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Climatic effects of 30 years of landscape change over the Greater Phoenix, Arizona, region: 1. Surface energy budget changes

This paper is part 1 of a two-part study that evaluates the climatic effects of recent landscape change for one of the nation's most rapidly expanding metropolitan complexes, the Greater Phoenix, Arizona, region. The region's landscape evolution over an approximate 30-year period since the early 1970s is documented on the basis of analyses of Landsat images and land use/land cover (LULC) data sets derived from aerial photography (1973) and Landsat (1992 and 2001). High-resolution, Regional Atmospheric Modeling System (RAMS), simulations (2-km grid spacing) are used in conjunction with consistently defined land cover data sets and associated biophysical parameters for the circa 1973, circa 1992, and circa 2001 time periods to quantify the impacts of intensive land use changes on the July surface temperatures and the surface radiation and energy budgets for the Greater Phoenix region. The main findings are as follows: since the early 1970s the region's landscape has been altered by a significant increase in urban/suburban land area, primarily at the expense of decreasing plots of irrigated agriculture and secondarily by the conversion of seminatural shrubland. Mean regional temperatures for the circa 2001 landscape were 0.12??C warmer than the circa 1973 landscape, with maximum temperature differences, located over regions of greatest urbanization, in excess of 1??C. The significant reduction in irrigated agriculture, for the circa 2001 relative to the circa 1973 landscape, resulted in dew point temperature decreases in excess of 1??C. The effect of distinct land use conversion themes (e.g., conversion from irrigated agriculture to urban land) was also examined to evaluate how the most important conversion themes have each contributed to the region's changing climate. The two urbanization themes studied (from an initial landscape of irrigated agriculture and seminatural shrubland) have the greatest positive effect on near-surface temperature, increasing maximum daily temperatures by 1??C. Overall, sensible heat flux differences between the circa 2001 and circa 1973 landscapes result in a 1 W m -2 increase in domain-wide sensible heating, and a similar order of magnitude decrease in latent heating, highlighting the importance of surface repartitioning in establishing near-surface temperature trends. In part 2 of this study, we address the role of the surface budget changes on the mesoscale dynamics/thermodynamics, in context of the large-scale environment. Copyright 2009 by the American Geophysical Union.

Journal of Geophysical Research D: Atmospheres

The effects of landscape position on plant species density: Evidence of past environmental effects in a coastal wetland

Here we propose that an important cause of variation in species density may be prior environmental conditions that continue to influence current patterns. In this paper we investigated the degree to which species density varies with location within the landscape, independent of contemporaneous environmental conditions. The area studied was a coastal marsh landscape subject to periodic storm events. To evaluate the impact of historical effects, it was assumed that the landscape position of a plot relative to the river's mouth ('distance from sea') and to the edge of a stream channel ('distance from shore') would correlate with the impact of prior storm events, an assumption supported by previous studies. To evaluate the importance of spatial location on species density, data were collected from five sites located at increasing distances from the river's mouth along the Middle Pearl River in Louisiana. At each site, plots were established systematically along transects perpendicular to the shoreline. For each of the 175 Plots, we measured elevation, soil salinity, percent of plot recently disturbed, percent of sunlight captured by the plant canopy (as a measure of plant abundance), and plant species density. Structural equation analysis ascertained the degree to which landscape position variables explained variation in species density that could not be explained by current environmental indicators. Without considering landscape variables, 54% of the variation in species density could be explained by the effects of salinity, flooding, and plant abundance. When landscape variables were included, distance from shore was unimportant but distance from sea explained an additional 12% of the variance in species density (R2 of final model = 66%). Based on these results it appears that at least some of the otherwise unexplained variation in species density can be attributed to landscape position, and presumably previous storm events. We suggest that future studies may gain additional insight into the factors controlling current patterns of species density by examining the effects of position within the landscape.

