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Vegetation type conversion in the US Southwest: Frontline observations and management responses

Forest and nonforest ecosystems of the western United States are experiencing major transformations in response to land-use change, climate warming, and their interactive effects with wildland fire. Some ecosystems are transitioning to persistent alternative types, hereafter called “vegetation type conversion” (VTC). VTC is one of the most pressing management issues in the southwestern US, yet current strategies to intervene and address change often use trial-and-error approaches devised after the fact. To better understand how to manage VTC, we gathered managers, scientists, and practitioners from across the southwestern US to collect their experiences with VTC challenges, management responses, and outcomes.

Arizona, California, Colorado, New Mexico

Modeled interactions of mountain pine beetle and wildland fire under future climate and management scenarios for three western US landscapes

Mountain pine beetle (MPB) is a native disturbance agent across most pine forests in the western US. Climate changes will directly and indirectly impact frequencies and severities of MPB outbreaks, which can then alter fuel characteristics and wildland fire dynamics via changes in stand structure and composition. To investigate the importance of MPB to past and future landscape dynamics, we used the mechanistic, spatially explicit ecosystem process model FireBGCv2 to quantify interactions among climate, MPB, wildfire, fire suppression, and fuel management under historical and projected future climates for three western US landscapes. We compared simulated FireBGCv2 output from three MPB modules (none, simple empirical, and complex mechanistic) using three focus variables and six exploratory variables to evaluate the importance of MPB to landscape dynamics.

Fire Ecology

Modeling of fire spread in sagebrush steppe using FARSITE: An approach to improving input data and simulation accuracy

Background: Model simulations of wildfire spread and assessments of their accuracy are needed for understanding and managing altered fire regimes in semiarid regions. The accuracy of wildfire spread simulations can be evaluated from post hoc comparisons of simulated and actual wildfire perimeters, but this requires information on pre-fire vegetation fuels that is typically not available. We assessed the accuracy of the Fire-Area Simulator (FARSITE) model parameterized with maps of fire behavior fuel models (FBFMs) obtained from the widely used LANDFIRE, as well as alternative means which utilized the classification of Rangeland Analysis Platform (RAP) satellite-derived vegetation cover maps to create FBFM maps. We focused on the 2015 Soda wildfire, which burned 113,000 ha of sagebrush steppe in the western USA, and then assessed the transferability of our RAP-to-FBFM selection process, which produced the most accurate reconstruction of the Soda wildfire, on the nearby 2016 Cherry Road wildfire. Results: Parameterizing FARSITE with maps of FBFMs from LANDFIRE resulted in low levels of agreement between simulated and observed area burned, with maximum Sorensen’s coefficient (SC) and Cohen’s kappa ( K ) values of 0.38 and 0.36, respectively. In contrast, maps of FBFMs derived from unsupervised classification of RAP vegetation cover maps led to much greater simulated-to-observed burned area agreement (SC = 0.70, K = 0.68). The FBFM map that generated the greatest simulated-to-observed burned area agreement for the Soda wildfire was then used to crosswalk FBFMs to another nearby wildfire (2016 Cherry Road), and this FBFM selection led to high FARSITE simulated-to-observed burned area agreement (SC = 0.80, K = 0.79). Conclusions: Using RAP to inform pre-fire FBFM selection increased the accuracy of FARSITE simulations compared to parameterization with the standard LANDFIRE FBFM maps, in sagebrush steppe. Additionally, the crosswalk method appeared to have regional generalizability. Flanking and backfires were the primary source of disagreements between simulated and observed fire spread in FARSITE, which are sources of error that may require modeling of lateral heterogeneity in fuels and fire processes at finer scales than used here.

