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USGS2016

Spatial genetic structure of Long-tailed Ducks (Clangula hyemalis) among Alaskan, Canadian, and Russian breeding populations

Arctic ecosystems are changing at an unprecedented rate. How Arctic species are able to respond to such environmental change is partially dependent on the connections between local and broadly distributed populations. For species like the Long-tailed Duck ( Clangula hyemalis ), we have limited telemetry and band-recovery information from which to infer population structure and migratory connectivity; however, genetic analyses can offer additional insights. To examine population structure in the Long-tailed Duck, we characterized variation at mtDNA control region and microsatellite loci among four breeding areas in Alaska, Canada, and Russia. We observed significant differences in the variance of mtDNA haplotype frequencies between the Yukon-Kuskokwim Delta (YKD) and the three Arctic locations (Arctic Coastal Plain in Alaska, eastern Siberia, and central Canadian Arctic). However, like most sea duck genetic assessments, our study found no evidence of population structure based on autosomal microsatellite loci. Long-tailed Ducks use multiple wintering areas where pair formation occurs with some populations using both the Pacific and Atlantic Oceans. This situation provides a greater opportunity for admixture across breeding locales, which would likely homogenize the nuclear genome even in the presence of female philopatry. The observed mtDNA differentiation was largely due to the presence of two divergent clades: (A) a clade showing signs of admixture among all breeding locales and (B) a clade primarily composed of YKD samples. We hypothesize that the pattern of mtDNA differentiation reflects some degree of philopatry to the YKD and isolation of two refugial populations with subsequent expansion and admixture. We recommend additional genetic assessments throughout the circumpolar range of Long-tailed Ducks to further quantify aspects of genetic diversity and migratory connectivity in this species.

Arctic Coastal Plain, central Canadian Arctic, eas↗

USGS2009

Migratory patterns and population structure among breeding and wintering red-breasted mergansers ( Mergus serrator ) and common mergansers ( M. merganser )

Philopatry has long been assumed to structure populations of waterfowl and other species of birds genetically, especially via maternally transmitted mitochondrial DNA (mtDNA), yet other migratory behaviors and nesting ecology (use of ground vs. cavity sites) may also contribute to population genetic structure. We investigated the effects of migration and nesting ecology on the population genetic structure of two Holarctic waterfowl, the Red-breasted Merganser ( Mergus serrator ) and Common Merganser ( M. merganser ), using mtDNA control-region sequence data. Red-breasted Mergansers (a ground-nesting species) exhibited lower levels of population differentiation across their North American range, possibly as a result of post-Pleistocene range expansion and population growth. By contrast, Common Mergansers (a cavity-nesting species) breeding in western and eastern North America were strongly differentiated, as were continental groups in North America and Europe. Our hypothesis that population differentiation of breeding female Common Mergansers results from limited migration during non-breeding periods was not supported, in that equally heterogeneous mtDNA lineages were observed in males and females on several wintering areas. The interspecific differences in mtDNA patterns for these two closely related species may have resulted from factors related to nesting ecology (ground vs. cavity nesting) and responses to historical climate change.

The Auk↗

USGS2012

Evidence for population bottlenecks and subtle genetic structure in the yellow rail

The Yellow Rail (Coturnicops noveboracencis) is among the most enigmatic and least studied North American birds. Nesting exclusively in marshes and wetlands, it breeds largely east of the Rocky Mountains in the northern United States and Canada, but there is an isolated population in southern Oregon once believed extirpated. The degree of connectivity of the Oregon population with the main population is unknown. We used mitochondrial DNA sequences (mtDNA) and six microsatellite loci to characterize the Yellow Rail's genetic structure and diversity patterns in six areas. Our mtDNA-based analyses of genetic structure identified significant population differentiation, but pairwise comparison of regions identified no clear geographic trends. In contrast, microsatellites suggested subtle genetic structure differentiating the Oregon population from those in the five regions sampled in the Yellow Rail's main breeding range. The genetic diversity of the Oregon population was also the lowest of the six regions sampled, and Oregon was one of three regions that demonstrated evidence of recent population bottlenecks. Factors that produced population reductions may include loss of wetlands to development and agricultural conversion, drought, and wildfire. At this time, we are unable to determine if the high percentage (50%) of populations having experienced bottlenecks is representative of the Yellow Rail's entire range. Further genetic data from additional breeding populations will be required for this issue to be addressed.

