Search USGSSearch

SEARCH · Search USGS

Results for “Ecological Modeling”

Search indexed USGS publications on groundwater, aquifers, geologic maps, mineral resources and earthquakes. Explore source records by subject and place.

Quote a phrase for an exact phrase match. Source license links do not imply unrestricted reuse.

At least 487 records · Page 27Linked to original sources

Integration of environmental simulation models with satellite remote sensing and geographic information systems technologies: case studies

Environmental modelers are testing and evaluating a prototype land cover characteristics database for the conterminous United States developed by the EROS Data Center of the U.S. Geological Survey and the University of Nebraska Center for Advanced Land Management Information Technologies. This database was developed from multi temporal, 1-kilometer advanced very high resolution radiometer (AVHRR) data for 1990 and various ancillary data sets such as elevation, ecological regions, and selected climatic normals. Several case studies using this database were analyzed to illustrate the integration of satellite remote sensing and geographic information systems technologies with land-atmosphere interactions models at a variety of spatial and temporal scales. The case studies are representative of contemporary environmental simulation modeling at local to regional levels in global change research, land and water resource management, and environmental simulation modeling at local to regional levels in global change research, land and water resource management and environmental risk assessment. The case studies feature land surface parameterizations for atmospheric mesoscale and global climate models; biogenic-hydrocarbons emissions models; distributed parameter watershed and other hydrological models; and various ecological models such as ecosystem, dynamics, biogeochemical cycles, ecotone variability, and equilibrium vegetation models. The case studies demonstrate the important of multi temporal AVHRR data to develop to develop and maintain a flexible, near-realtime land cover characteristics database. Moreover, such a flexible database is needed to derive various vegetation classification schemes, to aggregate data for nested models, to develop remote sensing algorithms, and to provide data on dynamic landscape characteristics. The case studies illustrate how such a database supports research on spatial heterogeneity, land use, sensitivity analysis, and scaling issues involving regional extrapolations and parameterizations of dynamic land processes within simulation models.

Pecora 12 Symposium

Modelling the distribution of chickens, ducks, and geese in China

Global concerns over the emergence of zoonotic pandemics emphasize the need for high-resolution population distribution mapping and spatial modelling. Ongoing efforts to model disease risk in China have been hindered by a lack of available species level distribution maps for poultry. The goal of this study was to develop 1 km resolution population density models for China's chickens, ducks, and geese. We used an information theoretic approach to predict poultry densities based on statistical relationships between poultry census data and high-resolution agro-ecological predictor variables. Model predictions were validated by comparing goodness of fit measures (root mean square error and correlation coefficient) for observed and predicted values for 1/4 of the sample data which were not used for model training. Final output included mean and coefficient of variation maps for each species. We tested the quality of models produced using three predictor datasets and 4 regional stratification methods. For predictor variables, a combination of traditional predictors for livestock mapping and land use predictors produced the best goodness of fit scores. Comparison of regional stratifications indicated that for chickens and ducks, a stratification based on livestock production systems produced the best results; for geese, an agro-ecological stratification produced best results. However, for all species, each method of regional stratification produced significantly better goodness of fit scores than the global model. Here we provide descriptive methods, analytical comparisons, and model output for China's first high resolution, species level poultry distribution maps. Output will be made available to the scientific and public community for use in a wide range of applications from epidemiological studies to livestock policy and management initiatives.

Agriculture, Ecosystems and Environment

Improving the effectiveness of ecological site descriptions: General state-and-transition models and the Ecosystem Dynamics Interpretive Tool (EDIT)

State-and-transition models (STMs) are useful tools for management, but they can be difficult to use and have limited content. STMs created for groups of related ecological sites could simplify and improve their utility. The amount of information linked to models can be increased using tables that communicate management interpretations and important within-group variability. We created a new web-based information system (the Ecosystem Dynamics Interpretive Tool) to house STMs, associated tabular information, and other ecological site data and descriptors. Fewer, more informative, better organized, and easily accessible STMs should increase the accessibility of science information.

