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At least 595 records · Page 33Linked to original sources

Avian movements and wetland connectivity in landscape conservation

The current conservation crisis calls for research and management to be carried out on a long-term, multi-species basis at large spatial scales. Unfortunately, scientists, managers, and agencies often are stymied in their effort to conduct these large-scale studies because of a lack of appropriate technology, methodology, and funding. This issue is of particular concern in wetland conservation, for which the standard landscape approach may include consideration of a large tract of land but fail to incorporate the suite of wetland sites frequently used by highly mobile organisms such as waterbirds (e.g., shorebirds, wading birds, waterfowl). Typically, these species have population dynamics that require use of multiple wetlands, but this aspect of their life history has often been ignored in planning for their conservation. We outline theoretical, empirical, modeling, and planning problems associated with this issue and suggest solutions to some current obstacles. These solutions represent a tradeoff between typical in-depth single-species studies and more generic multi-species studies. They include studying within- and among-season movements of waterbirds on a spatial scale appropriate to both widely dispersing and more stationary species; multi-species censuses at multiple sites; further development and use of technology such as satellite transmitters and population-specific molecular markers; development of spatially explicit population models that consider within-season movements of waterbirds; and recognition from funding agencies that landscape-level issues cannot adequately be addressed without support for these types of studies.

Conservation Biology↗

A spatial model to prioritize sagebrush landscapes in the intermountain west (U.S.A.) for restoration

The ecological integrity of Sagebrush (Artemisia spp.) ecosystems in the Intermountain West (U.S.A.) has been diminished by synergistic relationships among human activities, spread of invasive plants, and altered disturbance regimes. An aggressive effort to restore Sagebrush habitats is necessary if we are to stabilize or improve current habitat trajectories and reverse declining population trends of dependent wildlife. Existing economic resources, technical impediments, and logistic difficulties limit our efforts to a fraction of the extensive area undergoing fragmentation, degradation, and loss. We prioritized landscapes for restoring Sagebrush habitats within the intermountain western region of the United States using geographic information system (GIS) modeling techniques to identify areas meeting a set of conditions based on (1) optimum abiotic and biotic conditions favorable for revegetation of Sagebrush; (2) potential to increase connectivity of Sagebrush habitats in the landscape to benefit wildlife; (3) location of population strongholds for Greater Sage-Grouse (Centrocercus urophasianus, a species of conservation concern); and (4) potential impediments to successful restoration created by Cheatgrass (Bromus tectorum, an invasive exotic annual grass). Approximately 5.8 million ha in southwestern Idaho, northern Nevada, and eastern Oregon met our criteria for restoring Wyoming big sagebrush (Artemisia tridentata ssp. wyomingensis) and 5.1 million ha had high priority for restoring Mountain big sagebrush (A. tridentata ssp. vaseyana). Our results represent an integral component in a hierarchical framework after which site-specific locations for treatments can be focused within high-priority areas. Using this approach, long-term restoration strategies can be implemented that combine local-scale treatments and objectives with large-scale ecological processes and priorities. ?? 2008 Society for Ecological Restoration International.

Restoration Ecology↗

Direct and indirect effects of land use on floral resources and flower-visiting insects across an urban landscape

Although urban areas are often considered to have uniformly negative effects on biodiversity, cities are most accurately characterized as heterogeneous mosaics of buildings, streets, parks, and gardens that include both ‘good’ and ‘bad’ areas for wildlife. However, to date, few studies have evaluated how human impacts vary in direction and magnitude across a heterogeneous urban landscape. In this study, we assessed the distribution of floral resources and flower-visiting insects across a variety of land uses in New York City. We visited both green spaces (e.g. parks, cemeteries) and heavily developed neighborhood blocks (e.g. with high or low density residential zoning) and used structural equation modeling (SEM) to evaluate the direct and indirect effects of median income, vegetation, and development intensity on floral resources and insects in both settings. Abundance and taxonomic richness of flower-visiting insects was significantly greater in green spaces than neighborhood blocks. The SEM results indicated that heavily-developed neighborhoods generally had fewer flower-visiting insects consistent with reductions in floral resources. However, some low-density residential neighborhoods maintained high levels of floral resources and flower-visiting insects. We found that the effects of surrounding vegetation on floral resources, and thus indirect effects on insects, varied considerably between green spaces and neighborhood blocks. Along neighborhood blocks, vegetation consisted of a mosaic of open gardens and sparsely distributed trees and had a positive indirect effect on flower-visiting insects. In contrast, vegetation in urban green spaces was associated with increased canopy cover and thus had a negative indirect effect on flower-visiting insects through reductions in floral resources. In both neighborhood blocks and green spaces, vegetation had a positive direct effect on flower-visiting insects independent of the influence of vegetation on floral resources. Our results demonstrate how inter-related components of an urban ecosystem can vary with respect to one another across a heterogeneous urban landscape, suggesting that it is inappropriate to generalize about urban systems as a whole without first addressing differences among component land use types.