The Condor↗

USGS1999

Structure and stability of the midsummer fish communities in Chequamegon Bay, Lake Superior, 1973-1996

We analyzed the structure and stability of the summer fish communities of Chequamegon Bay, Lake Superior, during 1973–1996 from data collected with bottom trawls at 39 stations. Fifty‐three taxa were collected during the study, but we found that relative abundances for 20 taxa described most of the internal variability of the data for all taxa. Abundance data for the 20 species showed that two communities existed in the bay; one inhabited shallow water (≤3.0 m) whereas the other inhabited deeper water (>3.0 m). No temporal patterns of change were found in the structure of the shallow‐water community, whose variation was best described by abundances of 12 taxa. The deepwater community, whose variation was best described by eight taxa, underwent three periods ofstability; 1973–1978, 1979–1988, and 1989–1996. We conclude that the shallow‐water community was stable throughout the 24 years studied. Dynamics of the deepwater community were greatly affected by changes in stocking rates of lake trout Salvelinus namaycush and splake (hybrid of brook trout S. fontinalis × lake trout) and by rehabilitation of populations of lake herring Coregonus artedi and lake whitefish C. clupeaformis. Information on the existence, structure, stability, and habitats of fish communities in the bay will be useful for assessing changes in those communities that result from further changes in the bay or lake ecosystems.

Wisconsin↗

USGS2002

Population genetic structure in Lahontan cutthroat trout

We used 10 microsatellite loci to examine the genetic population structure of cutthroat trout Oncorhynchus clarki within the Lahontan Basin complex. Genetic diversity was analyzed for trout from Nevada, California, and Utah representing three putative subspecies: Lahontan O. c. henshawi , Paiute O. c. seleniris , and Humboldt (an unnamed subspecies) cutthroat trout. We found significant differences in microsatellite diversity among the three putative subspecies found in this area. Analysis of molecular variance partitioned microsatellite variation as 9.8% among subspecies, 27.7% among populations, and 62.5% within populations of Lahontan Basin cutthroat trout. Genetic distance analyses (Cavalli-Sforza-Edwards and F st ) supported unique population structure in cutthroat trout from the Humboldt and Pilot Peak drainages. Pairwise F st values for Lahontan cutthroat trout were not significantly correlated with geographic distance between population pairs ( r 2 = 0.008; P < 0.0001), suggesting that they are extremely isolated populations with small effective sizes that are vulnerable to extinction. Two extant hatchery strains of Lahontan cutthroat trout showed genetic associations with different geographic source populations. The Pyramid Lake hatchery strain was most closely associated genetically with fish from Summit Lake. The Pilot Peak hatchery strain was associated genetically with Pilot Peak wild trout (Utah) and Macklin Creek trout (California). The phylogeographic diversity depicted in this study supports unique population structure and suggests important evolutionary relationships needed to evaluate transplanted populations and hatchery supplementation within the basin.

California, Idaho, Nevada, Oregon↗

USGS2004

Age structure and mortality of walleyes in Kansas reservoirs: Use of mortality caps to establish realistic management objectives