Idaho, Oregon

Long-term recovery of Mexican spotted owl nesting habitat after fire in the Lincoln National Forest, New Mexico

Background Dry mixed-conifer forests of the southwestern United States are experiencing rapid, anthropogenically driven fire regime change. Prior to the Euro-American settlement, most of these forests experienced frequent surface fires but are now vulnerable to uncharacteristically large, high-severity fires. Fire directly influences the structure and composition of these forests and, in turn, the wildlife that inhabit them. Changing fire regimes result in a certain decline of some species and uncertain consequences for others. The Mexican spotted owl ( Strix occidentalis lucida ) is a federally listed threatened species of particular note in southwestern mixed-conifer forests. High-severity fire is cited as the owl’s primary threat in the revised species recovery plan, but uncertainties surround the impacts of high-severity fire on the habitat of the threatened owl, particularly across a timeframe longer than a few years. Our objective was to explore the long-term (100-year) effects of fire severity on elements of forest structure vital for Mexican spotted owl nesting. We quantified structural attributes for nest/roost habitat across mixed-conifer forests that burned at varying severity levels and time periods in the last century. We then examined the drivers of structural attributes by detecting statistical differences between severity classes and time periods through permutational multivariate analysis of variance. Results High-severity fire has the strongest deleterious impact on elements of forest structure (total basal area, percent medium tree basal area, percent large tree basal area, large tree density, and canopy cover) vital to Mexican spotted owl nesting, and although the structural differences between severity classes diminish with time, it took ≥ 80–100 years to reach the structural conditions desired for Mexican spotted owl nesting after stand-replacing fires. The most important attribute measured, canopy cover, required 90–100 years after high-severity fires to reach levels most suitable for Mexican spotted owls in the Lincoln National Forest. Conclusions As fires increase in frequency, severity, and size compared to the last century, the Lincoln National Forest is projected to face an overall decrease in the structural conditions needed for Mexican spotted owl nesting habitat in this region. Short intervals between uncharacteristically high-severity fires in particular pose an imminent threat to nesting habitat.

New Mexico

Connecting dryland fine-fuel assessments to wildfire exposure and natural resource values at risk

Background Wildland fire in arid and semi-arid (dryland) regions can intensify when climatic, biophysical, and land-use factors increase fuel load and continuity. To inform wildland fire management under these conditions, we developed high-resolution (10-m) estimates of fine fuel across the Altar Valley in southern Arizona, USA, which spans dryland, grass-dominated ecosystems that are administered by multiple land managers and owners. We coupled field measurements at the end of the 2021 growing season with Sentinel-2 satellite imagery and vegetation indices acquired during and after the growing season to develop predictions of fine fuel across the entire valley. We then assessed how climate, soil, vegetation, and land-use factors influenced the amount and distribution of fine fuels. We connected fine fuels to fire management points, past ignition history, and socio-economic vulnerability to evaluate wildfire exposure and assessed how fuel related to habitat of the endangered masked bobwhite quail ( Colinus virginianus ridgwayi ). Results The high amount of fine fuel (400–3600 kg/ha; mean = 1392 kg/ha) predicted by our remote sensing model ( R 2 = 0.63) for 2021 compared to previous years in the valley was stimulated by near-record high growing season precipitation that was 177% of the 1990–2020 mean. Fine fuel increased across the valley if it was contained within the wildlife refuge boundary and had lower temperature and vapor pressure deficit, higher soil organic content, and abundant annual plants and an invasive perennial grass ( R 2 = 0.24). The index of potential exposure to wildfire showed a clustering of high exposure centered around roads and low-density housing development distant from fire management points and extending into the upper elevations flanking the valley. Within the Buenos Aires National Wildlife Refuge, fine fuel increased with habitat suitability for the masked bobwhite quail within and adjacent to core habitat areas, representing a natural resource value at risk, accompanied with higher overall mean fine fuel (1672 kg/ha) in relation to 2015 (1347 kg/ha) and 2020 (1363 kg/ha) means. Conclusions By connecting high-resolution estimates of fine fuel to climatic, biophysical and land-use factors, wildfire exposure, and a natural resource value at risk, we provide a pro-active and adaptive framework for fire risk management within highly variable and rapidly changing dryland landscapes.

Arizona

Vegetation, fuels, and fire-behavior responses to linear fuel-break treatments in and around burned sagebrush steppe: Are we breaking the grass-fire cycle?