Rangelands

Applications of genetic data to improve management and conservation of river fishes and their habitats

Environmental variation and landscape features affect ecological processes in fluvial systems; however, assessing effects at management-relevant temporal and spatial scales is challenging. Genetic data can be used with landscape models and traditional ecological assessment data to identify biodiversity hotspots, predict ecosystem responses to anthropogenic effects, and detect impairments to underlying processes. We show that by combining taxonomic, demographic, and genetic data of species in complex riverscapes, managers can better understand the spatial and temporal scales over which environmental processes and disturbance influence biodiversity. We describe how population genetic models using empirical or simulated genetic data quantify effects of environmental processes affecting species diversity and distribution. Our summary shows that aquatic assessment initiatives that use standardized data sets to direct management actions can benefit from integration of genetic data to improve the predictability of disturbance–response relationships of river fishes and their habitats over a broad range of spatial and temporal scales.

Fisheries

Spatial capture-recapture models for jointly estimating population density and landscape connectivity

Population size and landscape connectivity are key determinants of population viability, yet no methods exist for simultaneously estimating density and connectivity parameters. Recently developed spatial capture–recapture (SCR) models provide a framework for estimating density of animal populations but thus far have not been used to study connectivity. Rather, all applications of SCR models have used encounter probability models based on the Euclidean distance between traps and animal activity centers, which implies that home ranges are stationary, symmetric, and unaffected by landscape structure. In this paper we devise encounter probability models based on “ecological distance,” i.e., the least-cost path between traps and activity centers, which is a function of both Euclidean distance and animal movement behavior in resistant landscapes. We integrate least-cost path models into a likelihood-based estimation scheme for spatial capture–recapture models in order to estimate population density and parameters of the least-cost encounter probability model. Therefore, it is possible to make explicit inferences about animal density, distribution, and landscape connectivity as it relates to animal movement from standard capture–recapture data. Furthermore, a simulation study demonstrated that ignoring landscape connectivity can result in negatively biased density estimators under the naive SCR model.

Ecology

Infectious hematopoietic necrosis virus virological and genetic surveillance 2000–2012

Surveillance records of the acute RNA pathogen of Pacific salmonid fish infectious hematopoietic necrosis virus are combined for the first time to enable landscape-level ecological analyses and modeling. The study area is the freshwater ecosystems of the large Columbia River watershed in the U.S. states of Washington, Oregon, and Idaho, as well as coastal rivers in Washington and Oregon. The study period is 2000–2012, and records were contributed by all five resource management agencies that operate conservation hatcheries in the study area. Additional records from wild fish were collected from the National Wild Fish Health Survey, operated by the U.S. Fish and Wildlife Survey. After curation and normalization, the data set consists of 6766 records, representing 1146 sample sites and 15 different fish hosts. The virus was found in an average of 12.4% of records, and of these 66.2% also have viral genetic analysis available. This data set is used to conduct univariate ecological and epidemiological analyses and develop a novel hierarchical landscape transmission model for an aquatic pathogen.

Ecology

Behavioral differences following ingestion of large meals and consequences for management of a harmful invasive snake: A field experiment

Many snakes are uniquely adapted to ingest large prey at infrequent intervals. Digestion of large prey is metabolically and aerobically costly, and large prey boluses can impair snake locomotion, increasing vulnerability to predation. Cessation of foraging and use of refugia with microclimates facilitating digestion are expected to be strategies employed by free‐ranging snakes to cope with the demands of digestion while minimizing risk of predation. However, empirical observations of such submergent behavior from field experiments are limited. The brown treesnake (Serpentes: Colubridae: Boiga irregularis ) is a nocturnal, arboreal, colubrid snake that was accidentally introduced to the island of Guam, with ecologically and economically costly consequences. Because tools for brown treesnake damage prevention generally rely on snakes being visible or responding to lures or baits while foraging, cessation of foraging activities after feeding would complicate management. We sought to characterize differences in brown treesnake activity, movement, habitat use, and detectability following feeding of large meals (rodents 33% of the snake's unfed body mass) via radio telemetry, trapping, and visual surveys. Compared to unfed snakes, snakes in the feeding treatment group showed drastic decreases in hourly and nightly activity rates, differences in refuge height and microhabitat type, and a marked decrease in detectability by trapping and visual surveys. Depression of activity lasted approximately 5–7 days, a period that corresponds to previous studies of brown treesnake digestion and cycles of detectability. Our results indicate that management strategies for invasive brown treesnakes need to account for cycles of unavailability and underscore the importance of preventing spread of brown treesnakes to new environments where large prey are abundant and periods of cryptic behavior are likely to be frequent. Characterization of postfeeding behavior changes provides a richer understanding of snake ecology and foraging models for species that consume large prey.