Écoscience

How landscape dynamics link individual- to population-level movement patterns: A multispecies comparison of ungulate relocation data

Aim To demonstrate how the interrelations of individual movements form large-scale population-level movement patterns and how these patterns are associated with the underlying landscape dynamics by comparing ungulate movements across species. Locations Arctic tundra in Alaska and Canada, temperate forests in Massachusetts, Patagonian Steppes in Argentina, Eastern Steppes in Mongolia. Methods We used relocation data from four ungulate species (barren-ground caribou, Mongolian gazelle, guanaco and moose) to examine individual movements and the interrelation of movements among individuals. We applied and developed a suite of spatial metrics that measure variation in movement among individuals as population dispersion, movement coordination and realized mobility. Taken together, these metrics allowed us to quantify and distinguish among different large-scale population-level movement patterns such as migration, range residency and nomadism. We then related the population-level movement patterns to the underlying landscape vegetation dynamics via long-term remote sensing measurements of the temporal variability, spatial variability and unpredictability of vegetation productivity. Results Moose, which remained in sedentary home ranges, and guanacos, which were partially migratory, exhibited relatively short annual movements associated with landscapes having very little broad-scale variability in vegetation. Caribou and gazelle performed extreme long-distance movements that were associated with broad-scale variability in vegetation productivity during the peak of the growing season. Caribou exhibited regular seasonal migration in which individuals were clustered for most of the year and exhibited coordinated movements. In contrast, gazelle were nomadic, as individuals were independently distributed and moved in an uncoordinated manner that relates to the comparatively unpredictable (yet broad-scale) vegetation dynamics of their landscape. Main conclusions We show how broad-scale landscape unpredictability may lead to nomadism, an understudied type of long-distance movement. In contrast to classical migration where landscapes may vary at broad scales but in a predictable manner, long-distance movements of nomadic individuals are uncoordinated and independent from other such individuals. Landscapes with little broad-scale variability in vegetation productivity feature smaller-scale movements and allow for range residency. Nomadism requires distinct integrative conservation strategies that facilitate long-distance movements across the entire landscape and are not limited to certain migration corridors.

Alaska, Massachusetts

Analysis of predator movement in prairie landscapes with contrasting grassland composition

Mammalian predation influences waterfowl breeding success in the U.S. northern Great Plains, yet little is known about the influence of the landscape on the ability of predators to find waterfowl nests. We used radiotelemetry to record nightly movements of red foxes ( Vulpes vulpes ) and striped skunks ( Mephitis mephitis ) in two 41.4-km 2 study areas in North Dakota. Study areas contained either 15–20% grassland (low grassland composition) or 45–55% grassland (high grassland composition). Grasslands included planted cover, pastureland, and hayland. We predicted that the type and composition of cover types in the landscape would influence both predator movement across the landscape (as measured by the fractal dimension and displacement ratio) as well as localized movement (as measured by the rate of movement and turning angle between locations) within patches of different cover types. Red fox movements were straighter (lower fractal dimensions and higher displacements) across landscapes with a low grassland composition, indicating directed movement between the more isolated patches of planted cover. Striped skunk movements did not differ between landscape types, illustrating their movement along wetland edges, which had similar compositions in both landscape types. The high variability in turning angles by red fox in planted cover and pastureland in both landscape types is consistent with restricted-area foraging. The high rate of movement by red foxes in planted cover and by striped skunks in wetland edges suggests that spatial memory may influence movement patterns. Understanding the behavior of predators in fragmented prairie landscape is essential for managing breeding habitat for grassland birds and for predicting the spatial and temporal dynamics of predators and their prey.