Age structure, total annual mortality, and mortality caps (maximum mortality thresholds established by managers) were investigated for walleye Sander vitreus (formerly Stizostedion vitreum) populations sampled from eight Kansas reservoirs during 1991-1999. We assessed age structure by examining the relative frequency of different ages in the population; total annual mortality of age-2 and older walleyes was estimated by use of a weighted catch curve. To evaluate the utility of mortality caps, we modeled threshold values of mortality by varying growth rates and management objectives. Estimated mortality thresholds were then compared with observed growth and mortality rates. The maximum age of walleyes varied from 5 to 11 years across reservoirs. Age structure was dominated (???72%) by walleyes age 3 and younger in all reservoirs, corresponding to ages that were not yet vulnerable to harvest. Total annual mortality rates varied from 40.7% to 59.5% across reservoirs and averaged 51.1% overall (SE = 2.3). Analysis of mortality caps indicated that a management objective of 500 mm for the mean length of walleyes harvested by anglers was realistic for all reservoirs with a 457-mm minimum length limit but not for those with a 381-mm minimum length limit. For a 500-mm mean length objective to be realized for reservoirs with a 381-mm length limit, managers must either reduce mortality rates (e.g., through restrictive harvest regulations) or increase growth of walleyes. When the assumed objective was to maintain the mean length of harvested walleyes at current levels, the observed annual mortality rates were below the mortality cap for all reservoirs except one. Mortality caps also provided insight on management objectives expressed in terms of proportional stock density (PSD). Results indicated that a PSD objective of 20-40 was realistic for most reservoirs. This study provides important walleye mortality information that can be used for monitoring or for inclusion into population models; these results can also be combined with those of other studies to investigate large-scale differences in walleye mortality. Our analysis illustrates the utility of mortality caps for monitoring walleye populations and for establishing realistic management goals.

North American Journal of Fisheries Management↗

USGS2008

Lake Erie Yellow perch age estimation based on three structures: Precision, processing times, and management implications

Yellow perch Perca flavescens support economically important recreational and commercial fisheries in Lake Erie and are intensively managed. Age estimation represents an integral component in the management of Lake Erie yellow perch stocks, as age-structured population models are used to set safe harvest levels on an annual basis. We compared the precision associated with yellow perch (N = 251) age estimates from scales, sagittal otoliths, and anal spine sections and evaluated the time required to process and estimate age from each structure. Three readers of varying experience estimated ages. The precision (mean coefficient of variation) of estimates among readers was 1% for sagittal otoliths, 5-6% for anal spines, and 11-13% for scales. Agreement rates among readers were 94-95% for otoliths, 71-76% for anal spines, and 45-50% for scales. Systematic age estimation differences were evident among scale and anal spine readers; less-experienced readers tended to underestimate ages of yellow perch older than age 4 relative to estimates made by an experienced reader. Mean scale age tended to underestimate ages of age-6 and older fish relative to otolith ages estimated by an experienced reader. Total annual mortality estimates based on scale ages were 20% higher than those based on otolith ages; mortality estimates based on anal spine ages were 4% higher than those based on otolith ages. Otoliths required more removal and preparation time than scales and anal spines, but age estimation time was substantially lower for otoliths than for the other two structures. We suggest the use of otoliths or anal spines for age estimation in yellow perch (regardless of length) from Lake Erie and other systems where precise age estimates are necessary, because age estimation errors resulting from the use of scales could generate incorrect management decisions. ?? Copyright by the American Fisheries Society 2008.

North American Journal of Fisheries Management↗

USGS2008

Fish assemblages in a western Iowa stream modified by grade control structures

Over 400 riprap grade control structures (GCSs) have been built in streams of western Iowa to reduce erosion and protect bridges, roads, and farmland. In conjunction with a companion study evaluating fish passage over GCSs in Turkey Creek, we evaluated the differences in fish assemblage and habitat characteristics in reaches immediately downstream from GCSs (GCS sites) and reaches at least 1 km from any GCS (non-GCS sites). The GCS sites were characterized by greater proportions of pool habitat, maximum depths, fish biomass, and abundance of juvenile largemouth bass Micropterus salmoides than were non-GCS sites. Index of biotic integrity (IBI) scores were poor or fair (<43 on a 0-100 scale) and not significantly different between the GCS and non-GCS sites. Additionally, we investigated both the longitudinal changes in fish assemblages in this GCS-fragmented stream and the changes in fish assemblages after slope modifications of three GCSs to facilitate fish passage. Thirteen fish species were present throughout the study area, whereas another 15 species exhibited truncated distributions not extending to the most upstream sampling location. After modification of the GCSs, IBI scores increased at seven of nine sites (mean increase =4.6 points). Also, channel catfish Ictalurus punctatus were detected 7.3 km upstream at sites where, 2 years before GCS modification, they had been absent from collections. Given the number and distribution of GCSs in western Iowa streams, understanding the effects of these structures is vital to the conservation and management of fish assemblages in this and other regions where GCSs or similar structures are used. ?? Copyright by the American Fisheries Society 2008.

