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Vascular flora of saline lakes in the southern high plains of Texas and eastern New Mexico

Saline lakes and freshwater playas form the principal surface hydrological feature of the High Plains of the Southern Great Plains. Saline lakes number less than 50 and historically functioned as discharge wetlands with relatively consistent water availability due to the presence of one or more springs. Currently, less than ten saline lakes contain functional springs. A survey of vascular plants at six saline lakes in the Southern High Plains of northwest Texas and one in eastern New Mexico during May and September 2009 resulted in a checklist of 49 species representing 16 families and 40 genera. The four families with the most species were Asteraceae (12), Amaranthaceae (8), Cyperaceae (5), and Poaceae (12). Non-native species (Bromus catharticus, Poa compressa, Polypogon monspeliensis, Sonchus oleraceus, Kochia scoparia, and Tamarix ramosissima) accounted for 10% of the total species recorded. Whereas nearly 350 species of vascular plants have been identified in playas in the Southern High Plains, saline lakes contain a fraction of this species richness. The Southern High Plains saline lake flora is regionally unique, containing taxa not found in playas, with species composition that is more similar to temperate desert wetlands of the Intermountain Region and Gulf Coastal Plain of North America.

New Mexico, Texas

Integrated rangeland fire management strategy actionable science plan completion assessment: Restoration topic, 2015–20

Loss and degradation of sagebrush rangelands due to an accelerated invasive annual grass-wildfire cycle and other stressors are substantial management, conservation, and economic issues in the western United States. These sagebrush rangelands comprise a unique biome spanning 11 states, support over 350 wildlife species, and provide important ecosystem services that include stabilizing the economies of western communities. Impacts to sagebrush ecosystem processes over large areas due to the annual grass-wildfire cycle necessitated the development of a coordinated, science-based strategy for improving efforts to achieve long-term protection, conservation, and restoration of sagebrush rangelands, which was framed in 2015 under the Integrated Rangeland Fire Management Strategy (IRFMS). Central to this effort was the development of an Actionable Science Plan (Plan) that identified 37 priority science needs (Needs) for informing the actions proposed under the 5 topics (Fire, Invasives, Restoration, Sagebrush and Sage-Grouse, Climate and Weather) that were part of the collective focus of the IRFMS. Notable keys to this effort were identification of the Needs co-produced by managers and researchers, and a focus on resulting science being “actionable.” Substantial investments aimed at fulfilling the Needs identified in the Plan have been made since its release in 2016. While the state of the science has advanced considerably, the extent to which knowledge gaps remain relative to identified Needs is relatively unknown. Moreover, new Needs have likely emerged since the original strategy as results from actionable science reveal new questions, and possible (yet untested) solutions. A quantifiable assessment of the progress made on the original science Needs can identify unresolved gaps and new information that can help inform prioritization of future research efforts. This report details a systematic literature review that evaluated how well peer-reviewed journal articles and formal technical reports published between January 1, 2015, and December 31, 2020, addressed 10 needs (hereinafter “Needs”) identified under the Restoration topic in the Plan. The topic outlined research Needs for improving restoration success in degraded sagebrush rangelands. We established the level of progress towards addressing each Need following a standardized set of criteria, and developed summaries detailing how research objectives nested within Needs identified in the Plan (“Next Steps”) were either addressed well, partially addressed or remain outstanding (that is, addressed poorly) in the literature through 2020. Our searches resulted in the inclusion of 371 science products that at least partially addressed a Need identified in the Restoration topic. The Needs that were well and partially addressed included: the development of methods to improve seeding success of native plants after fire; evaluation of short- and long-term plant responses relative to biotic and abiotic factors following restoration treatments; assessments of long-term effects on ecosystem processes following conifer removal treatments and development of decision support tools to aid in conifer management; determination of the costs of restoration treatments relative to the benefits to sage-grouse occupancy, survival, and populations; and assessments of the factors which contribute to soil degradation and development of soil enhancement methods that enhance seed germination and seedling survival while resisting nonnative plants. Needs that were not addressed well included: the development of methods for the rapid recovery of sagebrush and native herbaceous plants following wildfire and determination of whether treatments benefit sage-grouse and mitigate impacts from fires; determination of thresholds for successful restoration across a range of environmental conditions and development of decision support tools to assist with restoration effectiveness monitoring; and development and evaluation of the utility of seed transfer zones. The information provided in this assessment will assist updating the Plan along with other science strategies.