Background Linear fuel breaks are being implemented to moderate fire behavior and improve wildfire containment in semiarid landscapes such as the sagebrush steppe of North America, where extensive losses in perennial vegetation and ecosystem functioning are resulting from invasion by exotic annual grasses (EAGs) that foster large and recurrent wildfires. However, fuel-break construction can also pose EAG invasion risks, which must be weighed against the intended fire-moderation benefits of the treatments. We investigated how shrub reductions (mowing, cutting), pre-emergent EAG-herbicides, and/or drill seedings of fire-resistant perennial bunchgrasses (PBGs) recently applied to create a large fuel-break system affected native and exotic plant abundances and their associated fuel loading and predicted fire behavior. Results In heavily EAG-invaded areas, herbicides reduced EAG and total herbaceous cover without affecting PBGs for 2–3 years and reduced predicted fire behavior for 1 year (from the Fuel Characteristic Classification System). However, surviving post-herbicide EAG cover was still > 30%, which was sufficient fuel to exceed the conventional 1.2-m-flame length (FL) threshold for attempting wildfire suppression with hand tools. In less invaded shrubland, shrub reduction treatments largely reduced shrub cover and height by ~ half without increasing EAGs, but then redistributed the wood to ground level and increased total herbaceous cover. Herbicides and/or drill seeding after shrub reductions did not affect EAG cover, although drill seedings increased PBG cover and exotic forbs (e.g., Russian thistle). Fire behavior was predicted to be moderated in only one of the many yearly observations of the various shrub-reduction treatment combinations. Over all treatments and years, FLs were predicted to exceed 1.2 m in 13% of simulations under average (11 km h −1 ) or high (47 km h −1 ) wind speed conditions and exceed the 3.4-m threshold for uncontrollable fire in 11% of simulations under high-wind speeds only. Conclusions Predicted fire-moderation benefits over the first 4 years of fuel break implementation were modest and variable, but, generally, increases in EAGs and their associated fire risks were not observed. Nonetheless, ancillary evidence from shrublands would suggest that treatment-induced shifts from shrub to herbaceous fuel dominance are expected to improve conditions for active fire suppression in ways not readily represented in available fire models.

Idaho, Oregon

Recent large-scale prescribed fire treatments reduced Carr Fire severity at Whiskeytown National Recreation Area

Background Severe fire weather is becoming more common throughout the western United States. Changing conditions demand a better understanding of how prescribed fire treatments perform under extreme burning conditions, including the interactive influence of the age of treatments, vegetation, and fire weather. The Carr Fire of July 2018 burned nearly the entire land area of Whiskeytown National Recreation Area (NRA) under extreme fuel moisture and temperature conditions. Prior to the Carr Fire and since 1997, staff at Whiskeytown NRA treated 23% of the 15,756-ha NRA using large-scale prescribed fire (underburn) treatments ranging in size from 40 to 400 hectares. Methods We used simultaneous autoregressive (SAR) models to describe the effects of landscape-scale fuel treatments on wildfire severity under extreme burning conditions and across diverse biophysical settings at Whiskeytown NRA. Because vegetation type and structure are known drivers of fire severity in diverse ecosystems such as at Whiskeytown NRA, we also considered three different sources of vegetation structure data, including a 2006 physiognomic-floristic classification, a 2011 lidar-based forest structure classification, and a 2016 Landfire map of existing vegetation physiognomy-subclass. Results The greatest effect on 2018 Carr Fire severity was time since treatment of underburn treatments, but treatment effectiveness on fire severity dissipated rapidly—showing notable effectiveness within 5 years of underburning but virtually no effectiveness beyond 10 years post-treatment. Additional factors related to severity included vegetation structure type, topographic position index, aspect, slope, temperature, and wind gust speed. Model variance explained and model parameters, including the effect of underburn treatments, were similar regardless of the source of vegetation structure data. Conclusions Our results show that large-scale underburning treatments can reduce wildfire severity even under extreme fire weather conditions but suggest that frequent maintenance intervals are required to maintain treatment effectiveness ahead of severe wildfire events.