North American Journal of Fisheries Management↗

USGS2010

Influence of drought conditions on brown trout biomass and size structure in the Black Hills, South Dakota

We evaluated the influence of drought conditions on the biomass of brown trout Salmo trutta in Spearfish Creek, upper Rapid Creek, and lower Rapid Creek in the Black Hills of western South Dakota. Stream discharge, mean summer water temperature, the biomass of juvenile and adult brown trout, and brown trout size structure were compared between two time periods: early (2000–2002) and late drought (2005–2007). Mean summer water temperatures were similar between the early- and late-drought periods in Spearfish Creek (12.4°C versus 11.5°C), lower Rapid Creek (19.2°C versus 19.3°C), and upper Rapid Creek (9.8°C in both periods). In contrast, mean annual discharge differed significantly between the two time periods in Spearfish Creek (1.95 versus 1.50 m 3 /s), lower Rapid Creek (2.01 versus 0.94 m 3 /s), and upper Rapid Creek (1.41 versus 0.84 m 3 /s). The mean biomass of adult brown trout in all three stream sections was significantly higher in the early-drought than in the late-drought period (238 versus 69 kg/ha in Spearfish Creek, 272 versus 91 kg/ha in lower Rapid Creek, and 159 versus 32 kg/ha in upper Rapid Creek). The biomass of juvenile brown trout was similar (43 versus 23 kg/ha) in Spearfish Creek in the two periods, declined from 136 to 45 kg/ha in lower Rapid Creek, and increased from 14 to 73 kg/ha in upper Rapid Creek. Size structure did not differ between the early- and late-drought periods in lower Rapid and Spearfish creeks, but it did in upper Rapid Creek. In addition to drought conditions, factors such as angler harvest, fish movements, and the nuisance algal species Didymosphenia geminata are discussed as possible contributors to the observed changes in brown trout biomass and size structure in Black Hills streams.

South Dakota↗

USGS2005

Spatiotemporal patterns of fish assemblage structure in a river impounded by low-head dams

We studied spatiotemporal patterns of fish assemblage structure in the Neosho River, Kansas, a system impounded by low-head dams. Spatial variation in the fish assemblage was related to the location of dams that created alternating lotic and lentic stream reaches with differing fish assemblages. At upstream sites close to dams, assemblages were characterized by species associated with deeper, slower-flowing habitat. Assemblages at sites immediately downstream from dams had higher abundance of species common to shallow, swift-flowing habitat. Temporal variation in assemblage structure was stronger than spatial variation, and was associated with fish life history events such as spawning and recruitment, as well as seasonal changes in environmental conditions. Our results suggest that low-head dams can influence spatial patterns of fish assemblage structure in systems such as the Neosho River and that such assemblages also vary seasonally.

Copeia↗

USGS2004

Genetic structure of cougar populations across the Wyoming basin: Metapopulation or megapopulation

We examined the genetic structure of 5 Wyoming cougar (Puma concolor) populations surrounding the Wyoming Basin, as well as a population from southwestern Colorado. When using 9 microsatellite DNA loci, observed heterozygosity was similar among populations (HO = 0.49-0.59) and intermediate to that of other large carnivores. Estimates of genetic structure (FST = 0.028, RST = 0.029) and number of migrants per generation (Nm) suggested high gene flow. Nm was lowest between distant populations and highest among adjacent populations. Examination of these data, plus Mantel test results of genetic versus geographic distance (P ??? 0.01), suggested both isolation by distance and an effect of habitat matrix. Bayesian assignment to population based on individual genotypes showed that cougars in this region were best described as a single panmictic population. Total effective population size for cougars in this region ranged from 1,797 to 4,532 depending on mutation model and analytical method used. Based on measures of gene flow, extinction risk in the near future appears low. We found no support for the existence of metapopulation structure among cougars in this region.