North Dakota

Associations of grassland birds with landscape factors in southern Wisconsin

We investigated the association of grassland birds with field- and landscape-level habitat variables in south-central Wisconsin during 1985–1987. Landscape-level variables were measured and digitized at 200, 400 and 800 m from the perimeter of 38 200 m × 100 m strip transects. A mixture of field and landscape variables was associated with the density of savannah sparrow ( Passerculus sandwichensis ) and grasshopper sparrow ( Ammodramus savannarum ). Only landscape variables were associated with the density of bobolink ( Dolichonyx oryzivorus ), eastern meadowlark ( Sturnella magna ) and all birds that were grassland species of management concern. Field size was not an important predictor of bird density. Cover-type diversity of the surrounding area was commonly selected in the models for three species and all birds that were grassland species of management concern. Higher bird densities in the transects were associated with landscapes where the cover types were less diverse. Landscapes with low cover type diversity were dominated by grassland, pasture and hay. Field habitat, mean patch size of cover types and distance to woody vegetation were the next most common predictors of avian density. The density of some grassland birds increased as nonlinear woody features such as woodlots and shrub carrs decreased in patch size, decreased in total amount in the landscape and increased in distance from a transect. However, density of other species was positively associated with linear woody features such as the total amount and nearness of hedgerows. The composition of the surrounding landscape, at least out to 800 m, is important in grassland bird management.

Wisconsin

Effects of landscape features on waterbird use of rice fields

Literature is reviewed to determine the effects of landscape features on waterbird use of fields in regions where rice (Oryza sativa) is grown. Rice-growing landscapes often consist of diverse land uses and land cover, including rice fields, irrigation ditches, other agricultural fields, grasslands, forests and natural wetlands. Numerous studies indicate that local management practices, such as water depth and timing of flooding and drawdown, can strongly influence waterbird use of a given rice field. However, the effects of size and distribution of rice fields and associated habitats at a landscape scale have received less attention. Even fewer studies have focused on local and landscape effects simultaneously. Habitat connectivity, area of rice, distance to natural wetlands, and presence and distance to unsuitable habitat can be important parameters influencing bird use of rice fields. However, responses to a given landscape vary with landscape structure, scale of analysis, among taxa and within taxa among seasons. A lack of multi-scale studies, particularly those extending beyond simple presence and abundance of a given species, and a lack of direct tests comparing the relative importance of landscape features with in-field management activities limits understanding of the importance of landscape in these systems and hampers waterbird conservation and management.

Waterbirds

Does Mallard clutch size vary with landscape composition?

We studied Mallards ( Anas platyrhynchos ) nesting in artificial nesting structures in northeastern North Dakota and compared clutch size between landscapes where proportion of cropland was either high (mean = 68.9%, cropland landscapes) or low (mean = 30.2%, grassland landscapes). Mallard clutch size was significantly related to nest initiation date and landscape composition. Mean clutch size, controlled for nest initiation date, was 1.24 ± 0.33 SE eggs smaller on cropland landscapes than on grassland landscapes. Generality of this pattern across space, time, and type of nesting sites is unknown, as is causation. Demographic importance of variation in clutch size may be influenced by covariation with other demographic variables, such as nest success and abundance of breeding pairs, which also are negatively correlated with landscape proportion of cropland. We suggest that researchers examine relationships between clutch size and landscape composition in both structure-nesting and ground-nesting Mallards, in other geographic areas, and in other duck species.

The Wilson Bulletin

Landscape characteristics influence pond occupancy by frogs after accounting for detectability

Many investigators have hypothesized that landscape attributes such as the amount and proximity of habitat are important for amphibian spatial patterns. This has produced a number of studies focusing on the effects of landscape characteristics on amphibian patterns of occurrence in patches or ponds, most of which conclude that the landscape is important. We identified two concerns associated with these studies: one deals with their applicability to other landscape types, as most have been conducted in agricultural landscapes; the other highlights the need to account for the probability of detection. We tested the hypothesis that landscape characteristics influence spatial patterns of amphibian occurrence at ponds after accounting for the probability of detection in little-studied peatland landscapes undergoing peat mining. We also illustrated the costs of not accounting for the probability of detection by comparing our results to conventional logistic regression analyses. Results indicate that frog occurrence increased with the percent cover of ponds within 100, 250, and 1000 m, as well as the amount of forest cover within 1000 m. However, forest cover at 250 m had a negative influence on frog presence at ponds. Not accounting for the probability of detection resulted in underestimating the influence of most variables on frog occurrence, whereas a few were overestimated. Regardless, we show that conventional logistic regression can lead to different conclusions than analyses accounting for detectability. Our study is consistent with the hypothesis that landscape characteristics are important in determining the spatial patterns of frog occurrence at ponds. We strongly recommend estimating the probability of detection in field surveys, as this will increase the quality and conservation potential of models derived from such data.