California

Representing 3-dimensional fuels for physics-based fire behavior models: A general framework and case study in a type-converted post-fire shrubfield

Background Physics-based three-dimensional (3D) fire behavior models improve planning for prescribed fire application and wildfire mitigation, but require high spatial resolution 3D fuel models as inputs. While multiple methods and data sources for realistically representing 3D, heterogeneous fuels are available, no unifying framework exists to guide the use of these tools to create 3D fuel models across gradients of vegetation characteristics and data availability. Existing data and methods are most uncertain for mid-level fuels (e.g., shrubs and small trees), due to canopy obstruction of remotely sensed data and a relative lack of modeling efforts. Yet, mid-level fuels are especially important as potential ladder fuels and increasingly common as the dominant fuel in type-converted, post-fire, shrub-dominated landscapes. Results Here we introduce the Framework for Representing 3D Fuels (FR3D), a general framework for combining multiple data sources and methods to construct 3D fuel models for forested and unforested landscapes. We then demonstrate FR3D in a case study to build a 3D fuelbed model in a post-fire, shrub-dominated landscape using three new methods for deriving mid-level shrub fuels from: (1) Airborne Laser Scanning (ALS), (2) imputation of Terrestrial Laser Scanning (TLS), and (3) generative modeling of TLS. We compare the resulting fuel models and examine how they affected simulated 3D fire behavior using QUIC-Fire. While each method represented the broad landscape patterning of shrubs, differences in shrub loading, height, and cover highlighted advantages and drawbacks of the different methods. Modeled fire behavior was realistic for all fuel representation methods, but rate of spread and fine fuel consumption was sensitive to the different arrangements of shrubs. Conclusions The sensitivity of fire behavior to shrub modeling methods emphasizes the need for fuel models that faithfully represent local fuelbed characteristics and conditions, and highlights the value in testing a range of modeled fuels to understand the potential range of prescribed fire outcomes. FR3D and novel methods of modeling mid-level fuel provide a foundation for tool integration efforts and increased site-specificity of fuel representation for physics-based fire models.

New Mexico

Near real-time indicators of burn severity in the western U.S. from active fire tracking

Background Timely information on wildfire burn severity is critical to assess and mitigate potential post-fire impacts on soils, vegetation, and hillslope stability. Tracking individual fire spread and intensity using satellite active fire data provides a pathway to near real-time (NRT) information. Here, we generated a large database ( n = 2177) of wildfire events in the western United States (U.S.) between 2012 and 2021 using active fire detections from the Visible Infrared Imaging Radiometer Suite (VIIRS) sensor on the Suomi National Polar-orbiting Partnership (SNPP) satellite and the Fire Events Data Suite (FEDS) algorithm to track large fire growth every 12 h. We integrated fire tracking data with final fire perimeters and burn severity data from the Monitoring Trends in Burn Severity (MTBS) program to evaluate the relationship between burn severity and fire behavior metrics derived from the fire tracking approach, including the rate of fire spread and average fire radiative power (FRP) of fire detections for each 12-h growth increment. Results When stratified by vegetation type, FRP and rate of spread metrics were positively correlated with classified burn severity for each 12-h growth increment, highlighting the potential to rapidly identify areas of high and low severity burning. In forests, integrated measures of FRP over the fire lifetime captured persistent flaming and smoldering that compensated for initial differences between AM (01:30) and PM (13:30) fire detections. Predictive modeling of these relationships based on multiple fire behavior indicators and vegetation type from the LANDFIRE program yielded an accuracy of 78% for the separation of unburned/low and moderate/high burn severity classes. Conclusions These results demonstrate the ability to capture within-fire differences in burn severity using NRT indicators from fire tracking to assist with emergency management and disaster preparedness for post-fire hazards, such as landslides, debris flows, or changes in stream flow and water quality. As VIIRS data are available within minutes of each satellite overpass in the U.S., rapid estimates of burn severity based on fire tracking can be made days or weeks before a large wildfire is fully contained.

