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Position of the Triassic-Jurassic boundary and timing of the end-Triassic extinctions on land: Data from the Moenave Formation on the southern Colorado Plateau, USA

Strata of the Moenave Formation on and adjacent to the southern Colorado Plateau in Utah–Arizona, U.S.A., represent one of the best known and most stratigraphically continuous, complete and fossiliferous terrestrial sections across the Triassic–Jurassic boundary. We present a synthesis of new biostratigraphic and magnetostratigraphic data collected from across the Moenave Formation outcrop belt, which extends from the St. George area in southwestern Utah to the Tuba City area in northern Arizona. These data include palynomorphs, conchostracans and vertebrate fossils (including footprints) and a composite polarity record based on four overlapping magnetostratigraphic sections. Placement of the Triassic–Jurassic boundary in strata of the Moenave Formation has long been imprecise and debatable, but these new data (especially the conchostracans) allow us to place the Triassic–Jurassic boundary relatively precisely in the middle part of the Whitmore Point Member of the Moenave Formation, stratigraphically well above the highest occurrence of crurotarsan body fossils or footprints. Correlation to marine sections based on this placement indicates that major terrestrial vertebrate extinctions preceded marine extinctions across the Triassic–Jurassic boundary and therefore were likely unrelated to the Central Atlantic Magmatic Province (CAMP) volcanism.

Arizona, Utah

Distribution of ore deposits and spectrographic analyses of some rocks and ores on the Colorado Plateau

The geographic pattern of known igneous rocks and ore deposits on the Colorado Plateau suggests a zonal arrangement of several types of ore deposits around centers of igneous activity. Spectrographic analyses of rocks and ores on the Plateau have been obtained in an effort to determine the distribution of elements and to examine the relationships between types of ore deposits and between the ore deposits and igneous rocks. Over 170 analyses of rocks and ores are given in this report. A preliminary study of these analyses suggests that the proportion of uranium, vanadium, copper, and silver in the uranium ores varies geographically, and that the pattern of variation may be in part concentric about some of the major laccolithic intrusions. It is also suggested that the following ratios of metals contained in the uranium ores are possible guides to larger-than-average ore deposits: (1) lead/uranium greater than 1, (2) lead/zinc greater than 10, and (3) zinc/geometric mean of cobalt and nickel less than 10.

Colorado

Federally owned coal and federal lands in the Colorado Plateau region

F ederally owned coal plays a major role in the energy supply of the United States. About 1.1 billion tons of coal were produced in the United States in 1997 (U.S. Department of Energy, 1998). About 30 percent of that total, or about 330 million tons, came from Federal lands. Almost all of the Federal coal production is from Wyoming, Montana, and three States in the Colorado Plateau Region—Utah, Colorado, and New Mexico.

Fact Sheet

Federally owned coal, federal lands, and coal quality in the Colorado Plateau Region

Federally owned coal plays a major role in the energy supply of the United States. About 1.1 billion tons of coal were produced in the United States in 1997 (U.S. Department of Energy, 1998). About 30 percent of that total, or about 330 million tons,1 came from Federal lands. (See USGS Fact Sheet FS-012-98.) Almost all of the Federal coal production is from Wyoming, Montana, and three States in the Colorado Plateau region—Utah, Colorado, and New Mexico2 (see table below).

Arizona, Colorado, New Mexico, Utah

Jurassic Lake T'oo'dichi': A large alkaline, saline lake, Morrison Formation, eastern Colorado Plateau

A large alkaline, saline lake, Lake T'oo'dichi', occupied the entire eastern part of the Colorado Plateau region during deposition of the Brushy Basin Member of the Upper Jurassic Morrison Formation. The lake extended from near the site of Albuquerque, New Mexico, to near the site of Grand Junction, Colorado, and occupied a region that encompassed the San Juan and ancestral Paradox basins, making it the largest ancient alkaline, saline lake known.

Geological Society of America Bulletin

Generalizing ecological site concepts of the Colorado Plateau for landscape-level applications

Numerous ecological site descriptions in the southern Utah portion of the Colorado Plateau can be difficult to navigate, so we held a workshop aimed at adding value and functionality to the current ecological site system. We created new groups of ecological sites and drafted state-and-transition models for these new groups. We were able to distill the current large number of ecological sites in the study area (ca. 150) into eight ecological site groups that capture important variability in ecosystem dynamics. Several inventory and monitoring programs and landscape scale planning actions will likely benefit from more generalized ecological site group concepts.