Ecology and Evolution

The Hyper-Envelope Modeling Interface (HEMI): A Novel Approach Illustrated Through Predicting Tamarisk ( Tamarix spp.) Habitat in the Western USA

Habitat suitability maps are commonly created by modeling a species’ environmental niche from occurrences and environmental characteristics. Here, we introduce the hyper-envelope modeling interface (HEMI), providing a new method for creating habitat suitability models using Bezier surfaces to model a species niche in environmental space. HEMI allows modeled surfaces to be visualized and edited in environmental space based on expert knowledge and does not require absence points for model development. The modeled surfaces require relatively few parameters compared to similar modeling approaches and may produce models that better match ecological niche theory. As a case study, we modeled the invasive species tamarisk ( Tamarix spp.) in the western USA. We compare results from HEMI with those from existing similar modeling approaches (including BioClim, BioMapper, and Maxent). We used synthetic surfaces to create visualizations of the various models in environmental space and used modified area under the curve (AUC) statistic and akaike information criterion (AIC) as measures of model performance. We show that HEMI produced slightly better AUC values, except for Maxent and better AIC values overall. HEMI created a model with only ten parameters while Maxent produced a model with over 100 and BioClim used only eight. Additionally, HEMI allowed visualization and editing of the model in environmental space to develop alternative potential habitat scenarios. The use of Bezier surfaces can provide simple models that match our expectations of biological niche models and, at least in some cases, out-perform more complex approaches.

Environmental Management

Simulation modeling of complex climate, wildfire, and vegetation dynamics to address wicked problems in land management

Complex, reciprocal interactions among climate, disturbance, and vegetation dramatically alter spatial landscape patterns and influence ecosystem dynamics. As climate and disturbance regimes shift, historical analogs and past empirical studies may not be entirely appropriate as templates for future management. The need for a better understanding of the potential impacts of climate changes on ecosystems is reaching a new level of urgency, especially in highly perturbed or vulnerable ecological systems. Simulation models are extremely useful tools for guiding management decisions in an era of rapid change, thus providing potential solutions for wicked problems in land management—those that are difficult to solve and inherently resistant to easily definable solutions. We identify three experimental approaches for landscape modeling that address management challenges in the context of uncertain climate futures and complex ecological interactions: (1) an historical comparative approach, (2) a future comparative approach, and (3) threshold detection. We provide examples of each approach from previously published studies of simulated climate, disturbance, and landscape dynamics in forested landscapes of the western United States, modeled with the FireBGCv2 ecosystem process model. Cumulatively, model outcomes indicate that typical land management strategies will likely not be sufficient to counteract the impacts of rapid climate change and altered disturbance regimes that threaten the stability of ecosystems. Without implementation of new, adaptive management strategies, future landscapes are very likely to be different than historical or contemporary ones, with significant and sometimes persistent changes triggered by interactions of climate and wildfire.

Montana, New Mexico, Wyoming

Climate change and disturbance interactions: Workshop on climate change and disturbance interactions in western North America, Tucson, Ariz., 12-15 February 2007

Warming temperatures across western North America, coupled with increased drought, are expected to exacerbate disturbance regimes, particularly wildfires, insect outbreaks, and invasions of exotic species. Many ecologists and resource managers expect ecosystems to change more rapidly from disturbance effects than from the effects of a changing climate by itself. A particular challenge is to understand the interactions among disturbance regimes; for example, how will massive outbreaks of bark beetles, which kill drought-stressed trees by feeding on cambial tissues, increase the potential for large severe wildfires in a warming climate? Researchers in climatology, ecosystem science, fire and insect ecology, and landscape modeling from across western North America convened in Tucson, Ariz., for a 2 and a half day intensive workshop to identify new research directions in climate change and disturbance ecology. Four work groups focused on different aspects of the response of disturbance regimes to climate change: (1) extreme events and climatic variability (2) the effects of changing disturbance regimes on ecosystems, (3) disturbance interactions and cumulative effects, and (4) developing new landscape disturbance models. The workshop was structured with the analytic hierarchy process, a decision support method for achieving consensus from diverse groups of experts without sacrificing individual contributions.