Journal of Mammalogy↗

USGS2018

Interisland genetic structure of two endangered Hawaiian waterbirds: The Hawaiian Coot and Hawaiian Gallinule

Most of Hawaii's endemic avifauna are species of conservation concern. Some of Hawaii's endangered waterbirds, however, have increased in number as a result of intensive management of wetlands. To inform these conservation efforts, we examined interisland genetic structure and gene flow within 2 Hawaiian endemic waterbirds, the Hawaiian Coot ( Fulica alai ) and the Hawaiian subspecies of the Common Gallinule ( Gallinula galeata sandvicensis ), using microsatellite and mitochondrial loci. Hawaiian Coots and Hawaiian Gallinules occupy coastal wetlands and exhibit similar life history characteristics and generation times, although they may differ in dispersal propensity. Mark–resight data for Hawaiian Coot indicate interisland movements, whereas Hawaiian Gallinules are sedentary. Genetic diversity is partitioned across the landscape differently for Hawaiian Coots and Hawaiian Gallinules; patterns of variation are likely influenced by behavioral and ecological mechanisms. Hawaiian Coots exhibit low levels of structure at microsatellite loci ( F ST = 0.029) and high levels of gene flow among islands. Conversely, Hawaiian Gallinules are highly structured across marker types (microsatellite F ST = 0.205, mtDNA control region F ST = 0.370, mtDNA ND2 F ST = 0.087), with restricted recent gene flow. Patterns of gene flow have changed after the population declines in the early to mid-1900s. Gene flow estimates indicate historical dispersal from Kauai to Oahu in both species, while recent estimates show individual Hawaiian Coots dispersing from Oahu and restricted gene flow between islands for the Hawaiian Gallinule. Changes in gene flow through time suggest that patterns of dispersal may be an artifact of the availability of habitat, which may be indirectly associated with the synergistic influences of population density and wetland quality. Despite recent population size increases for both species, continued threats to Hawaiian waterbirds (i.e. nonnative mammalian predators and invasive plants, avian disease, altered hydrology, and saltwater inundation of freshwater wetlands) will likely require continued active management to maintain viable populations.

Hawaii↗

USGS2003

Comparison of wetland structural characteristics between created and natural salt marshes in southwest Louisiana, USA

The use of dredge material is a well-known technique for creating or restoring salt marshes that is expected to become more common along the Gulf of Mexico coast in the future. However, the effectiveness of this restoration method is still questioned. Wetland structural characteristics were compared between four created and three natural salt marshes in southwest Louisiana, USA. The created marshes, formed by the pumping of dredge material into formerly open water areas, represent a chronosequence, ranging in age from 3 to 19 years. Vegetation and soil structural factors were compared to determine whether the created marshes become more similar over time to the natural salt marshes. Vegetation surveys were conducted in 1997, 2000, and 2002 using the line-intercept technique. Site elevations were measured in 2000. Organic matter (OM) was measured in 1996 and 2002, while bulk density and soil particle-size distribution were determined in 2002 only. The natural marshes were dominated by Spartina alterniflora , as were the oldest created marshes; these marshes had the lowest mean site elevations (<30 cm NGVD). The six-year-old created marsh (formed in 1996) was higher in elevation (> 35 cm NGVD) and became dominated by high marsh ( S. patens, Distichlis spicata ) and shrub ( Baccharis halimifolia, Iva frutescens ) species. The higher elevation marsh seems to be following a different plant successional trajectory than the other marshes, indicating a relationship between marsh elevation and species composition. The soils in both the created and natural marshes contain high levels of clays (30–65 %), with sand comprising < 1 % of the soil distribution. OM was significantly greater and bulk density significantly lower in two of the natural marshes when compared to the created marshes. The oldest created marsh had significantly greater OM than the younger created marshes, but it may still take several decades before equivalency is reached with the natural marshes. Vegetation structural characteristics in the created marshes take only a few years to become similar to those in the natural marshes, just so long as the marshes are formed at a proper elevation. This agrees with other studies from North Carolina and Texas. However, it will take several decades for the soil characteristics to reach equivalency with the natural marshes, if they ever will.