western United States

Environmental tolerance of an invasive riparian tree and its potential for continued spread in the southwestern US

Questions: Exotic plant invasion may be aided by facilitation and broad tolerance of environmental conditions, yet these processes are poorly understood in species-rich ecosystems such as riparian zones. In the southwestern United States (US) two plant species have invaded riparian zones: tamarisk (Tamarix ramosissima, T. chinensis, and their hybrids) and Russian olive (Elaeagnus angustifolia). We addressed the following questions: (1) is Russian olive able to tolerate drier and shadier conditions than cottonwood and tamarisk? (2) Can tamarisk and cottonwood facilitate Russian olive invasion? Location: Arid riparian zones, southwestern US. Methods: We analyzed riparian tree seedling requirements in a controlled experiment, performed empirical field studies, and analyzed stable oxygen isotopes to determine the water sources used by Russian olive. Results: Russian olive survival was significantly higher in dense shade and low moisture conditions than tamarisk and cottonwood. Field observations indicated Russian olive established where flooding cannot occur, and under dense canopies of tamarisk, cottonwood, and Russian olive. Tamarisk and native riparian plant species seedlings cannot establish in these dry, shaded habitats. Russian olive can rely on upper soil water until 15 years of age, before utilizing groundwater. Conclusions: We demonstrate that even though there is little evidence of facilitation by cottonwood and tamarisk, Russian olive is able to tolerate dense shade and low moisture conditions better than tamarisk and cottonwood. There is great potential for continued spread of Russian olive throughout the southwestern US because large areas of suitable habitat exist that are not yet inhabited by this species. ?? 2010 International Association for Vegetation Science.

Journal of Vegetation Science

Fine-resolution land cover mapping over large and mountainous areas for Lāna‘i, Hawaii using posterior probabilities, and expert knowledge

The task of accurately mapping species-specific vegetation cover in remote and topographically complex regions like those found in Hawaiʻi presents unique challenges. This study leverages a machine learning approach to accurately classify vegetation into fine species-specific classes across the island of Lāna‘i, Hawaii, offering a novel methodology for tackling such challenges. Utilizing high-resolution WordView-2 satellite imagery, a neural network classifier and a custom lidar-based geometric correction, we introduced two new approaches to refine our high-resolution land cover classifications. This included the implementation of prior-based adjustments to class posterior probabilities to enhance land cover classification accuracy. Moreover, we developed mixed hierarchical classification maps that use class posterior probabilities to identify, at the pixel level, the finest land cover class that meets a user-defined confidence threshold. The resulting high-resolution land cover map for Lāna‘i captures the rich diversity and distribution of native and invasive plant species with high overall accuracy, generally exceeding 95%, based on independent ground control data. The capacity to produce wall-to-wall species-level vegetation maps provides a new window into monitoring vegetation dynamics on Lāna‘i and similarly remote and topographically complex regions, and contributes to our broader understanding of ecosystem responses to invasive species, climatic changes, and land management practices such as erosion and sediment control planning. Our approach offers a blueprint for similar efforts in other complex and remote ecosystems.

