Tundra Swan (Cygnus columbianus columbianus) parents provided simultaneous biparental care by foraging near each other and their cygnets, and cygnets spent more time foraging during bouts in which both parents were foraging nearby than when only one parent was foraging nearby. Parents spent nearly twice as much foraging time on land than did non-parents, a habitat in which cygnets foraged more intensely than parents (i.e., spent more time foraging during foraging bouts) and could graze on protein-rich sedges rather than use more difficult below-water foraging methods. Parents also spent more than twice as much time being vigilant and more than three times as much time defending their territory than non-parents, behaviors that presumably benefited cygents by decreasing predation risk and indirect foraging competition, respectively. Parents therefore incurred the costs of foraging less intensely during foraging bouts, spending more time interacting, more time in vigilance, and less time sleeping/preening than non-parents.

Waterbirds

The mating behavior of Iguana iguana

Over a 19 month period I observed the social behaviors of individually recognized green iguanas, Iguana iguana, at three sites in the llanos of Venezuela. The behavior of iguanas outside the mating season differed from that seen during the mating season in three major ways: (1) during normal waking hours outside the breeding season, adult iguanas spent the majority of time immobile, apparently resting; (2) their interactions involved fewer high intensity displays; and (3) their day to day movements were often nomadic. During the mating season, one site was watched continuously during daylight hours (iguanas sleep throughout the night), allowing a complete count of all copulation attempts (N &equals; 250) and territorial interactions. At all sites, dominant males controlled access to small mating territories. Within the territories there did not appear to be any resources needed by females or their offspring. Thus, females could choose mates directly on the basis of male phenotype. Females aggregated in the mating territories of the largest males and mated preferentially with them. Territorial males copulated only once per day, although on several occasions more than one resident female was receptive on the same day. A few small nonterritorial males exhibited pseudofemale behavior (i.e., they abstained from sexual competition), but most nonterritorial males stayed on the periphery of mating territories and attempted to force copulations on unguarded females (peripheral male behavior). Uncooperative females were mounted by as many as three males simultaneously. Females resisted 95% of the 200 observed mating attempts by peripheral males, but only 56% of the attempts by territorial males (N &equals; 43). The selectivity of the females probably increased the genetic representation of the territorial males in the next generation. During the mating season females maintained a dominance hierarchy among themselves. Low ranked females tended to be excluded from preferred mating territories. In this system, both sexes may be subjected to sexual selection. I hypothesize that the ecological factors responsible for the unusual mating system are related to the lack of defendable resources, the iguana's folivory, and the high density of iguanas present in preferred mating areas.

Smithsonian Contributions to Zoology

Ecology and behaviour of the Alder Flycatcher Empidonax alnorum, Tyrannidae in winter

The Alder Flycatcher Empidonax alnorum (Tyrannidae) winters from Colombia and Venezuela, south to northern Argentina. We studied the species in Manu National Park, Madre de Dios, Peru. In this area the species occupies riparian zones along lowland meander rivers as well as adjacent transition zone forest in the floodplain. These habitats have a number of age-related longitudinal zones (parallel to the river margin) that are short term (I-2 year) transitory stages in the succession to floodplain forest. The fast growing plant species that make up these habitats are suitable for commercial harvest for paper pulp. We examined habitat use of the species in an attempt to identify the effects of such harvest and resulting habitat loss on these birds. The Alder Flycatcher favours two longitudinal zones where it establishes territories. The territories may be long-term or transitory, and are often occupied and defended by two individuals, generally of different age. One bird tends to be more active in territory defence than the other. Defence involves patrolling, vocal duels, and chases. Birds sally for insects but also eat large quantities of fruit.

Book chapter

Ecology and behaviour of the Alder Flycatcher Empidonax alnorum, Tyrannidae in winter

The Alder Flycatcher Empidonax alnorum (Tyrannidae) winters from Colombia and Venezuela, south to northern Argentina. We studied the species in Manu National Park, Madre de Dios, Peru. In this area the species occupies riparian zones along lowland meander rivers as well as adjacent transition zone forest in the floodplain. These habitats have a number of age-related longitudinal zones (parallel to the river margin) that are short term (I-2 year) transitory stages in the succession to floodplain forest. The fast growing plant species that make up these habitats are suitable for commercial harvest for paper pulp. We examined habitat use of the species in an attempt to identify the effects of such harvest and resulting habitat loss on these birds. The Alder Flycatcher favours two longitudinal zones where it establishes territories. The territories may be long-term or transitory, and are often occupied and defended by two individuals, generally of different age. One bird tends to be more active in territory defence than the other. Defence involves patrolling, vocal duels, and chases. Birds sally for insects but also eat large quantities of fruit.