Oikos↗

Vegetation change over 140 years in a sagebrush landscape of the Rio Grande del Norte National Monument, New Mexico, USA

Questions Big sagebrush ( Artemisia tridentata ) ecosystems across the western United States have experienced many changes in ecosystem dynamics and vegetation composition over the last century due to livestock grazing, non-native species, and changing climate and fire regimes. We conducted the first systematic investigation of historical vegetation composition and vegetation change in a sagebrush landscape in the southwestern United States, asking whether sagebrush or grass dominated the landscape historically? Location The Rio Grande del Norte National Monument (RGDN), northern New Mexico, USA. Methods We combined General Land Office (GLO) surveys from 1881 with modern vegetation maps, field vegetation surveys, and sagebrush ages from growth ring analysis to test for changes in vegetation in the RGDN over the last 140 years. Results We found that big sagebrush presence across the study area increased significantly, from being present on 16% of section lines in 1881 to 79% in 2019, and only three section lines lost sagebrush presence during that period. Concurrently, the number of section lines with low grass index more than doubled since 1881, while moderate and high grass index declined. Grass declined equally in areas where sagebrush increased and areas with no change in sagebrush, suggesting that changes in both vegetation types were catalyzed by external factors, likely including overgrazing. The growth ring analysis of 93 sagebrush revealed a maximum age of 87 years and establishment in every decade since the 1930s, consistent with the GLO results. Conclusions The significant vegetation changes in the RGDN over the last century, including an increase of sagebrush, provide important context about the shifting mosaic of grasslands and shrublands relevant to current and future management and ecosystem dynamics.

New Mexico↗

Interacting effects of human presence and landscape modification on birds and mammals

Sustainable human–wildlife coexistence requires a mechanistic understanding of the many ways that humans affect animals. However, progress is hampered by the lack of accessible data measuring the dynamic presence of people. Here, we leverage mobile-device data to disentangle how human presence and landscape modification differentially influence the use of geographic and environmental space for 37 mammal and bird species across the United States. Human presence affected more than 65% of species, with substantial variation across species. For ~60% of species that responded to human activities, the effects were interdependent—animals tended to react more strongly to human presence in less modified habitats. Our results demonstrate that human presence and landscape modification have complex combined effects on wildlife, which need to be considered for effective management.

Science↗

Vegetation composition, nutrient, and sediment dynamics along a floodplain landscape

Forested floodplains are important landscape features for retaining river nutrients and sediment loads but there is uncertainty in how vegetation influences nutrient and sediment retention. In order to understand the role of vegetation in nutrient and sediment trapping, we quantified species composition and the uptake of nutrients in plant material relative to landscape position and ecosystem attributes in an urban, Piedmont watershed in Virginia, USA. We investigated in situ interactions among vegetative composition, abundance, carbon (C), nitrogen (N) and phosphorus (P) fluxes and ecosystem attributes such as water level, shading, soil nutrient mineralization, and sediment deposition. This study revealed strong associations between vegetation and nutrient and sediment cycling processes at the plot scale and in the longitudinal dimension, but there were few strong patterns between these aspects at the scale of geomorphic features (levee, backswamp, and toe-slope). Patterns reflected the nature of the valley setting rather than a simple downstream continuum. Plant nutrient uptake and sediment trapping were greatest at downstream sites with the widest floodplain and lowest gradient where the hydrologic connection between the floodplain and stream is greater. Sediment trapping increased in association with higher herbaceous plant coverage and lower tree canopy density that, in turn, was associated with a more water tolerant tree community found in the lower watershed but not at the most downstream site in the watershed. Despite urbanization effects on the hydrology, this floodplain functioned as an efficient nutrient trap. N and P flux rates of herbaceous biomass and total litterfall more than accounted for the N and P mineralization flux rate, indicating that vegetation incorporated nearly all mineralized nutrients into biomass.