Ecological Applications

Beyond the edge: Linking agricultural landscapes, stream networks, and best management practices

Despite much research and investment into understanding and managing nutrients across agricultural landscapes, nutrient runoff to freshwater ecosystems is still a major concern. We argue there is currently a disconnect between the management of watershed surfaces (agricultural landscape) and river networks (riverine landscape). These landscapes are commonly managed separately, but there is limited cohesiveness between agricultural landscape-focused research and river science, despite similar end goals. Interdisciplinary research into stream networks that drain agricultural landscapes is expanding but is fraught with problems. Conceptual frameworks are useful tools to order phenomena, reveal patterns and processes, and in interdisciplinary river science, enable the joining of multiple areas of understanding into a single conceptual–empirical structure. We present a framework for the interdisciplinary study and management of agricultural and riverine landscapes. The framework includes components of an ecosystems approach to the study of catchment–stream networks, resilience thinking, and strategic adaptive management. Application of the framework is illustrated through a study of the Fox Basin in Wisconsin, USA. To fully realize the goal of nutrient reduction in the basin, we suggest that greater emphasis is needed on where best management practices (BMPs) are used within the spatial context of the combined watershed–stream network system, including BMPs within the river channel. Targeted placement of BMPs throughout the riverine landscape would increase the overall buffering capacity of the system to nutrient runoff and thus its resilience to current and future disturbances.

Journal of Environmental Quality

A landscape indicator approach to the identification and articulation of the consequences of land-cover change in the Mid-Atlantic Region, 1973-2001

Landscape indicators, derived from land-use and land-cover data, hydrology, nitrate deposition, and elevation data, were used by Jones and others (2001a) to calculate the ecological consequences of land-cover change. Nitrate loading and physical bird habitat were modeled from 1973 and 1992 land-cover and other spatial data for the Mid-Atlantic region. Utilizing the same methods, this study extends the analysis another decade with the use of the 2001 National Land Cover Dataset. Land-cover statistics and trends are calculated for three time periods: 1973-1992, 1992-2001 and 1973-2001. In addition, high-resolution aerial photographs (1 meter or better ground-sample distance) were acquired and analyzed for thirteen pairs of adjacent USGS 7.5 minute quadrangle maps in areas where distinct positive or negative changes to nitrogen loading and bird habitat were previously calculated. During the entire 30 year period, the data show that there was extensive loss of agriculture and forest area and a major increase in urban land-cover classes. However, the majority of the conversion of other classes to urban occurred during the 1992-2001 period. During the 1973-1992 period, there was only moderate increase in urban area, while there was an inverse relationship between agricultural change and forest change. In general, forest gain and agricultural loss was found in areas of improving landscape indicators, and forest loss and agricultural gain was found to occur in areas of declining indicators related to habitat and nitrogen loadings, which was generally confirmed by the aerial photographic analysis. In terms of the specific model results, bird habitat, which is mainly related to the extent of forest cover, declined overall with forest extent, but was also affected more in the decline of habitat quality. Nitrate loading, which is mainly related to agricultural land cover actually improved from 1992-2001, and in the overall study, mainly due to the conversion of agriculture to forests and urban. The high-resolution imagery analysis was significant in that it confirmed, at a very local level, the specific land-cover changes that were driving the landscape metrics and model results that were calculated from moderate resolution land-cover data and models. These were generally subtle changes in patch size of agriculture, forest, and urban areas, but had substantial effects on bird habitat and nitrogen loadings. This analysis of high-resolution imagery demonstrates and confirms the important ability of moderate-resolution land-cover data to capture significant landscape-level activity that is directly related to specific metrics of ecological significance. It also demonstrates consistent landscape-scale relationships between data derived from high-resolution, moderate-resolution and landscape-model sources. Finally, many of the areas of improvement and decline in bird habitat and nitrogen loadings appear to be potentially regional in nature and likely reflect some local trend in landscape activity. Although the use of ecoregions as sampling units has been criticized in recent years, these results show that basic changes in Level 1 land-cover categories, such as forest and agriculture, may still reflect ecoregional patterns and considerations at some scale of mapping and analysis. This is a potentially important area for future landscape-indicator research. This and other follow-on research opportunities are discussed.