Eos, Earth and Space Science News

Evaluating the impact of watershed development and climate change on stream ecosystems: A Bayesian network modeling approach

A continuous-variable Bayesian network (cBN) model is used to link watershed development and climate change to stream ecosystem indicators. A graphical model, reflecting our understanding of the connections between climate change, weather condition, loss of natural land cover, stream flow characteristics , and stream ecosystem indicators is used as the basis for selecting flow metrics for predicting macroinvertebrate-based indicators. Selected flow metrics were then linked to variables representing watershed development and climate change. We fit the model to data from two river basins in southeast US and the resulting model was used to simulate future stream ecological conditions using projected future climate and development scenarios. The three climate models predicted varying ecological condition trajectories, but similar worst-case ecological conditions. The established modeling approach couples mechanistic understanding with field data to develop predictions of management-relevant variables across a heterogeneous landscape. We discussed the transferability of the modeling approach.

North Carolina, South Carolina, Virginia

Users' guide to system dynamics model describing Coho salmon survival in Olema Creek, Point Reyes National Seashore, Marin County, California

The system dynamics model described in this report is the result of a collaboration between U.S. Geological Survey (USGS) scientists and National Park Service (NPS) San Francisco Bay Area Network (SFAN) staff, whose goal was to develop a methodology to integrate inventory and monitoring data to better understand ecosystem dynamics and trends using salmon in Olema Creek, Marin County, California, as an example case. The SFAN began monitoring multiple life stages of coho salmon (Oncorhynchus kisutch) in Olema Creek during 2003 (Carlisle and others, 2013), building on previous monitoring of spawning fish and redds. They initiated water-quality and habitat monitoring, and had access to flow and weather data from other sources. This system dynamics model of the freshwater portion of the coho salmon life cycle in Olema Creek integrated 8 years of existing monitoring data, literature values, and expert opinion to investigate potential factors limiting survival and production, identify data gaps, and improve monitoring and restoration prescriptions. A system dynamics model is particularly effective when (1) data are insufficient in time series length and/or measured parameters for a statistical or mechanistic model, and (2) the model must be easily accessible by users who are not modelers. These characteristics helped us meet the following overarching goals for this model: Summarize and synthesize NPS monitoring data with data and information from other sources to describe factors and processes affecting freshwater survival of coho salmon in Olema Creek. Provide a model that can be easily manipulated to experiment with alternative values of model parameters and novel scenarios of environmental drivers. Although the model describes the ecological dynamics of Olema Creek, these dynamics are structurally similar to numerous other coastal streams along the California coast that also contain anadromous fish populations. The model developed for Olema can be used, at least as a starting point, for other watersheds. This report describes each of the model elements with sufficient detail to guide the primary target audience, the NPS resource specialist, to run the model, interpret the results, change the input data to explore hypotheses, and ultimately modify and improve the model. Running the model and interpreting the results does not require modeling expertise on the part of the user. Additional companion publications will highlight other aspects of the model, such as its development, the rationale behind the methodological approach, scenario testing, and discussions of its use. System dynamics models consist of three basic elements: stocks , flows , and converters . Stocks are measurable quantities that can change over time, such as animal populations. Flows are any processes or conditions that change the quantity in a stock over time (Ford, 1999), are expressed in the model as a rate of change, and are diagrammed as arrows to or from stocks. Converters are processes or conditions that change the rate of flows. A converter is connected to a flow with an arrow indicating that it alters the rate of change. Anything that influences the rate of change (such as different environmental conditions, other external factors, or feedbacks from other stocks or flows) is modeled as a converter. For example, the number of fish in a population is appropriately modeled as a stock. Mortality is modeled as a flow because it is a rate of change over time used to determine the number of fish in the population. The density-dependent effect on mortality is modeled as a converter because it influences the rate of morality. Together, the flow and converter change the number, or stock, of juvenile coho. The instructions embedded in the stocks, flows, converters, and the sequence in which they are linked are processed by the simulation software with each completed sequence composing a model run. At each modeled time step within the model run, the stock counts will go up, down, or stay the same based on the modeled flows and the influence of converters on those flows. The model includes a user-friendly interface to change model parameters, which allows park staff and others to conduct sensitivity analyses, incorporate future knowledge, and implement scenarios for various future conditions. The model structure incorporates place holders for relationships that we hypothesize are significant but data are currently lacking. Future climate scenarios project stream temperatures higher than any that have ever been recorded at Olema Creek. Exploring climate change impacts on coho survival is a high priority for park staff, therefore the model provides the user with the option to experiment with hypothesized effects and to incorporate effects based on future observations.