Virginia↗

Eroding Cascadia— Sediment and solute transport and landscape denudation in western Oregon and northwestern California

Riverine measurements of sediment and solute transport give empirical basin-scale estimates of bed-load, suspended-sediment, and silicate-solute fluxes for 100,000 km 2 of northwestern California and western Oregon. This spatially explicit sediment budget shows the multifaceted control of geology and physiography on the rates and processes of fluvial denudation. Bed-load transport is greatest for steep basins, particularly in areas underlain by the accreted Klamath terrane. Bed-load flux commonly decreases downstream as clasts convert to suspended load by breakage and attrition, particularly for softer rock types. Suspended load correlates strongly with lithology, basin slope, precipitation, and wildfire disturbance. It is highest in steep regions of soft rocks, and our estimates suggest that much of the suspended load is derived from bed-load comminution. Dissolution, measured by basin-scale silicate-solute yield, constitutes a third of regional landscape denudation. Solute yield correlates with precipitation and is proportionally greatest in low-gradient and wet basins and for high parts of the Cascade Range, where undissected Quaternary volcanic rocks soak in 2−3 m of annual precipitation. Combined, these estimates provide basin-scale erosion rates ranging from ∼50 t ∙ km −2 ∙ yr −1 (approximately equivalent to 0.02 mm ∙ yr −1 ) for low-gradient basins such as the Willamette River to ∼500 t ∙ km −2 ∙ yr −1 (∼0.2 mm ∙ yr −1 ) for steep coastal drainages. The denudation rates determined here from modern measurements are less than those estimated by longer-term geologic assessments, suggesting episodic disturbances such as fire, flood, seismic shaking, and climate change significantly add to long-term landscape denudation.

California, Oregon, Washington↗

New thermochronometric constraints on the Tertiary landscape evolution of the central and eastern Grand Canyon, Arizona

Thermal histories are modeled from new apatite (U-Th)/He and apatite fission-track data in order to quantitatively constrain the landscape evolution of the Grand Canyon region. Fifty new samples and their associated thermochronometric ages are presented here. Samples span from Lee’s Ferry in the east to Quartermaster Canyon in the west and include four age-elevation transects into Grand Canyon and borehole samples from the Coconino Plateau. Twenty-seven samples are inversely modeled to provide continuous thermal histories. This represents the most extensive and complete dataset on patterns of long-term exhumation in the Grand Canyon region, and it enables us to constrain the timing and magnitude of erosion and also discriminate between canyon incision and broader planation. The new data suggest that the early Cenozoic landscape in eastern Grand Canyon was low in relief and does not indicate the presence of an early Cenozoic precursor to the modern Grand Canyon. However, there is evidence for the incision of a smaller-scale canyon across the Kaibab Uplift at 28–20 Ma. This middle-Cenozoic denudation event was accompanied by the removal of a majority of remaining Mesozoic strata west of the Kaibab Uplift. In contrast, just upstream in the area of Lee’s Ferry, ∼2 km of Mesozoic strata remained over the middle Cenozoic and were removed after 10 Ma.

Arizona↗

Landscape-scale evaluation of genetic structure among barrier-isolated populations of coastal cutthroat trout, Oncorhynchus clarkii clarkii

Relationships among landscape structure, stochastic disturbance, and genetic diversity were assessed by examining interactions between watershed-scale environmental factors and genetic diversity of coastal cutthroat trout (Oncorhynchus clarkii clarkii) in 27 barrier-isolated watersheds from western Oregon, USA. Headwater populations of coastal cutthroat trout were genetically differentiated (mean FST = 0.33) using data from seven microsatellite loci (2232 individuals), but intrapopulation microsatellite genetic diversity (mean number of alleles per locus = 5, mean He = 0.60) was only moderate. Genetic diversity of coastal cutthroat trout was greater (P = 0.02) in the Coast Range ecoregion (mean alleles = 47) than in the Cascades ecoregion (mean alleles = 30), and differences coincided with indices of regional within-watershed complexity and connectivity. Furthermore, regional patterns of diversity evident from isolation-by-distance plots suggested that retention of within-population genetic diversity in the Coast Range ecoregion is higher than that in the Cascades, where genetic drift is the dominant factor influencing genetic patterns. Thus, it appears that physical landscape features have influenced genetic patterns in these populations isolated from short-term immigration. ?? 2008 NRC.