Rangelands

High-resolution receiver function imaging reveals Colorado Plateau lithospheric architecture and mantle-supported topography

After maintaining elevations near sea level for over 500 million years, the Colorado Plateau (CP) has a present average elevation of 2 km. We compute new receiver function images from the first dense seismic transect to cross the plateau that reveal a central CP crustal thickness of 42–50 km thinning to 30–35 km at the CP margins. Isostatic calculations show that only approximately 20% of central CP elevations can be explained by thickened crust alone, with the CP edges requiring nearly total mantle compensation. We calculate an uplift budget showing that CP buoyancy arises from a combination of crustal thickening, heating and alteration of the lithospheric root, dynamic support from mantle upwelling, and significant buoyant edge effects produced by small-scale convecting asthenosphere at its margins.

Arizona, Colorado, New Mexico, Texas, Utah

Lungfish burrows in the Upper Triassic Chinle and Dolores Formations, Colorado Plateau

Vertical-to-inclined, cylindrical trace fossils that occur in the Upper Triassic Chinle and Dolores Formations on the Colorado Plateau are interpreted to be the casts of lungfish burrows. The casts, which are as much as 11 cm in diameter and as much as 1.6 m long, were formed by passive silicilastic and carbonate sedimentation into apparently abandoned lungfish burrows. Locally, the burrow fillings are overwhelmingly abundant, and many intersect and have destroyed former burrow fillings. Superposition of bioturbation episodes has obliterated most primary sedimentary structures. This bioturbation has contributed to the mottled coloration and the knobby-weathering texture of the rocks. The burrow-fillings occur ubiquitously in three lithofacies, comprising 1) purple- and white-mottled, silicified sandstone and siltstone, 2) red and brown siltstone and mudstone, and 3) pink and green limestone. These strata were deposited in a continental environment that included fluvial channels and floodplains, sand sheets and playa mudflats, and lacustrine basins, marshes, and deltas. The identification of the trace fossils as the positive casts of lungfish burrows is based on their morphologic similarity to previously identified lungfish burrows and to available hand specimens. The widespread occurrence of the lungfish burrows in the Chinle and Dolores Formations attests to the extensive habitat that supported lungfish in the Late Triassic and to conditions favorable for burrow preservation. Analogy with the environments that support modern lungfish populations suggests that the Late Triassic climate in the study area provided sufficient moisture to support large populations of lungfish and that this climate was probably punctuated by seasonally dry periods.

Journal of Sedimentary Petrology

Small-Scale Vertical Distribution of Bacterial Biomass and Diversity in Biological Soil Crusts from Arid Lands in the Colorado Plateau

We characterized, at millimeter resolution, bacterial biomass, diversity, and vertical stratification of biological soil crusts in arid lands from the Colorado Plateau. Microscopic counts, extractable DNA, and plate counts of viable aerobic copiotrophs (VAC) revealed that the top centimeter of crusted soils contained atypically large bacterial populations, tenfold larger than those in uncrusted, deeper soils. The plate counts were not always consistent with more direct estimates of microbial biomass. Bacterial populations peaked at the immediate subsurface (1-2 mm) in light-appearing, young crusts, and at the surface (0-1 mm) in well-developed, dark crusts, which corresponds to the location of cyanobacterial populations. Bacterial abundance decreased with depth below these horizons. Spatially resolved DGGE fingerprints of Bacterial 16S rRNA genes demonstrated the presence of highly diverse natural communities, but we could detect neither trends with depth in bacterial richness or diversity, nor a difference in diversity indices between crust types. Fingerprints, however, revealed the presence of marked stratification in the structure of the microbial communities, probably a result of vertical gradients in physicochemical parameters. Sequencing and phylogenetic analyses indicated that most of the naturally occurring bacteria are novel types, with low sequence similarity (83-93%) to those available in public databases. DGGE analyses of the VAC populations indicated communities of lower diversity, with most types having sequences more than 94% similar to those in public databases. Our study indicates that soil crusts represent small-scale mantles of fertility in arid ecosystems, harboring vertically structured, little-known bacterial populations that are not well represented by standard cultivation methods.