western United States

The state of the giant sequoias: Losses, risks, and opportunities

Background Giant sequoias, an iconic tree found only in a narrow band along the western slope of the Sierra Nevada of California, are facing unprecedented threats. Despite being adapted to fire and resistant to both drought and insect attack, the species has exhibited signs of vulnerability to these stressors in recent years, with the most substantial impacts coming from wildfire. Recently, several unprecedentedly large and severe fires have resulted in the deaths of many large trees and, in some cases, limited postfire regeneration. These impacts have led to heightened efforts by managers to implement treatments to protect those places that are still vulnerable. Here, we provide a comprehensive, range-wide assessment of the condition of the species, including an evaluation of the trends in wildfire and its effects on large giant sequoia mortality and the potential for local extirpation, treatment patterns, and vulnerability to future fire. Results From 2015 to 2024, 82% of giant sequoia grove area burned in wildfire, compared to 24% between 1910 and 2014. We conservatively estimate that 17.6% of all large giant sequoias have been killed by wildfire since 1984, with most dying in 2020 and 2021. Due to very severe wildfire, roughly 13% of the range is at some risk of local extirpation due to the loss of mature trees and limited predicted regeneration. Treatment activity has increased since 1995, with treatment type and amount varying by jurisdiction. The combination of treatment and wildfire has created a mosaic of resistance to severe fire, with 26% of grove area at high resistance, 38% at moderate resistance, 2% at low resistance, and 16% having no resistance. The remaining 18% are areas where mature giant sequoia forest was lost due to high severity fire. Conclusions Giant sequoias have experienced substantial losses due to recent wildfire, with large areas at risk of grove loss. However, most burned area had lower severity effects, creating opportunities to leverage wildfire as a treatment. When combined with areas of active management, these opportunities can promote giant sequoia conservation in a time of rapid change.

California

Relationship of satellite-derived fire severity to archaeological fire severity and fire effects in the Jemez Mountains, New Mexico, USA

Background Archaeological attributes important for determining cultural affiliation, site age, or artifact origin can be damaged or destroyed by wildfires. Satellite-derived fire severity products are used in post-fire rapid response to prioritize and guide stabilization and preservation activities, but their ability to predict archaeological fire impacts has not yet been tested. We compared the severity of fire from Burned Area Reflectance Classification (BARC) maps with ground-based observations of fire severity (“archaeological fire severity”) and archaeological impacts for 904 prehistoric or historic sites within three wildfires in the Jemez Mountains of north-central New Mexico, U.S. Results from this unique study inform archaeological preservation in the context of wildfires and fire management. Results We found a statistically significant, moderately strong association between BARC and archaeological fire severity and a high likelihood that sites with a low BARC fire severity also had a low archaeological fire severity designation. Sites of moderate or high BARC severity were less likely to have a matching severity classification in post-fire assessments, and BARC maps tended to identify as unburned areas where archaeologists determined that sites were burned at low severity. Wildfire of any severity resulted in measurable impacts to archaeological architecture, features, or artifacts at some sites, with increasing proportion of sites impacted at moderate versus low BARC severity, but fire impacts were also observed at sites classified by BARC as unburned. Conclusions Generally, BARC maps provide good assessment of the severity of fire and its effects on archaeological resources, particularly for areas burned at low severity. In areas of moderate or high fire severity, BARC maps may not consistently identify sites with a high need for assessment, and archaeological fire effects can also be present at sites classified by BARC as unburned. Adoption of standardized post-fire archaeological assessments across wildfires and regions, exploration of alternative fire mapping products, and continued research on the fire environments that produce archaeological fire effects can enhance archaeological preservation and post-fire response.

New Mexico

Effects of repeat prescribed burning in dry coniferous forests in national parks of California

Background Prescribed fire is a common approach to reduce fuels and mitigate fire hazards. The accumulation of live and dead fuels following initial treatment means that repeated application of prescribed fire could be used to maintain this benefit. However, the effect of repeated prescribed fires is not well documented in many dry coniferous forests in the western United States. Here, we present observations of changes in live trees and surface fuels following two prescribed fires in dry coniferous forests in national parks of California. Results Changes in forest structure and accumulation of surface fuels were similar over time following initial-entry and second-entry fires. An exception was that repeated fires were associated with substantial reductions in stem density. There were smaller changes in live tree basal area and stem biomass. Conclusions Our results indicate that following initial-entry fires, subsequent burning maintained reductions in surface fuel loads without major inadvertent losses of live tree basal area and stem biomass, implying the survival of large trees.