Canadian Journal of Fisheries and Aquatic Sciences↗

Cold-climate slope deposits and landscape modifications of the Mid-Atlantic Coastal Plain, Eastern USA

The effects of Pleistocene cold-climate geomorphology are distributed across the weathered and eroded Mid-Atlantic Coastal Plain uplands from the Wisconsinan terminal moraine south to Tidewater Virginia. Cold-climate deposits and landscape modifications are superimposed on antecedent landscapes of old, weathered Neogene upland gravels and Pleistocene marine terraces that had been built during warm periods and sea-level highstands. In New Jersey, sequences of surficial deposits define a long history of repeating climate change events. To the south across the Delmarva Peninsula and southern Maryland, most antecedent topography has been obscured by Late Pleistocene surficial deposits. These are spatially variable and are collectively described as a cold-climate alloformation. The cold-climate alloformation includes time-transgressive details of climate deterioration from at least marine isotope stage (MIS) 4 through the end of MIS 2. Some deposits and landforms within the alloformation may be as young as the Younger Dryas. Southwards along the trend of the Potomac River, these deposits and their climatic affinities become diffused. In Virginia, a continuum of erosion and surficial deposits appears to be the product of ‘normal’ temperate, climate-forced processes. The cold-climate alloformation and more temperate deposits in Virginia are being partly covered by Holocene alluvium and bay mud.

Geological Society Special Publication↗

Cross-scale modeling of surface temperature and tree seedling establishment inmountain landscapes

Abstract: Introduction: Estimating surface temperature from above-ground field measurements is important for understanding the complex landscape patterns of plant seedling survival and establishment, processes which occur at heights of only several centimeters. Currently, future climate models predict temperature at 2 m above ground, leaving ground-surface microclimate not well characterized. Methods: Using a network of field temperature sensors and climate models, a ground-surface temperature method was used to estimate microclimate variability of minimum and maximum temperature. Temperature lapse rates were derived from field temperature sensors and distributed across the landscape capturing differences in solar radiation and cold air drainages modeled at a 30-m spatial resolution. Results: The surface temperature estimation method used for this analysis successfully estimated minimum surface temperatures on north-facing, south-facing, valley, and ridgeline topographic settings, and when compared to measured temperatures yielded an R2 of 0.88, 0.80, 0.88, and 0.80, respectively. Maximum surface temperatures generally had slightly more spatial variability than minimum surface temperatures, resulting in R2 values of 0.86, 0.77, 0.72, and 0.79 for north-facing, south-facing, valley, and ridgeline topographic settings. Quasi-Poisson regressions predicting recruitment of Quercus kelloggii (black oak) seedlings from temperature variables were significantly improved using these estimates of surface temperature compared to air temperature modeled at 2 m. Conclusion: Predicting minimum and maximum ground-surface temperatures using a downscaled climate model coupled with temperature lapse rates estimated from field measurements provides a method for modeling temperature effects on plant recruitment. Such methods could be applied to improve projections of species’ range shifts under climate change. Areas of complex topography can provide intricate microclimates that may allow species to redistribute locally as climate changes.

Ecological Processes↗

Potential influence of wildfire in modulating climate-induced forest redistribution in a central Rocky Mountain landscape

Introduction Climate change is expected to impose significant tension on the geographic distribution of tree species. Yet, tree species range shifts may be delayed by their long life spans, capacity to withstand long periods of physiological stress, and dispersal limitations. Wildfire could theoretically break this biological inertia by killing forest canopies and facilitating species redistribution under changing climate. We investigated the capacity of wildfire to modulate climate-induced tree redistribution across a montane landscape in the central Rocky Mountains under three climate scenarios (contemporary and two warmer future climates) and three wildfire scenarios (representing historical, suppressed, and future fire regimes). Methods Distributions of four common tree species were projected over 90 years by pairing a climate niche model with a forest landscape simulation model that simulates species dispersal, establishment, and mortality under alternative disturbance regimes and climate scenarios. Results Three species (Douglas-fir, lodgepole pine, subalpine fir) declined in abundance over time, due to climate-driven contraction in area suitable for establishment, while one species (ponderosa pine) was unable to exploit climate-driven expansion of area suitable for establishment. Increased fire frequency accelerated declines in area occupied by Douglas-fir, lodgepole pine, and subalpine fir, and it maintained local abundance but not range expansion of ponderosa pine. Conclusions Wildfire may play a larger role in eliminating these conifer species along trailing edges of their distributions than facilitating establishment along leading edges, in part due to dispersal limitations and interspecific competition, and future populations may increasingly depend on persistence in locations unfavorable for their establishment.