Microbial Ecology

Comparison of the isotopic abundance of U235 and U238 and the radium activity ratios in Colorado Plateau uranium ores

The isotopic abundances of uranium and the radium activity ratios of eleven samples of uranium ore from the Colorado Plateau have been measured. No significant variation in the isotopic abundance of the uranium was noted; with'in the experimental error, the average U 235 /U 238 ratio is 137.7. There is a significant variation in the Ra 226 /Ra 223 activity ratios (0.048−0.143), which indicates a relatively recent alteration of the ore samples. The variations do not, however, explain the lead-uranium and lead-lead age discrepancies.

Geochimica et Cosmochimica Acta

Insights into post-Miocene uplift of the western margin of the Colorado Plateau from the stratigraphic record of the lower Colorado River

The spatial and temporal distribution of Pliocene to Holocene Colorado River deposits (southwestern USA and northwestern Mexico) form a primary data set that records the evolution of a continental-scale river system and helps to delineate and quantify the magnitude of regional deformation. We focus in particular on the age and distribution of ancestral Colorado River deposits from field observations, geologic mapping, and subsurface studies in the area downstream from Grand Canyon (Arizona, USA). A new 4.73 ± 0.17 Ma age is reported for a basalt that flowed down Grand Wash to near its confluence with the Colorado River at the eastern end of what is now Lake Mead (Arizona and Nevada). That basalt flow, which caps tributary gravels, another previously dated 4.49 ± 0.46 Ma basalt flow that caps Colorado River gravel nearby, and previously dated speleothems (2.17 ± 0.34 and 3.87 ± 0.1 Ma) in western Grand Canyon allow for the calculation of long-term incision rates. Those rates are ~90 m/Ma in western Grand Canyon and ~18–64 m/Ma in the eastern Lake Mead area. In western Lake Mead and downstream, the base of 4.5–3.5 Ma ancestral Colorado River deposits, called the Bullhead Alluvium, is generally preserved below river level, suggesting little if any bedrock incision since deposition. Paleoprofiles reconstructed using ancestral river deposits indicate that the lower Colorado River established a smooth profile that has been graded to near sea level since ca. 4.5 Ma. Steady incision rates in western Grand Canyon over the past 0.6–4 Ma also suggest that the lower Colorado River has remained in a quasi–steady state for millions of years with respect to bedrock incision. Differential incision between the lower Colorado River corridor and western Grand Canyon is best explained by differential uplift across the Lake Mead region, as the overall 4.5 Ma profile of the Colorado River remains graded to Pliocene sea level, suggesting little regional subsidence or uplift. Cumulative estimates of ca. 4 Ma offsets across faults in the Lake Mead region are similar in magnitude to the differential incision across the area during the same approximate time frame. This suggests that in the past ~4 Ma, vertical deformation in the Lake Mead area has been localized along faults, which may be a surficial response to more deep-seated processes. Together these data sets suggest ~140–370m of uplift in the past 2–4 Ma across the Lake Mead region.

Arizona, California, Nevada

Appraisal of the accuracy of U.S. Geological Survey ore reserve estimates for uranium-vanadium deposits on the Colorado Plateau

The U.S. Geological Survey has made estimates of the reserves of uranium and vanadium in the carnotite deposits explored by Geological Survey drilling on the Colorado Plateau. This report presents an appraisal of the accuracy of the reserve estimates for deposits in the Uravan mineral belt, the causes of inaccuracy, and the significance of the estimates in terms of the total known reserves of the region.