Hawaii

Biogeochemical and ecological impacts of livestock grazing in semi-arid southeastern Utah, USA

Relatively few studies have examined the ecological and biogeochemical effects of livestock grazing in southeastern Utah. In this study, we evaluated how grazing has affected soil organic carbon and nitrogen to a depth of 50 cm in grasslands located in relict and actively-grazed sites in the Canyonlands physiographic section of the Colorado Plateau. We also evaluated differences in plant ground cover and the spatial distribution of soil resources. Results show that areas used by domestic livestock have 20% less plant cover and 100% less soil organic carbon and nitrogen compared to relict sites browsed by native ungulates. In actively grazed sites, domestic livestock grazing also appears to lead to clustered, rather than random, spatial distribution of soil resources. Magnetic susceptibility, a proxy for soil stability in this region, suggests that grazing increases soil erosion leading to an increase in the area of nutrient-depleted bare ground. Overall, these results, combined with previous studies in the region, suggest that livestock grazing affects both plant cover and soil fertility with potential long-term implications for the sustainability of grazing operations in this semi-arid landscape. ?? 2007 Elsevier Ltd. All rights reserved.

Journal of Arid Environments

Habitat and landscape effects on abundance of Missouri's grassland birds

Of 6 million ha of prairie that once covered northern and western Missouri, <36,500&thinsp;ha remain, with planted, managed, and restored grasslands comprising most contemporary grasslands. Most grasslands are used as pasture or hayfields. Native grasses largely have been replaced by fescue ( Festuca spp.) on most private lands (almost 7 million ha). Previously cropped fields set aside under the Conservation Reserve Program (CRP) varied from a mix of cool-season grasses and forbs, or mix of native warm-season grasses and forbs, to simple tall-grass monocultures. We used generalized linear mixed models and distance sampling to assess abundance of 8 species of breeding grassland birds on 6 grassland types commonly associated with farm practices in Missouri and located in landscapes managed for grassland-bird conservation. We selected Bird Conservation Areas (BCAs) for their high percentage of grasslands and grassland-bird species, and for <5% forest cover. We used an information-theoretic approach to assess the relationship between bird abundance and 6 grassland types, 3 measures of vegetative structure, and 2 landscape variables (% grassland and edge density within a 1-km radius). We found support for all 3 levels of model parameters, although there was less support for landscape than vegetation structure effects likely because we studied high-percentage-grassland landscapes (BCAs). Henslow's sparrow ( Ammodramus henslowii ) counts increased with greater percentage of grassland, vegetation height-density, litter depth, and shrub cover and lower edge density. Henslow's sparrow counts were greatest in hayed native prairie. Dickcissel ( Spiza americana ) counts increased with greater vegetation height-density and were greatest in planted CRP grasslands. Grasshopper sparrow ( A. savannarum ) counts increased with lower vegetation height, litter depth, and shrub cover. Based on distance modeling, breeding densities of Henslow's sparrow, dickcissel, and grasshopper sparrow in the 6 grassland types ranged 0.9&ndash;2.6, 1.4&ndash;3.2, and 0.1&ndash;1.5&thinsp;birds/ha, respectively. We suggest different grassland types and structures (vegetation height, litter depth, shrub cover) are needed to support priority grassland-bird species in Missouri.

Journal of Wildlife Management

Grazing effects on plant community succession of early- and mid-seral seeded grassland compared to shortgrass steppe

Questions: Grazing may speed or slow secondary succession, and the direction may depend on seral stage and relative tolerance of native perennial grasses compared with annual invasive species. How does grazing affect succession where undisturbed communities have a long evolutionary history of grazing by native herbivores and are tolerant to livestock grazing? Location: Shortgrass steppe, North American Great Plains, Colorado (40°49′N, 104°46′W), USA. Methods: Ungrazed and grazed early-seral (4–6 yr) and mid-seral (18–20 yr) seeded fields (Conservation Reserve Program) and traditionally grazed native steppe were compared for effects on plant composition in relation to changes expected from regional succession models. Results: Recovery towards undisturbed native shortgrass steppe for early- and mid-seral communities, respectively, was 19% and 36% for total vegetation cover, 5% and 21% for planted native species, 10% and 88% for non-planted native perennial grasses, only 0.2% and 13% for short grasses, and overall dissimilarity in community species compositions was 97% and 68%. In general, grazing effects were neutral or most often not significant in all years and/or were small in overall community magnitude. The early-seral community displayed more changes indicative of a slowing of succession with grazing (total vegetative and grass basal cover) rather than reducing invasive species (species targeted by timing of grazing), although drought had limited the establishment of grazing-tolerant short grasses. The mid-seral community showed more changes consistent with advancing successional recovery with grazing (overall community dissimilarity, forbs, planted native perennial grasses, tall grasses and target species). However, non-planted native perennial grasses responded negatively to grazing in the mid-seral community and positively in native shortgrass steppe where outside seed would originate. Conclusions: Grazing effects on particular functional groups and species were not the same across seral stages, were mixed in terms of speeding or slowing succession, and were generally not large at the community level. Evolutionary history of grazing may serve as a general guide but decisions on whether to graze successional grasslands may best be made after assessing whether tolerant perennial short grass species are significant components. Monitoring may then be necessary to determine species responses in particular community matrixes and effects on subsequent immigration of non-seeded native perennial species.