Idaho↗

Chapter 4. Predicting post-fire erosion and sedimentation risk on a landscape scale

Historic fire suppression efforts have increased the likelihood of large wildfires in much of the western U.S. Post-fire soil erosion and sedimentation risks are important concerns to resource managers. In this paper we develop and apply procedures to predict post-fire erosion and sedimentation risks on a pixel-, catchment-, and landscape-scale in central and western Colorado. Our model for predicting post-fire surface erosion risk is conceptually similar to the Revised Universal Soil Loss Equation (RUSLE). One key addition is the incorporation of a hydrophobicity risk index (HY-RISK) based on vegetation type, predicted fire severity, and soil texture. Post-fire surface erosion risk was assessed for each 90-m pixel by combining HYRISK, slope, soil erodibility, and a factor representing the likely increase in soil wetness due to removal of the vegetation. Sedimentation risk was a simple function of stream gradient. Composite surface erosion and sedimentation risk indices were calculated and compared across the 72 catchments in the study area. When evaluated on a catchment scale, two-thirds of the catchments had relatively little post-fire erosion risk. Steeper catchments with higher fuel loadings typically had the highest post-fire surface erosion risk. These were generally located along the major north-south mountain chains and, to a lesser extent, in west-central Colorado. Sedimentation risks were usually highest in the eastern part of the study area where a higher proportion of streams had lower gradients. While data to validate the predicted erosion and sedimentation risks are lacking, the results appear reasonable and are consistent with our limited field observations. The models and analytic procedures can be readily adapted to other locations and should provide useful tools for planning and management at both the catchment and landscape scale.

Journal of Sustainable Forestry↗

Honey bee (Apis mellifera) colonies benefit from grassland/ pasture while bumble bee (Bombus impatiens) colonies in the same landscapes benefit from non-corn/soybean cropland

Agriculturally important commercially managed pollinators including honey bees ( Apis mellifera L., 1758) and bumble bees ( Bombus impatiens Cresson, 1863) rely on the surrounding landscape to fulfill their dietary needs. A previous study in Europe demonstrated that managed honey bee foragers and unmanaged native bumble bee foragers are associated with different land uses. However, it is unclear how response to land use compares between managed honey bees and a managed native bumble bee species in the United States, where honey bees are an imported species. Furthermore, to our knowledge, no such direct comparisons of bee responses to land use have been made at the colony level. To better understand how two different social bees respond to variation in land use, we monitored the weights of A . mellifera and B . impatiens colonies placed in 12 apiaries across a range of land use in Michigan, United States in 2017. Bombus impatiens colonies gained more weight and produced more drones when surrounded by diverse agricultural land (i.e., non-corn/soybean cropland such as tree fruits and grapes), while honey bee colonies gained more weight when surrounded by more grassland/pasture land. These findings add to our understanding of how different bee species respond to agricultural landscapes, highlighting the need for further species-specific land use studies to inform tailored land management.

Michigan↗

Influence of trees in the landscape on parasitism rates of grassland passerine nests in Southeastern North Dakota

Woody vegetation has been linked to increased rates of Brown-headed Cowbird ( Molothrus ater ) parasitism for some grassland hosts. In northern North Dakota, however, studies reported that parasitism of grassland passerine nests was lower in landscapes with trees than in those without trees. We looked for evidence of this pattern elsewhere, using data from two studies conducted on the Sheyenne National Grassland in southeastern North Dakota. Specifically, we examined the probability of parasitism relative to percent tree cover within 2 km of a nest. We found a negative relationship for grassland passerine nests of all species tested. Our results support the suggestion that cowbirds are less likely to parasitize nests of grassland passerines where tree cover on the landscape is greater. This pattern could be explained by cowbirds switching to alternative hosts in woodlands, but this hypothesis needs further testing.