Louisiana↗

USGS2012

Spatio-temporal variations in age structures of a partially re-established population of northern river otters ( Lontra canadensis )

Examination of age structures and sex ratios is useful in the management of northern river otters ( Lontra canadensis ) and other furbearers. Reintroductions and subsequent recolonizations of river otters have been well documented, but changes in demographics between expanding and established populations have not been observed. As a result of reintroduction efforts, immigration from Arkansas and northeastern Texas, and other efforts, river otters have become partially reestablished throughout eastern and central Oklahoma. Our objective was to examine age structures of river otters in Oklahoma and identify trends that relate to space (watersheds, county) and time (USDA Animal and Plant Health Inspection Service county trapping records). We predicted that river otters in western areas of the state were younger than river otters occurring farther east. From 2005–2007, we obtained salvaged river otter carcasses from federal and state agencies, and we live-captured other river otters using leg hold traps. Seventy-two river otters were sampled. Overall, sex ratios were skewed toward females (1F∶0.8M), but they did not differ among spatiotemporal scales examined. Teeth were removed from salvaged and live-captured river otters (n = 63) for aging. One-year old river otters represented the largest age class (30.2%). Proportion of juveniles (<1 y old) in Oklahoma (19.0%) was less than other states. Mean age of river otters decreased from east-to-west in the Arkansas River and its tributaries. Mean age of river otters differed between the Canadian River Watershed (0.8 y) and the Arkansas River Watershed (2.9 y) and the Canadian River Watershed and the Red River Watershed (2.4 y). Proportion of juveniles did not differ among spatiotemporal scales examined. Similar to age structure variations in other mammalian carnivores, colonizing or growing western populations of river otters in Oklahoma contained younger ages than more established eastern populations.

American Midland Naturalist↗

USGS2010

Population structure and relatedness among female Northern Pintails in three California wintering regions

Female Northern Pintails ( Anas acuta ) were sampled in California's three main Central Valley wintering regions (Sacramento Valley, Suisun Marsh, San Joaquin Valley) during September–October before most regional movements occur and microsatellite and mitochondrial DNA were analyzed to examine population structure and relatedness. Despite reportedly high rates of early-fall pairing and regional fidelity, both sets of markers indicated that there was little overall genetic structuring by region. Pintails from Suisun Marsh did exhibit higher relatedness among individuals and capture groups than in the Sacramento or San Joaquin Valleys, likely reflecting a sample comprised of a greater proportion of local breeders. The lack of genetic structuring among regions indicates that a high degree of movement and interchange occurs among pintails wintering in the Central Valley. Thus, although maintaining the existing distribution of pintails among Central Valley regions is important for other reasons, it does not appear to be critical to retain current patterns of population genetic variation. Because of potential lack of independence among highly related study individuals, researchers should consider regional differences in relatedness when designing sampling schemes and interpreting research findings.

California↗

USGS2005

Upper-crustal structure of the inner Continental Borderland near Long Beach, California

A new P-wave velocity/structural model for the inner Continental Borderland (ICB) region was developed for the area near Long Beach, California. It combines controlled-source seismic reflection and refraction data collected during the 1994 Los Angeles Region Seismic Experiment (LARSE), multichannel seismic reflection data collected by the U.S. Geological Survey (1998-2000), and nearshore borehole stratigraphy. Based on lateral velocity contrasts and stratigraphic variation determined from borehole data, we are able to locate major faults such as the Cabrillo, Palos Verdes, THUMS-Huntington Beach, and Newport Inglewood fault zones, along with minor faults such as the slope fault, Avalon knoll, and several other yet unnamed faults. Catalog seismicity (1975-2002) plotted on our preferred velocity/structural model shows recent seismicity is located on 16 out of our 24 faults, providing evidence for continuing concern with respect to the existing seismic-hazard estimates. Forward modeling of P-wave arrival times on the LARSE line 1 resulted in a four-layer model that better resolves the stratigraphy and geologic structures of the ICB and also provides tighter constraints on the upper-crustal velocity structure than previous modeling of the LARSE data. There is a correlation between the structural horizons identified in the reflection data with the velocity interfaces determined from forward modeling of refraction data. The strongest correlation is between the base of velocity layer 1 of the refraction model and the base of the planar sediment beneath the shelf and slope determined by the reflection model. Layers 2 and 3 of the velocity model loosely correlate with the diffractive crust layer, locally interpreted as Catalina Schist.