California

Mapping predicted ecological states at landscape scales using remote sensing data and machine learning

Dryland ecosystems, covering 45% of the Earth's land and supporting over one-third of the global population, face significant threats from land degradation and ecological state change. Managing these ecosystems is complex, and science-based frameworks like Ecological Site Descriptions and state-and-transition models are essential tools for guiding decisions to support ecological health while maintaining stakeholder values such as grazing, wildlife, and recreation. However, alignment of these frameworks with smaller scale soil survey maps limits their applicability to broader ecological processes. Here, we extend these frameworks to larger landscapes with a machine learning approach that integrates large-scale, high-resolution vegetation data with identified ecological states from a data-driven state-and-transition model developed for a landscape-scale Ecological Site Group. A “global” model, which used combined inputs from multiple remotely sensed datasets, outperformed individual dataset models based on evaluation with independent data. Ecological state maps generated through this approach broaden the utility of state-and-transition models across Ecological Site Groups, providing a more spatially robust tool for land management at watershed and larger landscape scales. These methods, and the associated ecological state maps, can help meet critical needs for improved land condition assessments that support development of resource management plans and help identify priority areas for restoration and conservation.

Arizona, Colorado, New Mexico, Utah, Wyoming

Applying local and global sensitivity analysis to inform bigheaded carp management

Natural resource managers commonly use population-level models to aid in understanding the status of target populations or the potential implications of management actions. Sensitivity analyses, specifically, local sensitivity analysis (LSA) and global sensitivity analysis (GSA), exist as tools to improve understanding of these models, the importance of specific parameters to model outcomes, and the assumptions inherent to population models. However, applied ecology and closely related fields such as fisheries and wildlife management rarely use LSA or GSA. Here, we apply LSA and GSA to the Spatially Explicit Invasive Carp Population model, a population-level model used to simulate and compare management actions on the invasive silver carp Hypophthalmichthys molitrix population in the Illinois River. The spatial structure within the model arises from the navigation “pools” that exist between navigation lock-and-dam in the river and are also the focus of fisheries management in the river. We completed LSA and GSA for two reasons: (1) to compare the importance of model input parameters on simulated model populations and (2) to demonstrate the use of LSA and GSA for population-level models. In our system, the most important sensitivity values generally were detected in Alton Pool. This pool is the farthest downstream and includes a direct connection with the Mississippi River. These results indicate that the population dynamics in Alton Pool are an important driver of the population dynamics throughout the Illinois River. Management outcomes in this system will, therefore, be strongly dependent on the dynamics of Alton Pool.

Illinois

Combining resilience and resistance with threat-based approaches for prioritizing management actions in sagebrush ecosystems

The sagebrush biome is a dryland region in the western United States experiencing rapid transformations to novel ecological states. Threat-based approaches for managing anthropogenic and ecosystem threats have recently become prominent, but successfully mitigating threats depends on the ecological resilience of ecosystems. We used a spatially explicit approach for prioritizing management actions that combined a threat-based model with models of resilience to disturbance and resistance to annual grass invasion. The threat-based model assessed geographic patterns in sagebrush ecological integrity (SEI) to identify core sagebrush, growth opportunity, and other rangeland areas. The resilience and resistance model identified ecologically relevant climate and soil water availability indicators from process-based ecohydrological models. The SEI areas and resilience and resistance indicators were consistent—the resilience and resistance indicators showed generally positive relationships with the SEI areas. They also were complementary—SEI areas provided information on intact sagebrush areas and threats, while resilience and resistance provided information on responses to disturbances and management actions. The SEI index and resilience and resistance indicators provide the basis for prioritizing conservation and restoration actions and determining appropriate strategies. The difficulty and time required to conserve or restore SEI areas increase as threats increases and resilience and resistance decrease.