Colorado

Potential environmental effects of pack stock on meadow ecosystems of the Sierra Nevada, USA

Pack and saddle stock, including, but not limited to domesticated horses, mules, and burros, are used to support commercial, private and administrative activities in the Sierra Nevada. The use of pack stock has become a contentious and litigious issue for land management agencies in the region inter alia due to concerns over effects on the environment. The potential environmental effects of pack stock on Sierra Nevada meadow ecosystems are reviewed and it is concluded that the use of pack stock has the potential to influence the following: (1) water nutrient dynamics, sedimentation, temperature, and microbial pathogen content; (2) soil chemistry, nutrient cycling, soil compaction and hydrology; (3) plant individuals, populations and community dynamics, non-native invasive species, and encroachment of woody species; and (4) wildlife individuals, populations and communities. It is considered from currently available information that management objectives of pack stock should include the following: minimise bare ground, maximise plant cover, maintain species composition of native plants, minimise trampling, especially on wet soils and stream banks, and minimise direct urination and defecation by pack stock into water. However, incomplete documentation of patterns of pack stock use and limited past research limits current understanding of the effects of pack stock, especially their effects on water, soils and wildlife. To improve management of pack stock in this region, research is needed on linking measurable monitoring variables (e.g. plant cover) with environmental relevancy (e.g. soil erosion processes, wildlife habitat use), and identifying specific environmental thresholds of degradation along gradients of pack stock use in Sierra Nevada meadows.

Rangeland Journal

Habitat selection and management of the Hawaiian crow

The abundance and range of the Hawaiian crow, or alala, ( Corvus hawaiiensis ) have decreased drastically since the 1890's. Fewer than 10 breeding pairs remained in the wild in 1985. A sample of 82 nests during 1970-82 were used to determine habitat associations. Two hundred firty-nine alala observations were used to estimate densities occurring in different vegetation types in 1978. Compared to available habitat, more nests and higher bird densities during the breeding season occurred in areas where: (1) canopy cover was > 60%; (2) koa ( Acacia koa ) and ohia ( Metrosideros polymorpha ) were dominant species in the crown layer; (3) native plants constituted > 75% of the understory cover; and (4) the elevation was 1,100-1,500 m. Compared to breeding habitat, nonbreeding habitat tended to lie at lower elevations and in wetter forests having the crown layer dominated by ohia but lacking koa. Habitat loss is a major factor underlying the decline of this species although predation on fledgings, avian disease, and shooting also have reduced the population. Remaining key habitat areas have little or no legal protection through zoning and land ownership. Preserves should be established to encompass the location of existing pairs and to assure the provision of optimum breeding habitat and suitable nonbreeding habitat.

Hawaii

Introduced species: A significant component of human-caused global change

Biological invasions are a widespread and significant component of human-caused global environmental change. The extent of invasions of oceanic islands, and their consequences for native biological diversity, have long been recognized. However, invasions of continental regions also are substantial. For example, more than 2,000 species of alien plants are established in the continental United States. These invasions represent a human-caused breakdown of the regional distinctiveness of Earth's flora and fauna—a substantial global change in and of itself. Moreover, there are well- documented examples of invading species that degrade human health and wealth, alter the structure and functioning of otherwise undisturbed ecosystems, and/or threaten native biological diversity. Invasions also interact synergistically with other components of global change. notably land use change. People and institutions working to understand, prevent, and control invasions are carrying out some of the most important—and potentially most effective—work on global environmental change.