Bulletin of the Seismological Society of America↗

USGS2019

Rupture branching structure of the 2014 Mw 6.0 South Napa, California earthquake inferred from explosion-generated fault-zone trapped waves

We present evidence for multiple fault branches of the West Napa fault zone (WNFZ) based on fault‐zone trapped waves (FZTWs) generated by two explosions that were detonated within the main surface rupture zone produced by the 24 August 2014 Mw 6.0 South Napa earthquake. The FZTWs were recorded by a 15‐kilometer‐long dense (100 m spacing) linear seismic array consisting of 155 4.5‐hertz three‐component seismometers that were deployed across the surface ruptures and adjacent faults in Napa Valley in the summer of 2016. The two explosions were located ∼3.5 km north and ∼5 km south of the 2016 recording array. Prominent FZTWs, with large amplitudes and long wavetrains following the P and S waves, are observed on the seismograms. We analyzed FZTW waveforms in both time and frequency domains to characterize the branching structure of subsurface rupture zones along the WNFZ. The 2014 surface rupture zone was ∼12 km in length along the main trace of the WNFZ, which appears to form an ∼400–600‐meter‐wide low‐velocity waveguide to depths in excess of 5–7 km. Seismic velocities within the main rupture are reduced by 40%–50% relative to the surrounding‐rock velocities. Within 1.5 km of the main trace of the WNFZ, there are at least two subordinate fault traces that formed 3‐ to 6‐kilometer‐long surface breaks during the 2014 mainshock. Our modeling suggests that these subordinate fault traces are also low‐velocity waveguides that connect with the main rupture at depths of ∼2–3 km⁠ , forming a flower structure. FZTWs were also recorded at seismic stations across the Carneros fault (CF), which is ∼1 km west of the WNFZ; this suggests that the CF connects with the WNFZ at shallow depths, even though the CF did not experience surface rupture during the 2014 Mw 6.0 mainshock. 3D finite‐difference simulations of recorded FZTWs imply a branching structure along multiple fault strands associated with the WNFZ.

California↗

USGS1985

Crustal structure of the southern Calaveras fault zone, central California, from seismic refraction investigations

A magnitude 5.7 earthquake on 6 August 1979, within the Calaveras fault zone, near Coyote Lake of west-central California, motivated a seismic-refraction investigation in this area. A northwest-southeast profile along the fault, as well as two fan profiles across the fault were recorded to examine the velocity structure of this region. The analysis of the data reveals a complicated upper crustal velocity structure with strong lateral variations in all directions. The near-surface layers consist of recent sediments with seismic P -wave velocities of 2.6 to 3.2 km/sec. These are underlain by rocks of the Great Valley Sequence which have an average velocity of 4.5 km/sec. The Great Valley Sequence is present along the whole profile; depths range from 4.3 to 4.8 km in the northwest near Anderson Lake and in the southeast of the profile line near Hollister. In the middle of the profile near Coyote Lake, however, this layer thins and we find a laterally limited higher velocity layer (5.1 km/sec) between depths of 2.8 to 4.8 km. The high-velocity zone, which coincides with a gravity high, also correlates spatially with the hypocentral area of the 6 August earthquake and its aftershocks and may therefore represent an asperity on the fault. Velocities within the fault zone were determined from the fan profiles. Near Anderson Lake, a pronounced delay of first arrivals on the fan records indicates a vertical 1- to 2-km-wide near-surface, low-velocity zone along the fault. Near Coyote Lake, the delays observed in the fan records correlate with two subsurface en-echelon fault planes which have been previously identified from lineations in the seismicity pattern. The structure of the lower crust is similar to the neighboring Diablo Range: a 8- to 9-km-thick upper crustal layer with a seismic velocity of 5.7 to 6.3 km/sec is underlain by a 3-km-thick layer with velocity 6.8 km/sec. In accordance with previous studies of the Diablo Range, there are indications of a pronounced lower crustal low-velocity zone between a depth of 17 and 23 km. The presence of this low-velocity zone suggests that higher velocity (igneous?) rocks of the middle crust have been thrust over sedimentary rocks.

California↗