Open-File Report

A comparison of hydrocarbon-related landscape disturbance patterns along the New York-Pennsylvania border, 2004–2013

Executive Summary The New York-Pennsylvania area has a long history of hydrocarbon extraction, and the addition of shale gas extraction methods contributes to landscape disturbance borne by previously developed oil and non-shale gas resources. The main unconventional extraction method used to extract shale gas from the Marcellus Shale located in New York and Pennsylvania is hydraulic fracturing, or “fracking,” although other conventional methods are used extensively. All forms of hydrocarbon extraction disturb the surrounding landscape to some extent, primarily in the form of land clearance and degradation, road construction, and pipeline development, although the effects of these disturbances are not fully understood. In this study, landscape-change metrics and indicators are used to analyze change in a 10-county region along the New York-Pennsylvania border—the New York counties of Allegany, Steuben, Chemung, Tioga, and Broome, and the Pennsylvania counties of McKean, Potter, Tioga, Bradford, and Susquehanna. This 10-county region was selected due to the differences in policies between the States of New York and Pennsylvania. While fracking occurred extensively in Pennsylvania over the past 10 years or more, the State of New York issued a temporary moratorium against hydraulic fracturing in 2010—citing repercussions that might affect air quality, water quality, and public health—and officially banned hydraulic fracturing in June 2015. The quantification of landscape disturbance due to hydrocarbon extraction activities is presented in this report as land-use and land-cover (LULC) change between 2004 and 2013 and defined using specific disturbance categories (including well sites, roads, and pipelines) to compare the disturbances and changes, by county, on both sides of the New York-Pennsylvania border. The quantification was accomplished by gathering the signatures of disturbance from high-resolution aerial images, comparing the derived totals of disturbance, and then computing landscape metrics in a geographic information system (GIS) environment. The collected data represent a summation of landscape disturbance from oil and gas development, as some of the data represented were established decades earlier. The Analytical Tools Interface for Landscape Assessments (ATtILA) software was used to calculate land-cover area and landscape metrics for each shale gas, non-shale gas, oil, and other infrastructure types associated with hydrocarbons across each county and both five-county regions in the study area. The three primary metrics used to describe changes in forest structure were (1)forest area, (2) interior forest area, and (3) forest edge area. The changes in metrics were subsequently evaluated using the Pearson correlation coefficient. Overall, the disturbed-area footprint in the Pennsylvania region is considerably larger than the disturbed-area footprint in the New York region (13,687.9 hectares [ha] in Pennsylvania; 3,840.5 ha in New York). Disturbance per site is similar, with 1.2 disturbed ha per site in New York and 1.6 disturbed ha per site in Pennsylvania. In the New York-Pennsylvania 10-county region, hydrocarbon-development and extraction disturbance strongly correlate with a reduction in the percentage of forest for the entire region. This observation also appears to be true in the New York five-county region for forest area. This form of disturbance in the New York five-county region shows significantly correlated changes in forest metrics (–0.4 percent total forest area), particularly in the percentage of interior forest (–1.2 percent total area) and forest edge (+0.7 percent total area). On the other hand, gas and hydrocarbon-development and extraction disturbance (1.0 percent total area) in the Pennsylvania five-county region strongly correlates with a total decline in forest area and agricultural land area (–0.8 percent combined total area) but not with either land-cover class separately.