Colorado

Sandstone-hosted uranium deposits of the Colorado Plateau, USA

More than 4,000 sandstone-hosted uranium occurrences host over 1.2 billion pounds of mined and in situ U 3 O 8 throughout the Colorado Plateau. Most of the resources are in two distinct mineral systems with deposits hosted in the Triassic Chinle and Jurassic Morrison Formations. In the Chinle mineral system, base metal sulfides typically accompany mineralization. The Morrison mineral system is characterized by V/U ratios up to 20. The uranium source was likely volcanic ash preserved as bentonitic mudstones in the Brushy Basin Member of the Morrison Formation, and lithic volcanic clasts, ash shards, and bentonitic clay in the lower part of the Chinle Formation. Vanadium originated from two possible sources: iron–titanium oxides that are extensively altered in bleached rock near deposits or from similar minerals in variably bleached red beds interbedded with and beneath the Morrison. In Chinle-hosted deposits, in addition to volcanic ash, a contributing source of both vanadium and uranium is proposed here for the first time to be underlying red beds in the Moenkopi and Cutler Formations that have undergone a cycle of reddening-bleaching-reoxidation. Transport in both systems was likely in groundwater through the more permeable sandstones and conglomerate units. The association of uranium minerals with carbonate and more rarely apatite, suggests that transport of uranium was as a carbonate or phosphate complex. The first comprehensive examination of paleoclimate, paleotopography, and subsurface structure of aquifers coupled with analysis of the geochronology of deposits suggests that that there were distinct pulses of uranium mineralization/redistribution during the period from about 259 Ma to 12 Ma when oxidized mineralizing fluids were intermittently rejuvenated in the Plateau in response to changes in tectonic regime and climate. Multiple lines of evidence indicate that deposits formed at ambient temperatures of about 25 °C to no greater than about 140 °C. In both systems, deposits formed where groundwater flow slowed and was subject to evaporative concentration. Stagnant conditions allowed for prolonged interaction of U- and V-enriched groundwater with ferrous iron-bearing reductants, such as illite and iron–titanium oxides, and more rarely organic material such as plant debris. Paragenetically late in the sequence, reducing fluids introduced additional organic matter to some deposits. Reducing fluids and introduced organic matter (now amorphous and altered by radiolysis) may originate from regional petroleum systems where peak oil and gas generation was from ∼ 82 to ∼ 5 Ma. Our novel analysis indicates that these reducing fluids bleached rock and protected affected deposits from remobilization during exposure and weathering that followed uplift of the Plateau (∼80 to 40 Ma).

Arizona, Colorado, Nevada, New Mexico, Utah

Molluscan paleoecology and sedimentation patterns of the Cenomanian-Turonian extinction interval in the southern Colorado Plateau region

Lithology, carbonate and organic carbon geochemistry, and molluscan macrofauna were investigated in fine-grained clay-dominated strata spanning the Cenomanian-Turonian Stage boundary in the southern Colorado Plateau region. Analyses of these data show basinal and onshore to offshore changes in sedimentologic and molluscan assemblage trends through the upper Cenomanian Sciponoceras gracile and Neocardioceras juddii and lower Turonian Watinoceras coloradoense Biozones. Diverse infaunal/epifaunal bivalve assemblages are best developed in the Sciponoceras gracile Biozone and at the more clay-rich nearshore sections. Gastropod and infaunal bivalve abundance and diversity decrease in an offshore and up-section direction as carbonate and organic carbon percentages generally increase. The Neocardioceras juddii Biozone precedes the stage boundary and is characterized in Utah and Arizona by pulsed increases in organic carbon and detrital-feeding gastropods, and by a shift in infaunal bivalve dominance toward species adapted for soft-substrate conditions. These data suggest that pulses of increased organic detrital resources and decreased substrate firmness were associated with the stage boundary extinctions. The stage boundary is marked by a change to depauperate epifaunal assemblages at western sections. At all sections, the base of the Watinoceras coloradoense Biozone is reflected by an abrupt increase in carbonate and decrease in organic carbon percentages as well as by the immigration of Mytiloides bivalves. Periods of progressive recovery through this biozone were associated with carbonate-rich depositional periods.

Arizona, Colorado, New Mexico, Utah

Chemical composition as a guide to the size of sandstone-type uranium deposits in the Morrison Formation on the Colorado Plateau