California

Size of modern megafire not unprecedented and a cumulative fire deficit persists

Background The area burned by wildfires in western North American forests has grown since the 1980s, largely driven by individual “megafires.” Despite breaking state records for fire size, it remains uncertain whether these large fires are unprecedented, owing to limited knowledge of the area burned during historical fire regimes (pre-1900). We used new methods and 14 tree-ring fire-scar sites to evaluate whether the area burned by Arizona’s largest fire on record – the 217,741 ha 2011 Wallow Fire – was unprecedented in a multi-century context. Using the 375-year reconstruction of annual area burned, we also assessed the departure of the modern fire regime by introducing cumulative fire deficit, calculated as the cumulative sum of 20-year area burned anomalies. Results The size of the Wallow Fire was exceptional in the modern era, but comparable to the extent of burning in 1748, 1847, and 1851 on the same landscape. Notably, years of extensive fire (50,000–100,000 ha) were common on this landscape prior to 1900. The average return interval for fires exceeding 100,000 ha was just 20 years. By contrast, the 2011 Wallow Fire was the only modern fire > 10,000 ha in our study area ( n = 129, 1970–2024), leading to an 85% reduction in average annual area burned. This exceptional departure from the historical fire regime generated a cumulative fire deficit exceeding 2.0 million hectares by 2011, which continues to increase despite the occurrence of the large Wallow Fire. Conclusions Although the modern wildfire crisis is characterized by large, severe wildfires, historical fire regimes also included extensive fires. On an Arizona landscape, the sizes of several historical fires were on par with the largest wildfire on record. Similar historical precedent of large fires is likely true elsewhere in dry conifer forests of western North America. These findings underscore that future wildfire events could be even larger, and likely more severe, as fuels accumulate and the climate grows warmer and drier. Forest restoration aimed at reestablishing characteristic disturbance regimes, including frequent, large, low-severity fires, could be used to assist recovery of these ecosystems resulting from the cumulative fire deficit.

Arizona

Recovery trajectories of surface fuels and forest trees following prescribed fire in low elevation conifer forests of California and southern Oregon

Background Prescribed fire is commonly used to manage surface fuels and stand structure in dry coniferous forests in the western United States. While the effectiveness of prescribed fire to manage fuel loads and live forest structure has been well documented, less is known about the shape of recovery trajectories more than a decade following treatment. We used up to 30 years of longitudinal observations from > 180 monitoring plots treated with a single prescribed fire-only treatment across six national parks in California and Oregon to compare competing recovery trajectory models: generalized linear models (GLM) that presumed exponential trajectories over time, generalized additive models (GAM) that had maximum flexibility to fit responses, and generalized Michaelis–Menten models that describe an asymptotic recovery trajectory. Results The Michaelis–Menten model had the lowest median RMSE of the fuel recovery models, with predicted recovery times and maximal fuel loads tending to increase with increasing fuel particle size. However, predictive performance was similar between model forms. GAM best predicted forest structure, showing that stem density of live overstory trees (> 15 cm stem diameter) declined immediately following prescribed fire without appreciable postfire accumulation. Results for live stem biomass were similar, but with less pronounced reductions. Conclusions We found that reductions in surface fuel biomass from a single prescribed fire-only treatment can last for more than a decade for some fuel types. Our analysis also demonstrates that asymptotic models reasonably describe surface fuel recovery trajectories. However, we lack the evidence to definitively conclude that asymptotic models best describe postfire recovery, even with repeated observations of up to three decades postfire. We also found that stem density and biomass of live trees with stem diameter > 15 cm both declined in the first few years immediately following the fire, after which they remained relatively unchanged for the remainder of the observation period.