Book chapter

Pre-nesting and nesting behavior of the Swainson's warbler

The Swainson?s Warbler is one of the least known of southern birds. Although fairly common in some parts of its summer range, observations of its breeding biology have been made by very few persons. The present study was conducted mostly at Macon, Georgia; Pendleton Ferry, Arkansas; and Dismal Swamp, Virginia....In central Georgia and east-central Arkansas, Swainson?s Warblers usually arrive on their territories during the first two weeks in April. Territories in several localities ranged in size from 0.3 to 4.8 acres. A color-marked Arkansas male occupied the same territory for at least four months. Hostile encounters between territorial male Swainson?s Warblers usually take place along the boundary of adjacent territories. Paired males were more aggressive than unpaired males. Toward the end of an encounter one of the two males would usually perform a display in which the wing and tail feathers were spread and the tail vibrated. Following boundary encounters males drifted back onto their territories and usually sang unbroken courses of songs for several minutes.....During pre-nesting at Macon, a mated pair spent the day mostly on the ground within 20 feet of each other, often foragin g 3 to 4 feet apart. What may have been a form of courtship display, in which the male flew from a perch down to the female and either pecked her rump or pounced on her, occurred about three times each hour throughout the day. During this period the male sang less than at other times during the breeding season.....First nests are usually built by the first week in May. Although other investigators reported finding nests of this species outside of the defended territory, all nests that I have found were within the territory. The large, bulky nest of this species usually is placed 2-6 feet above the ground. It is built by the female from materials gathered close to the nest site; and takes two or three days to complete.....Three and occasionally four white eggs are laid. The female incubates for 14-15 days. The Cowbird parasitizes nests in some parts of the breeding range of the Swainson?s Warbler.....During incubation the female spends about 78 per cent of daylight time on the nest. Both sexes feed young and clean nest. Young remain in nest from 10-12 days.

The Wilson Bulletin

Demography and movements of the endangered akepa and Hawaii creeper

We studied populations of the endangered Akepa (Loxops coccineus coccineus) and Hawaii Creeper (Oreomystis mana) at four sites on the island of Hawaii. Mean monthly density (? SL) of Akepa was 5.74 t 0.87, 1.3? 0.41, 0.96 -? 0.13, and 0.76 ? 0.12 Akepa/ha at Kau Forest, Hamakua, Keauhou Ranch, and Kilauea Forest study areas, respectively. Hawaii Creepers were found at densities of 1.68 ? 0.53, 1.79 ? 0.42, 0.48 ? 0.06, and 0.54 2 ? 0.08 birds/ha, respectively, at the four study areas. Highest capture rates and numbers of birds counted from stations occurred from August through November and February through March. Hatching-year birds were captured from May through December for Akepa and April through December for Hawaii Creeper. Annual survival for adults at Keauhou Ranch was 0.70 ? 0.27 SE for 61 Akepa and 0.73 ? 0.12 SE for 49 Hawaii Creepers. Lowest rates of mortality and emigration occurred between May and August. Both species appeared to defend Type-B territories typical of cardueline finches, retained mates for more than one year, and showed strong philopatry. Home ranges for Hawaii Creepers (X = 7.48 ha) were larger than those for Akepa (X = 3.94 No difference was found between home range sizes of males and females for either species.

The Wilson Bulletin

Aggressive interactions between the invasive Rio Grande cichlid (Herichthys cyanoguttatus) and native bluegill (Lepomis macrochirus), with notes on redspotted sunfish (Lepomis miniatus)

The Rio Grande cichlid (Herichthys cyanoguttatus) has been established in the Greater New Orleans Metropolitan area for at least 20 years, and its effect on native fishes is unknown. Behavioral trials were performed to determine if aggressive interactions occur between invasive H. cyanoguttatus and native bluegill (Lepomis macrochirus). When defending a territory as the resident, L. macrochirus were markedly aggressive, averaging 11.6 aggressive actions per lO-min behavioral trial. In contrast, L. macrochirus were extremely passive as invaders, with 0.5 aggressive actions per trial. Herichthys cyanoguttatus were equally aggressive as residents and as invaders, averaging 4.9 and 6.0 aggressive actions per trial, respectively. Herichthys cyanoguttatus interacted aggressively with native species whether they held territory or not, indicating that this invasive species may have fundamentally different strategies of aggression compared with native L. macrochirus. These differences may explain the continued success of H. cyanoguttatus as an invasive fish in southeastern Louisiana.

Journal of Ethology

Sidescan sonar examination of deteriorated revetments and bulkheads along Chicago's lake front

Lakefill for parks along Chicago's lake front is primarily defended with aged, deteriorated revetments. Survey by 100- and 500-kHz sidescan sonar documents that structural failure of the revetments is not related to lake-bottom undermining. The size and weight of the rock fill in the cribs is identified as a critical factor. Long-term effects of wave surge combined with ice action and gravity can remove rock fill either through breaks in the timber crib wall, or where revetment design offers lakeward exposure of the rock fill. Capstones are displaced when they lose underlying support. Revetments with a lakeward face of steel sheetpile, or toe protection mounded high against the structure, have no significant capstone displacement and are models for improved design. This study supports a rebuilding project that will bury the old revetments with new structures having a lakeward face of either steel sheetpile or a rubble mound.

Conference Paper