The concentrations of uranium, yttrium, sodium, iron, zirconium, manganese, calcium, and nickel in 75 mill-pulp samples of uranium deposits in the Salt Wash member of the Morrison formation on the Colorado Plateau have been found, by statistical tests, to be significantly related to the size of the deposits represented by the samples. The elements mentioned above are related to the formation of the deposits in a variety of ways. Zirconium is an intrinsic element, contained principally in the detrital syngenetic fraction of the host sandstone. Calcium, manganese, and sodium are intrinsic elements contained principally in epigenetic (diagenetic) carbonate in the host sandstone. Uranium, yttrium, and nickel are principally extrinsic elements, introduced into the host sandstone by uranium mineralization or related processes. Somewhat more than half of the iron is probably intrinsic and the remainder is extrinsic. Three methods can be used to estimate the size of uranium deposits in the Salt Wash, member within broad limits. Method 1 is based on simple linear-regression theory; method 2 is based on multiple-regression theory (long method); and method 3, on multiple-regression theory (short method). For methods 1 and 2 the estimated log size of each deposit can be computed from tables showing the known concentration of uranium, yttrium, sodium, iron, zirconium, manganese, calcium, and nickel in the deposits, as determined by semiquantitative spectrographic analysis. For method 3 the estimated size or log size can be read directly from a table showing known concentration of uranium and yttrium only. About 80 percent of the tonnage-size estimates from method 1 will be within a factor of 13 (12-14) of the true sizes. The precision of the size estimates from method 2, the long multiple-regression method, is highly variable. Some estimates from method 2 will be within a factor of 12 of the true size at the 80-percent confidence level; others, within a factor of 40 at the 80-percent confidence level. About 80 percent of the tonnage-size estimates from method 3, the short multiple-regression method, will be within a factor of 15 (13-16) of the true size. A group of 40 deposits of known size was used to test the theoretical derivation of the confidence intervals given above. It was concluded from the test that the confidence intervals describe the precision of the methods correctly. The methods for estimating the size of uranium deposits are useful where the ore is poorly exposed or where an independent estimate is desired. The error of the estimates may be quite large, as indicated by the confidence limits given above; but the estimates can be used to, at least, distinguish very large from very small deposits. They also may serve to encourage or discourage further expenditures in the development and exploration of ore bodies. The methods of size estimation can be particularly useful in attempts to appraise or compare groups of deposits or mining districts, inasmuch as the average estimate of size of deposits in a group is more precise than any single estimate. The methods for estimating size are established only for deposits in the Salt Wash member of the Morrison formation. Tests indicate that the equations calculated for deposits in the Salt Wash fail completely if applied to deposits in other stratigraphic units, such as the Moss Back and Shinarump members of the Chinle formation. A further restriction, not completely evaluated at present, is that semiquantitative spectrographic analyses of mill-pulp samples are required. No tests have been made to determine the precision and accuracy of the methods when other types of samples, such as drill core, are used.

Arizona, Colorado, New Mexico, Utah

Field Evaluations of Sampling Methods for Long-Term Monitoring of Upland Ecosystems on the Colorado Plateau

To inform planning for long-term ecological monitoring, we sampled vegetation and soil-surface attributes across a range of terrestrial ecosystems (physiognomic types) in seven National Park Service units on the Colorado Plateau. Primary objectives were (1) to evaluate a suite of sampling methods according to measures of repeatability, efficiency, and impacts on plot conditions; and (2) to characterize within- and among-plot variability in monitoring measures. This work was designed to support NPS staff in selecting the combination of methods that best meets their monitoring objectives and resource constraints. We found no differences among cover-estimation techniques in terms of repeatability between observers (measurement precision). Estimates for total live understory canopy cover, cover of individual species, and cover of soil-surface features were highly repeatable between observers for 10-m2 quadrats, 1-m2 quadrats, and line-point intercept sampling methods. Estimates of shrub and tree density in 10-m2 quadrats also were repeatable between observers, although sample sizes for were small for many species. At 10 of 11 ecological sites, we found that sampling with 10-m2 quadrats was the most efficient cover-estimation technique with respect to within-plot variability in cover estimates and numbers of subsamples required to estimate plot-level cover with 20 percent precision. According to these same measures, sampling with 1-m2 quadrats was the least efficient cover-estimation technique at eight of 11 ecological sites. The line-point technique was most efficient at eight of 11 ecological sites in terms of the amount of time required to estimate total plot-level cover with 20 percent precision - largely because 10-m2 quadrats were more time consuming and 1-m2 quadrats had greater within-plot variability relative to line-point sampling. However, there was no statistical difference among methods with respect to median subsampling times for 20 percent precision. There also were no differences among methods with respect to mean and median measures of among-plot variability in total live understory canopy cover. But among-plot variability was least for the line-point technique at seven of 11 ecological sites. Sampling activities had greatest impacts on plot conditions at macroplots where there was a high degree of cover by biological and physical soil crusts. Of all sampling procedures, 10-m2 quadrat sampling, line-point sampling, and gap-intercept sampling had the most impacts on soil conditions due to trampling of soil crusts by the field team.

Open-File Report

Geologic guides to prospecting for carnotite deposits on the Colorado Plateau

This report describes the geologic features that can be used to appraise the favorability of ground in guiding diamond-drill exploration for carnotite deposits in the Upper Jurassic Morrison formation on the Colorado Plateau. It is based on a statistical study of the geologic logs of about 2,500 holes drilled by the Geological Survey. The most useful features consist of the thickness of the ore-bearing sandstone, the color of the ore-bearing sandstone, the altered mudstone associated with the ore-bearing sandstone, and the abundance of carbonaceous material in the ore-bearing sandstone. Although each feature can be used alone to appraise the favorability of ground, an appraisal based on all of them together is even more useful. A method of expressing this favorability in numerical values is suggested. The results obtained by the Geological Survey using these geologic guides appear to be at least twice as favorable as the drilling results obtained with little or no geologic guidance.