New York, Pennsylvania

High spatial resolution WorldView-2 imagery for mapping NDVI and its relationship to temporal urban landscape evapotranspiration factors

Evapotranspiration estimation has benefitted from recent advances in remote sensing and GIS techniques particularly in agricultural applications rather than urban environments. This paper explores the relationship between urban vegetation evapotranspiration (ET) and vegetation indices derived from newly-developed high spatial resolution WorldView-2 imagery. The study site was Veale Gardens in Adelaide, Australia. Image processing was applied on five images captured from February 2012 to February 2013 using ERDAS Imagine. From 64 possible two band combinations of WorldView-2, the most reliable one (with the maximum median differences) was selected. Normalized Difference Vegetation Index (NDVI) values were derived for each category of landscape cover, namely trees, shrubs, turf grasses, impervious pavements, and water bodies. Urban landscape evapotranspiration rates for Veale Gardens were estimated through field monitoring using observational-based landscape coefficients. The relationships between remotely sensed NDVIs for the entire Veale Gardens and for individual NDVIs of different vegetation covers were compared with field measured urban landscape evapotranspiration rates. The water stress conditions experienced in January 2013 decreased the correlation between ET and NDVI with the highest relationship of ET-Landscape NDVI (Landscape Normalized Difference Vegetation Index) for shrubs (r 2 = 0.66) and trees (r 2 = 0.63). However, when the January data was excluded, there was a significant correlation between ET and NDVI. The highest correlation for ET-Landscape NDVI was found for the entire Veale Gardens regardless of vegetation type (r 2 = 0.95, p > 0.05) and the lowest one was for turf (r 2 = 0.88, p > 0.05). In support of the feasibility of ET estimation by WV2 over a longer period, an algorithm recently developed that estimates evapotranspiration rates based on the Enhanced Vegetation Index (EVI) from MODIS was employed. The results revealed a significant positive relationship between ETMODIS and ETWV2 (r 2 = 0.9857, p > 0.05). This indicates that the relationship between NDVI using high resolution WorldView-2 imagery and ground-based validation approaches could provide an effective predictive tool for determining ET rates from unstressed mixed urban landscape plantings.

Adelaide

Geographic context affects the landscape change and fragmentation caused by wind energy facilities

Wind energy generation causes transformation of landscapes as new roads, pads, and transmission lines are constructed. Limiting the landscape change and fragmentation caused by these facilities likely minimizes impacts to biodiversity and sensitive wildlife species. We examined the effects of wind energy facilities’ geographic context on changes in landscape patterns. We digitized the footprints of 39 wind facilities and the surrounding land cover using high-resolution imagery. We then measured landscape pattern before and after facility construction using 3 metrics associated with species responses to habitat loss and fragmentation (amount, core area, and connectivity of undeveloped land) within 1km around newly constructed turbines and roads. New facilities decreased the amount of undeveloped land, on average, by just 1.8% while changes in metrics of landscape pattern were relatively much larger (50 to 140%). Statistical models indicated levels of pre-construction development were a key factor explaining the impact of new wind facilities on landscape metrics, with pre-construction road networks, turbine spacing, and topography having smaller influences. As the proportion of developed land around facilities increased, a higher proportion of the facility utilized pre-construction developed land and a lower density of new roads were built, resulting in smaller impacts to undeveloped landscapes. Building of new road networks was also a predictor of landscape fragmentation. Density of new roads was higher in places with little pre-existing development and may be influenced by facility design. Utilizing existing development and carefully placing turbines may provide opportunities to minimize the impacts of new wind energy facilities.