New Zealand Journal of Ecology

Monitoring restoration impacts to endemic plant communities in soil inclusions of arid environments

Soil inclusions are small patches of soil with different properties than the surrounding, dominant soil. In arid areas of western North America, soil inclusions called slickspot soils are saltier than adjacent soil and support different types of native vegetation. Traditional sagebrush restoration efforts, such as using drills to plant seeds or herbicides to control invasive vegetation, may damage sensitive slickspot soil and supporting vegetation. USGS scientists David Pyke and Scott Shaff and collaborators monitored slickspot size and cover of endangered slickspot peppergrass for two years to see if they were affected by the application of the herbicide glyphosate or by a minimum-till drill in the Snake River Plain, ID. The researchers examined the use of aerial photographs versus on-the-ground measurements and concluded that slickspot sizes were not affected by these treatments. Remote sensing using aerial photographs proved a useful method for mapping slickspot soils.

International Journal of Agriculture & Biology

Microbial endophytes and compost improve plant growth in two contrasting types of hard rock mining waste

The re-vegetation of mining wastes with native plants is a comparatively low-cost solution for mine reclamation. However, re-vegetation fails when extreme pH values, low organic matter, or high concentrations of phytotoxic elements inhibit plant establishment and growth. Our aim was to determine whether the combined addition of municipal waste compost and diazotrophic endophytes (i.e., microorganisms that fix atmospheric N 2 and live within plants) could improve plant growth, organic matter accumulation, and phytostabilization of trace element contaminants in two types of hard rock mine waste. We grew a widespread native perennial grass, Bouteloua curtipendula , for one month in alkaline waste rock (porphyry copper mine) and tailings (Ag-Pb-Au mine, amended with dolomite) sourced from southeastern Arizona, United States. B. curtipendula tolerated elevated concentrations of multiple phytotoxic trace elements in the tailings (Mn, Pb, Zn), stabilizing them in roots without foliar translocation. Adding compost and endophyte seed coats improved plant growth, microbial biomass, and organic matter accumulation despite stark differences in the geochemical and physical characteristics of the mining wastes. The widespread grass B. curtipendula is a potential candidate for re-vegetating mine wastes when seeded with soil additives to increase pH and with microbial and organic amendments to increase plant growth.

International Journal of Phytoremediation

Synthesis of common management concerns associated with dam removal

Managers make decisions regarding if and how to remove dams in spite of uncertainty surrounding physical and ecological responses, and stakeholders often raise concerns about certain negative effects, regardless of whether or not these concerns are warranted at a particular site. We used a dam-removal science database supplemented with other information sources to explore seven frequently-raised concerns, herein Common Management Concerns (CMCs). We investigate the occurrence of these concerns and the contributing biophysical controls. The CMCs addressed are: degree and rate of reservoir sediment erosion, excessive channel incision upstream of reservoirs, downstream sediment aggradation, elevated downstream turbidity, drawdown impacts on local water infrastructure, colonization of reservoir sediments by non-native plants, and expansion of invasive fish. Biophysical controls emerged for some of the concerns, providing managers with information to assess whether a given concern is likely to occur at a site. To fully assess CMC risk, managers should concurrently evaluate site conditions and identify the ecosystem or human uses that will be negatively affected if the biophysical phenomenon producing the CMC occurs. We show how many CMCs have one or more controls in common, facilitating the identification of multiple risks at a site, and demonstrate why CMC risks should be considered in the context of other factors like natural watershed variability and disturbance history.