Colorado

Field-based evaluations of sampling techniques to support long-term monitoring of riparian ecosystems along wadeable streams on the Colorado Plateau

To better plan for and implement long-term ecological monitoring, we measured riparian vegetation and fluvial geomorphic features at pilot study sites on four wadeable perennial stream reaches, representative of drainages across the Colorado Plateau. Our primary objectives were to (1) collect field data, (2) evaluate the efficiency and effectiveness of various ecological measures and measurement techniques for riparian ecosystems, and (3) use field-based sampling to inform and refine the development of standard operating procedures for use in implementing integrated, long-term monitoring of riparian ecosystems. Ultimately, this work was aimed at providing NPS staff with some of the information and methods needed to design and implement long-term monitoring of NPS riparian resources, which is both relevant to management, and fully operational within institutional resource constraints. Our results suggest that selecting sampling reaches and establishing a sampling frame of 11 transects, across a range of stream types, is feasible given a limited set of decision rules. A distinctive feature of richness across all sites was the high percentage of rare species, defined here as species having a single occurrence at a site. Rare species represented from 33 percent to 47 percent of the species total across the four pilot sites. Our data show that the two smallest quadrat sizes, 0.01 m 2 and 0.1 m 2 , rarely had any species that occurred in the desired frequency range and can be omitted from the monitoring protocol. Few species fell within the 30&ndash;70 percent range in the 1-m 2 quadrats, but this quadrat size appears to be useful at the Tsaile Creek (CACH) site. We recommend continuing to collect information at the 1-m 2 scale and reevaluating its usefulness after more data are available from different types of sites. The 10-m 2 quadrat is adequate for monitoring changes in frequencies of very common species at all sites. Based on pilot study results, we conclude that at sites with low total species numbers (< 60 species), 40&ndash;60, 10-m 2 quadrats, would be sufficient to characterize overall species diversity for relatively common species. At sites with higher total numbers of species (> 100), 60&ndash;80, 10-m 2 quadrats would be required to characterize overall species diversity. Rare species of interest should be monitored using alternative approaches, such as a site inventory and/or mapping (see Elzinga and others, 1998). A large number of the systematically placed 10-m 2 quadrats span two or more geomorphic surfaces, especially adjacent to the channel. This makes resolution of species affinities with distinct geomorphic landforms difficult. Thus, we provide an amendment to improve characterization of herbaceous and shrub species on narrow, near-channel surfaces by sampling additional 0.5-m by 1-m quadrats on those surfaces. It appears that for sites in narrow valley settings where riparian zones average less than approximately 40 m, the number of 10-m 2 quadrats systematically placed on 11 transects will not provide shrub cover estimates at 20 percent precision. In such cases, additional sample reaches should be added in order to attain a minimum of 130 to 140 10-m 2 shrub quadrats. The line-intercept technique can provide a relatively rapid, reach-scale quantification of proportional cover for woody vegetation and geomorphic surface types and that variance in these measures stabilizes by the eighth or ninth transect sampled. An overlay of the distribution of geomorphic surface data derived from line-intercept sampling on topographic survey information indicates that delineation of geomorphic surfaces could be done in conjunction with the topographic survey of each transect, obviating the need to record surface breaks using the line intercept. To include geomorphic surface identifications with the topographic survey, surface breaks and transitional surfaces should be included and identified in the survey, in addition to systematically placed survey points. Compared to 5-m by 20-m tree quadrats, belt transects were shown to provide similar estimates of stand structure (stem density and stand basal area) in less than 30 percent of the time. Further, for the streams sampled, there were no statistically significant differences in stem density and basal area estimates between 10-m and 20-m belt transects and the smaller belts took approximately half the time to sample. There was, however, high variance associated with estimates of stand structure for infrequently occurring stems, such as large, relict or legacy riparian trees. Legacy riparian trees occurred in limited numbers at all sites sampled. A reachscale population census of these trees indicated that the 10-m belt transects tended to underestimate both stem density and basal area for these riparian forest elements and that a complete reach-scale census of legacy trees averaged less than one hour per site.

Colorado Plateau