PeerJ

Proximate and landscape factors influence grassland bird distributions

Ecologists increasingly recognize that birds can respond to features well beyond their normal areas of activity, but little is known about the relative importance of landscapes and proximate factors or about the scales of landscapes that influence bird distributions. We examined the influences of tree cover at both proximate and landscape scales on grassland birds, a group of birds of high conservation concern, in the Sheyenne National Grassland in North Dakota, USA. The Grassland contains a diverse array of grassland and woodland habitats. We surveyed breeding birds on 2015 100 m long transect segments during 2002 and 2003. We modeled the occurrence of 19 species in relation to habitat features (percentages of grassland, woodland, shrubland, and wetland) within each 100-m segment and to tree cover within 200-1600 m of the segment. We used information-theoretic statistical methods to compare models and variables. At the proximate scales, tree cover was the most important variable, having negative influences on 13 species and positive influences on two species. In a comparison of multiple scales, models with only proximate variables were adequate for some species, but models combining proximate with landscape information were best for 17 of 19 species. Landscape-only models were rarely competitive. Combined models at the largest scales (800-1600 m) were best for 12 of 19 species. Seven species had best models including 1600-m landscapes plus proximate factors in at least one year. These were Wilson's Phalarope (Phalaropus tricolor), Sedge Wren (Cistothorus platensis), Field Sparrow (Spizella pusilla), Grasshopper Sparrow (Ammodramus savannarum), Bobolink (Dolychonix oryzivorus), Red-winged Blackbird (Agelaius phoeniceus), and Brown-headed Cowbird (Molothrus ater). These seven are small-bodied species; thus larger-bodied species do not necessarily respond most to the largest landscapes. Our findings suggest that birds respond to habitat features at a variety of scales. Models with only landscape-scale tree cover were rarely competitive, indicating that broad-scale modeling alone, such as that based solely on remotely sensed data, is likely to be inadequate in explaining species distributions. ?? 2006 by the Ecological Society of America.

Ecological Applications

Ring-necked Pheasant brood habitat selection and movements in an intensive agricultural landscape

Management of row crops can greatly influence wildlife populations in an agriculturally intensive landscape. Many upland gamebird populations, including Phasianus colchicus L. (Ringnecked Pheasant; hereafter pheasant) are experiencing contemporary population declines in such landscapes throughout the Midwest United States. Reduced availability of quality brood habitat may be a factor in these declines. Alternative practices, such as spring cover crops, may increase brood survival and benefit local pheasant populations. Our objectives were to follow radio-tagged females and assess pheasant brood 1) movements among available habitat patches within landscapes including spring cover crops and Conservation Reserve Program (CRP) fields, 2) relative use of available cover types within a landscape, 3) selection of vegetation structure, vegetation composition, and invertebrate community structure by brood-rearing hens in landscapes dominated by row-crop agriculture, and 4) survival rates. Broods were found primarily in grassy areas (native grass, pasture, train track right-of-way, and grass strips) and spring cover crop fields even though cover crops were the least common cover type in all study areas. Brood movements were limited with broods staying near nest locations for 30 days after hatch. Though movements were small, broods were found in multiple cover types, averaging 2.8 out of 6 available cover types. There was no difference between used and random locations for invertebrate metrics including total counts, biomass, and richness; order-specific biomass; and order-specific counts. Visual obstruction and vegetation composition were similar between used and random locations. Across our study sites, we found little support for point-site selection (i.e., within-patch 4th order selection) but significant support for patch-site selection by female pheasants attending broods. Spring cover crops (<5%) and CRP (<15%) comprised a small percentage of the landscape area, but were selected by females attending broods as each contained approximately 25% of brood locations. Apparent survival of pheasant broods was low compared to other studies. Female pheasants selected for spring cover crops and CRP when attending broods, both are alternatives to current row-crop farming practices. As pheasants continue to respond to changes in western Kansas landscapes, homogeneity of cover types found in agricultural landscapes can be detrimental if practices continue to shift from quality pheasant habitat but can be advantageous if practices shift towards favorable management practices.