Journal of the American Water Resources Associatio

Establishment of non-native plant species after wildfires: Effects of fuel treatments, abiotic and biotic factors, and post-fire grass seeding treatments

Establishment and spread of non-native species following wildfires can pose threats to long-term native plant recovery. Factors such as disturbance severity, resource availability, and propagule pressure may influence where non-native species establish in burned areas. In addition, pre- and post-fire management activities may influence the likelihood of non-native species establishment. In the present study we examine the establishment of non-native species after wildfires in relation to native species richness, fire severity, dominant native plant cover, resource availability, and pre- and post-fire management actions (fuel treatments and post-fire rehabilitation treatments). We used an information-theoretic approach to compare alternative hypotheses. We analysed post-fire effects at multiple scales at three wildfires in Colorado and New Mexico. For large and small spatial scales at all fires, fire severity was the most consistent predictor of non-native species cover. Non-native species cover was also correlated with high native species richness, low native dominant species cover, and high seeded grass cover. There was a positive, but non-significant, association of non-native species with fuel-treated areas at one wildfire. While there may be some potential for fuels treatments to promote non-native species establishment, wildfire and post-fire seeding treatments seem to have a larger impact on non-native species. ?? IAWF 2006.

Colorado, New Mexico

Fire and vegetation history on Santa Rosa Island, Channel Islands, and long-term environmental change in southern California

The long-term history of vegetation and fire was investigated at two locations – Soledad Pond (275 m; from ca. 12 000 cal. a BP) and Abalone Rocks Marsh (0 m; from ca. 7000 cal. a BP) – on Santa Rosa Island, situated off the coast of southern California. A coastal conifer forest covered highlands of Santa Rosa during the last glacial, but by ca. 11 800 cal. a BP Pinus stands, coastal sage scrub and grassland replaced the forest as the climate warmed. The early Holocene became increasingly drier, particularly after ca. 9150 cal. a BP, as the pond dried frequently, and coastal sage scrub covered the nearby hillslopes. By ca. 6900 cal. a BP grasslands recovered at both sites. Pollen of wetland plants became prominent at Soledad Pond after ca. 4500 cal. a BP, and at Abalone Rocks Marsh after ca. 3465 cal. a BP. Diatoms suggest freshening of the Abalone Rocks Marsh somewhat later, probably by additional runoff from the highlands. Introduction of non-native species by ranchers occurred subsequent to AD 1850. Charcoal influx is high early in the record, but declines during the early Holocene when minimal biomass suggests extended drought. A general increase occurs after ca. 7000 cal. a BP, and especially after ca. 4500 cal. a BP. The Holocene pattern closely resembles population levels constructed from the archaeological record, and suggests a potential influence by humans on the fire regime of the islands, particularly during the late Holocene.

California

Habitat manipulation influences northern bobwhite resource selection on a reclaimed surface mine

More than 600,000&thinsp;ha of mine land have been reclaimed in the eastern United States, providing large contiguous tracts of early successional vegetation that can be managed for northern bobwhite ( Colinus virginianus ). However, habitat quality on reclaimed mine land can be limited by extensive coverage of non-native invasive species, which are commonly planted during reclamation. We used discrete-choice analysis to investigate bobwhite resource selection throughout the year on Peabody Wildlife Management Area, a 3,330-ha reclaimed surface mine in western Kentucky. We used a treatment-control design to study resource selection at 2 spatial scales to identify important aspects of mine land vegetation and whether resource selection differed between areas with habitat management (i.e., burning, disking, herbicide; treatment) and unmanaged units (control). Our objectives were to estimate bobwhite resource selection on reclaimed mine land and to estimate the influence of habitat management practices on resource selection. We used locations from 283 individuals during the breeding season (1 Apr&ndash;30 Sep) and 136 coveys during the non-breeding season (1 Oct&ndash;Mar 31) from August 2009 to March 2014. Individuals were located closer to shrub cover than would be expected at random throughout the year. During the breeding season, individuals on treatment units used areas with smaller contagion index values (i.e., greater interspersion) compared with individuals on control units. During the non-breeding season, birds selected areas with greater shrub-open edge density compared with random. At the microhabitat scale, individuals selected areas with increased visual obstruction >1&thinsp;m aboveground. During the breeding season, birds were closer to disked areas (linear and non-linear) than would be expected at random. Individuals selected non-linear disked areas during winter but did not select linear disked areas (firebreaks) because they were planted to winter wheat each fall and lacked cover during the non-breeding season. Individuals also selected areas treated with herbicide to control sericea lespedeza ( Lespedeza cuneata ) throughout the year. During the breeding season, bobwhites avoided areas burned during the previous dormant season. Habitat quality of reclaimed mine lands may be limited by a lack of shrub cover and extensive coverage of non-native herbaceous vegetation. Managers aiming to increase bobwhite abundance should focus on increasing interspersion of shrub cover, with no area >100&thinsp;m from shrub cover. We suggest disking and herbicide application to control invasive species and improve the structure and composition of vegetation for bobwhites.