California, Oregon

Responses in bird communities to wildland fires in southern California

There is a growing body of literature covering the responses of bird species to wildland fire events. Our study was unique among these because we investigated the effects of large-scale wildland fires on entire bird communities across multiple vegetation types. We conducted avian point counts during the breeding seasons for two years before and two years after the Cedar and Otay Fires in 2003 in southern California. Our balanced sampling effort took place at two sites, one low-elevation and one high-elevation, each containing replicate stations (burned and unburned) within five vegetation types: chaparral, coastal sage scrub, grassland, oak woodland, and riparian. Although fire caused some degree of change in the vegetation structure at all of our impacted survey points, we found that the post-fire shrub and tree cover was significantly lower in only two of the vegetation types within the low-elevation site, coastal sage scrub and chaparral. We found no significant changes in cover at the high-elevation site. Using univariate and multivariate analyses, we tested whether the fires were associated with a change in bird species diversity, community structure, and the relative abundance of individuals within a species. We found that species diversity changed in only one circumstance: it increased in coastal sage scrub at the low-elevation site. Multivariate analyses revealed significant differences in the post-fire bird community structure in the low-elevation chaparral, low-elevation coastal sage scrub, and the high-elevation grassland communities. Vegetation characteristics altered by fire, such as decreases in shrub and tree cover, influenced the changes we observed in the bird communities. The relative abundance of some species (lazuli bunting [ Passerina amoena ] and horned lark [ Eremophila alpestris ]) significantly increased after the fires, while other species declined significantly (Anna's hummingbird [ Calypte anna ], wrentit [ Chamaea fasciata ], and bushtit [ Psaltriparus minimus ]). We detected mixed results for the spotted towhee ( Pipilo maculatus ), which increased in burned chaparral and declined in burned coastal sage scrub within the low-elevation site. We suggest that the observed responses of birds to these fires may be attributed to: (1) the availability of nearby unburned refugia, (2) the continued suitability of post-fire vegetation at the study sites, and (3) the generally high mobility of this taxon.

California

Negligible influence of spatial autocorrelation in the assessment of fire effects in a mixed conifer forest

Fire is an important feature of many forest ecosystems, although the quantification of its effects is compromised by the large scale at which fire occurs and its inherent unpredictability. A recurring problem is the use of subsamples collected within individual burns, potentially resulting in spatially autocorrelated data. Using subsamples from six different fires (and three unburned control areas) we show little evidence for strong spatial autocorrelation either before or after burning for eight measures of forest conditions (both fuels and vegetation). Additionally, including a term for spatially autocorrelated errors provided little improvement for simple linear models contrasting the effects of early versus late season burning. While the effects of spatial autocorrelation should always be examined, it may not always greatly influence assessments of fire effects. If high patch scale variability is common in Sierra Nevada mixed conifer forests, even following more than a century of fire exclusion, treatments designed to encourage further heterogeneity in forest conditions prior to the reintroduction of fire will likely be unnecessary.

Fire Ecology

The effects of raking on sugar pine mortality following prescribed fire in Sequoia and Kings Canyon National Parks, California, USA

Prescribed fire is an important tool for fuel reduction, the control of competing vegetation, and forest restoration. The accumulated fuels associated with historical fire exclusion can cause undesirably high tree mortality rates following prescribed fires and wildfires. This is especially true for sugar pine ( Pinus lambertiana Douglas), which is already negatively affected by the introduced pathogen white pine blister rust ( Cronartium ribicola J.C. Fisch. ex Rabenh). We tested the efficacy of raking away fuels around the base of sugar pine to reduce mortality following prescribed fire in Sequoia and Kings Canyon national parks, California, USA. This study was conducted in three prescribed fires and included 457 trees, half of which had the fuels around their bases raked away to mineral soil to 0.5 m away from the stem. Fire effects were assessed and tree mortality was recorded for three years after prescribed fires. Overall, raking had no detectable effect on mortality: raked trees averaged 30% mortality compared to 36% for unraked trees. There was a significant effect, however, between the interaction of raking and average pre-treatment forest floor fuel depth: the predicted probability of survival of a 50 cm dbh tree was 0.94 vs. 0.96 when average pre-treatment fuel depth was 0 cm for a raked and unraked tree, respectively. When average pre-treatment forest floor fuel depth was 30 cm, the predicted probability of survival for a raked 50 cm dbh tree was 0.60 compared to only 0.07 for an unraked tree. Raking did not affect mortality when fire intensity, measured as percent crown volume scorched, was very low (0% scorch) or very high (>80% scorch), but the raking treatment significantly increased the proportion of trees that survived by 9.6% for trees that burned under moderate fire intensity (1% to 80% scorch). Raking significantly reduced the likelihood of bole charring and bark beetle activity three years post fire. Fuel depth and anticipated fire intensity need to be accounted for to maximize the effectiveness of the treatments. Raking is an important management option to reduce tree mortality from prescribed fire, but is most effective under specific fuel and burning conditions.

California