Condor↗

Ecology and conservation of a rare, old-growth-associated canopy lichen in a silvicultural landscape

Nephroma occultum Wetm. is a rare, epiphytic lichen associated with old-growth forests of northwestern North America. We describe its distribution, abundance, and habitat within the managed landscape of a southwestern Oregon watershed. Because this species is found mainly in the canopy, we used direct canopy access (tree climbing) in combination with ground (litter) searches for our surveys. We recommend this dual approach when confident determination of presence or absence of a canopy species is needed. Our surveys confirm that N. occultum is a rare old-growth associate within the area of this study, the southernmost extension of its known range. It was both rare across the landscape and uncommon in our primary study site, a 500 yr old stand. It was most often found growing close to the trunk on branches of large Pseudotsuga menziesii , but its distribution was sporadic even within an old-growth stand. The scarcity of very old stands, in combination with the limited ability of this species to disperse within and between stands, has likely contributed to its rarity within the watershed. Management for N. occultum should focus on populations and habitat needs rather than on individuals. Our calculations show that cutting with retention of individual trees surrounded by small buffers could result in the eventual loss of N. occultum from the study area.

Oregon↗

Local adaptation in Trinidadian guppies alters stream ecosystem structure at landscape scales despite high environmental variability

While previous studies have shown that evolutionary divergence alters ecological processes in small-scale experiments, a major challenge is to assess whether such evolutionary effects are important in natural ecosystems at larger spatial scales. At the landscape scale, across eight streams in the Caroni drainage, we found that the presence of locally adapted populations of guppies ( Poecilia reticulata ) is associated with reduced algal biomass and increased invertebrate biomass, while the opposite trends were true in streams with experimentally introduced populations of non-locally adapted guppies. Exclusion experiments conducted in two separate reaches of a single stream showed that guppies with locally adapted phenotypes significantly reduced algae with no effect on invertebrates, while non-adapted guppies had no effect on algae but significantly reduced invertebrates. These divergent effects of phenotype on stream ecosystems are comparable in strength to the effects of abiotic factors (e.g., light) known to be important drivers of ecosystem condition. They also corroborate the results of previous experiments conducted in artificial streams. Our results demonstrate that local adaptation can produce phenotypes with significantly different effects in natural ecosystems at a landscape scale, within a tropical watershed, despite high variability in abiotic factors: five of the seven physical and chemical parameters measured across the eight study streams varied by more than one order of magnitude. Our findings suggest that ecosystem structure is, in part, an evolutionary product and not simply an ecological pattern.

Copeia↗

Response of Tiger Salamanders (Ambystoma t. tigrinum) to wetland restoration in a midwestern agricultural landscape, U.S.A.

Since the early 1990s, > 3,000 ha of wetlands (and adjacent prairie) have been restored on the row-crop agricultural landscape of Winnebago County, Iowa, U.S.A. From 2014–2016, we surveyed 45 wetlands among 19 easements for occupancy by Eastern Tiger Salamanders ( Ambystoma tigrinum tigrinum ) and used radio-telemetry to measure their patterns of movement and habitat use. Rates of occupancy increased with wetland age, from < 25% for wetlands 1–2 years old to ∼75% for wetlands > 11 years old. A two-year survey (2014 and 2015) of ten wetlands restored in 2013 showed that nine were occupied after two years; we did not find a relationship between distance to the nearest salamander population and occupancy of newly restored wetlands by salamanders. We tracked 30 salamanders after they left their breeding wetlands for an average of 69±37 d (range = 14–109 d) and relocated them a total of 393 times. Typically, once a salamander left its breeding wetland, it traveled 50–350 m over several days, found a suitable burrow, then remained for much of the rest of the season. Mean daily distances traveled by salamanders were 7.9±5.6 m (range = 0–135 m); the range of maximum straight-line distances moved was 26–659 m; only one individual salamander traveled in a statistically linear path, relative to a random walk. While ∼90% of the landscape was composed of row-crop fields, salamanders used protective grassy habitats (e.g., restored prairie, road ditches) on ∼88% of our observations. Only three salamanders used row-crop fields, and two of them were killed by heavy equipment. Regardless of the terrestrial habitat types used by salamanders, we found them underground on 336 (84.8%) of our observations.

Iowa↗