Kentucky

A test of vegetation-related indicators of wetland quality in the prairie pothole region

This study was part of an effort by the U.S. Environmental Protection Agency to quantitatively assess the environmental quality or 'health' of wetland resources on regional and national scales. During a two-year pilot study, we tested selected indicators of wetland quality in the U.S. portion of the prairie pothole region (PPR). We assumed that the amount of cropland versus non-cropland (mostly grassland) in the plots containing these basins was a proxy for their quality. We then tested indicators by their ability to discriminate between wetlands at the extremes of that proxy. Amounts of standing dead vegetation were greater in zones of greater water permanence. Depth of litter was greater in zones of greater water permanence and in zones of basins in poor-quality watersheds. Amounts of unvegetated bottom were greater in basins in poor-quality watersheds; lesser amounts occurred in all wetlands during a wetter year. Greater amounts of open water occurred during a wetter year and in zones of greater water permanence. When unadjusted for areas (ha) of communities, plant taxon richness was higher in wet-meadow and shallow-marsh zones in good-quality watersheds than in similar zones in poor-quality watersheds. Wet-meadow zones in good-quality watersheds had greater numbers of native perennials than those in poor-quality watersheds. This relation held when we eliminated all communities in good-quality watersheds larger than the largest communities in poor-quality watersheds from the data set. We conclude that although amounts of unvegetated bottom and plant taxon richness in wet-meadow zones were useful indicators of wetland quality during our study, the search for additional such indicators should continue. The value of these indicators may change with the notoriously unstable hydrological conditions in the PPR. Most valuable would be indicators that could be photographed or otherwise remotely sensed and would remain relatively stable under various hydrological conditions. An ideal set of indicators could detect the absence of stressors, as well as the presence of structures or functions, of known value to major groups of organisms.

Journal of Aquatic Ecosystem Health

Ramet spacing of Elymus lanceolatus (thickspike wheatgrass) in response to neighbour density

Many plants exploit patchy resources through clonal foraging. Plants established in field plots were used to determine if Elymus lanceolatus ssp. lanceolatus (Scribner et J.G. Smith) Gould (thickspike wheatgrass) showed a clonal foraging response to neighbour densities, as it had previously shown to patchy soil nutrients. Neighbours consisted of the rhizomatous E. lanceolatus ssp. lanceolatus and the bunchgrass Elymus lanceolatus ssp. wawawaiensis (Scribner et Gould) J.R. Carlson et D.R. Dewey (proposed name), which are both native to the semiarid western U.S.A., and their ratios as well as total densities varied. Rather than an increase in spacing of exploratory ramets at high densities, as expected with clonal foraging, there was a decrease in spacing in both years of the experiment. Fewer target plants produced exploratory ramets at higher densities only in the second year. These reductions in exploratory clonal growth at higher neighbour densities, which were opposite to E . lanceolatus ssp. lanceolatus ' response to low-resource patches, occurred perhaps because soil resource levels were too low overall to support rhizome production, and this condition was more pronounced in the second year. Physical resistance from neighbour roots perhaps also reduced rhizome production. However, rhizome growth may not be beneficial in such cases, and plants may be adapted to produce exploratory rhizomes only when some high-resource patches are encountered by the clone.

Canadian Journal of Botany