Conservation Science and Practice

Evaluating the sagebrush conservation design through the lens of a sagebrush indicator species

Sagebrush ecosystems support a suite of unique species such as the emblematic greater sage-grouse ( Centrocercus urophasianus ; sage-grouse) but are under increasing pressure from anthropogenic stressors such as annual grass invasion, conifer encroachment, altered wildfire regimes, and land use change. We examined the ability of an ecosystem-based framework for sagebrush conservation, the sagebrush conservation design (SCD) strategy, and the associated model of sagebrush ecological integrity (SEI), to identify and rank priority habitats for sage-grouse, a sagebrush indicator species. We compared sage-grouse population trends from 1996–2021 across the three ranked SEI categories. We then modeled those trends directly as a function of the same landcover predictors underlying SEI, used the median trend estimates to recategorize the sage-grouse’s range, and used spatial correlation methods to compare our sage-grouse performance categories with those of SEI. Finally, we compared the sage-grouse condition categories, predicted by our landcover-based model, to empirical trends derived from population count data. We found that the SCD and SEI were effective tools for identifying and ranking priority habitats for sage-grouse. Population trends were stable in the core areas identified by SEI but declining in the lower (i.e., growth and other) condition categories. As a result, core areas encompassed an increasingly larger share of the total sage-grouse population in a disproportionately smaller area. Our model supports the general functional relationships between landcover and sage-grouse performance suggested by SEI. We found strong spatial congruence between our categories of predicted sage-grouse population performance, the condition categories of SEI, and empirical trends derived from population count data. Our analysis demonstrates that proactive ecosystem-based approaches to the conservation of the sagebrush biome can help optimize the return on limited conservation resources and benefits for sagebrush obligate species and help reduce some of the real and perceived conflicts inherent in single-species management.

Arizona, California, Colorado, Idaho, New Mexico,

Identifying western yellow-billed cuckoo breeding habitat with a dual modelling approach

The western population of the yellow-billed cuckoo ( Coccyzus americanus ) was recently listed as threatened under the federal Endangered Species Act. Yellow-billed cuckoo conservation efforts require the identification of features and area requirements associated with high quality, riparian forest habitat at spatial scales that range from nest microhabitat to landscape, as well as lower-suitability areas that can be enhanced or restored. Spatially explicit models inform conservation efforts by increasing ecological understanding of a target species, especially at landscape scales. Previous yellow-billed cuckoo modelling efforts derived plant-community maps from aerial photography, an expensive and oftentimes inconsistent approach. Satellite models can remotely map vegetation features (e.g., vegetation density, heterogeneity in vegetation density or structure) across large areas with near perfect repeatability, but they usually cannot identify plant communities. We used aerial photos and satellite imagery, and a hierarchical spatial scale approach, to identify yellow-billed cuckoo breeding habitat along the Lower Colorado River and its tributaries. Aerial-photo and satellite models identified several key features associated with yellow-billed cuckoo breeding locations: (1) a 4.5 ha core area of dense cottonwood-willow vegetation, (2) a large native, heterogeneously dense forest (72 ha) around the core area, and (3) moderately rough topography. The odds of yellow-billed cuckoo occurrence decreased rapidly as the amount of tamarisk cover increased or when cottonwood-willow vegetation was limited. We achieved model accuracies of 75–80% in the project area the following year after updating the imagery and location data. The two model types had very similar probability maps, largely predicting the same areas as high quality habitat. While each model provided unique information, a dual-modelling approach provided a more complete picture of yellow-billed cuckoo habitat requirements and will be useful for management and conservation activities.

Ecological Modelling

Normality of raw data in general linear models: The most widespread myth in statistics

In years of statistical consulting for ecologists and wildlife biologists, by far the most common misconception we have come across has been the one about normality in general linear models. These comprise a very large part of the statistical models used in ecology and include t tests, simple and multiple linear regression, polynomial regression, and analysis of variance (ANOVA) and covariance (ANCOVA). There is a widely held belief that the normality assumption pertains to the raw data rather than to the model residuals. We suspect that this error may also occur in countless published studies, whenever the normality assumption is tested prior to analysis. This may lead to the use of nonparametric alternatives (if there are any), when parametric tests would indeed be appropriate, or to use of transformations of raw data, which may introduce hidden assumptions such as multiplicative effects on the natural scale in the case of log-transformed data. Our aim here is to dispel this myth. We very briefly describe relevant theory for two cases of general linear models to show that the residuals need to be normally distributed if tests requiring normality are to be used, such as t and F tests. We then give two examples demonstrating that the distribution of the response variable may be nonnormal, and yet the residuals are well behaved. We do not go into the issue of how to test normality; instead we display the distributions of response variables and residuals graphically.

Bulletin of the Ecological Society of America