Kansas

Scale-specific habitat relationships influence patch occupancy: defining neighborhoods to optimize the effectiveness of landscape-scale grassland bird conservation

Context Beyond the recognized importance of protecting large areas of contiguous habitat, conservation efforts for many species are complicated by the fact that patch suitability may also be affected by characteristics of the landscape within which the patch is located. Currently, little is known about the spatial scales at which species respond to different aspects of the landscape surrounding an occupied patch. Objectives Using grassland bird point count data, we describe an approach to evaluating scale-specific effects of landscape composition on patch occupancy. Methods We used data from 793 point count surveys conducted in idle and grazed grasslands across Wisconsin, USA from 2012 to 2014 to evaluate scale-dependencies in the response of grassland birds to landscape composition. Patch occupancy models were used to evaluate the relationship between occupancy and landscape composition at scales from 100 to 3000 m. Results Bobolink ( Dolichonyx oryzivorus ) exhibited a pattern indicating selection for grassland habitats in the surrounding landscape at all spatial scales while selecting against other habitats. Eastern Meadowlark ( Sturnella magna ) displayed evidence of scale sensitivity for all habitat types. Grasshopper Sparrow ( Ammodramus savannarum ) showed a strong positive response to pasture and idle grass at all scales and negatively to cropland at large scales. Unlike other species, patch occupancy by Henslow’s Sparrow ( A. henslowii ) was primarily influenced by patch area. Conclusions Our results suggest that both working grasslands (pasture) and idle conservation grasslands can play an important role in grassland bird conservation but also highlight the importance of considering species-specific patch and landscape characteristics for effective conservation.

Landscape Ecology

Landscape complementation revealed through bipartite networks: An example with the Florida manatee

Context Landscape complementation, or how landscapes that contain two or more non-substitutable and spatially separated resources facilitate resource use, is critical for many populations. Implicit to the problem of landscape complementation is the movement of individuals to access multiple resources. Conventional measures of complementation, such as habitat area or distance between habitats, do not consider the spatial configuration of resources or how landscape features impede movement. Objectives We advanced a bipartite network approach to capture the spatial configuration and connectivity of two habitat types and contrasted this framework to conventional approaches in a habitat selection model. Methods Using satellite-telemetry of the Florida manatee (Trichechus manatus latirostris), a marine mammal that relies on two distinct, spatially separate habitats for foraging and thermoregulating, we parameterized and compared mixed conditional logistic models with covariates describing classic habitat selection metrics, conventional measures of landscape complementation, and bipartite network metrics. Results The models best supported included habitat area, resistance distance between habitats, and the bipartite network metric eigenvector centrality. The connectivity between habitats and the spatial configuration of one habitat type relative to other types better described habitat selection than conventional measures of landscape complementation alone. The type of habitat, i.e. seagrass or thermal refuge, influenced both the direction and magnitude of the response. Conclusions Landscape complementation is an important predictor of selection and thus classic complementation measures are not sufficient in describing the process. Formalization of complementation with bipartite network can therefor reveal effects potentially missed with conventional measures.

Florida

Assessing large landscape patterns of potential fire connectivity using circuit methods

Context Minimizing negative impacts of wildfire is a major societal objective in fire-prone landscapes. Models of fire connectivity can aid in understanding and managing wildfires by analyzing potential fire spread and conductance patterns. We define ‘fire connectivity’ as the landscape’s capacity to facilitate fire transmission from one point on the landscape to another. Objectives Our objective was to develop an approach for modeling fire connectivity patterns representing potential fire spread and relative flow across a broad landscape extent, particularly in the management-relevant context of fuel breaks. Methods We applied an omnidirectional circuit theory algorithm to model fire connectivity in the Great Basin of the western United States. We used predicted rates of fire spread to approximate conductance and calculated current densities to identify connections among areas with high spread rates. We compared existing and planned fuel breaks with fire connectivity patterns. Results Fire connectivity and relative flow outputs were characterized by spatial heterogeneity in the landscape’s capacity to transmit fire. We found that existing fuel break networks were denser in areas with relatively diffuse and impeded flow patterns, rather than in locations with channelized flow. Conclusions This approach could be paired with traditional fire behavior and risk analyses to better understand wildfire spread as well as direct strategic placement of individual fuel breaks within larger networks to constrain fire spread. Thus, our findings may offer local- to landscape-level support for management actions that aim to disrupt fire spread and mitigate the costs of fire on the landscape.

California, Idaho, Nevada, Oregon, Utah