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At least 451 records · Page 25Linked to original sources

USGS1997

Excavation of red squirrel middens by grizzly bears in the whitebark pine zone

Whitebark pine seeds Pinus albicaulis are an important food of grizzly Ursus arctos horribilis bears wherever whitebark pine is abundant in the contiguous United States of America; availability of seeds affects the distribution of bears, and the level of conflict between bears and humans. Almost all of the seeds consumed by bears are excavated from middens where red squirrels Tamiasciurus hudsonicus have cached whitebark pine cones. Relationships among the occupancy of middens by squirrels, the excavation of middens by bears, and site features were investigated in this study. Data were collected from radio-marked bears and from middens located from line transects on two study sites in the Yellowstone ecosystem. Densities of active middens were positively related to lodgepole pine Pinus contorta basal area and negatively related to steepness of slope. The probability that a midden was occupied by a squirrel (i.e. active) was positively related to lodgepole pine basal area in the surrounding stand, size of the midden and size of the whitebark pine cone crop, and negatively related to elevation and to bear excavation during the previous 2-12 months. The probability that a midden had been excavated by a bear during the previous 12 months was positively related to size of the midden, and to whitebark pine basal area and cone crop, and negatively related to nearness of roads and town sites. The influence of midden size on bear use was attributable to a positive relationship with the number of excavated cones. The positive association between bear excavations and whitebark pine basal area or cone crops was attributable to availability of pine seeds. Grizzly bears would benefit from the minimization of roads and other human facilities in the whitebark pine zone and from increases in the availability of whitebark pine seeds, potentially achieved by increasing the numbers of cone-producing whitebark pine trees, especially in lower elevations of the whitebark pine zone where red squirrels are more abundant.

Journal of Applied Ecology↗

USGS1992

Distribution of Yellowstone grizzly bears during the 1980s

Grizzly bear ( Ursus arctos horribilis ) females with young occupied a greater proportion of area east of Yellowstone National Park during 1980-1989 compared to 1973-1979. Occupancy by all bears and females with young was lower in all peripheral areas compared to the Park, but greater east and south compared to north and south of the Park. Observed changes reflect not only a static or slightly increasing population trend, but may also reflect biases in data collection. These biases are discussed and distributions of all observations and females with young are presented. Methodological problems implicit in analyzing changes in distribution of grizzly bears are also discussed.

Idaho, Montana, Wyoming↗

USGS1986

Monitoring grizzly bear population trends

A simple different equation model was developed to provide additional perspective on observed mortality and trend data on Yellowstone grizzly bears ( Ursus arctos horribilis ). Records of mortalities of adult females from 1959 to 1985 were utilized, in conjunction with data on females with cubs. The overall downward trend of observed numbers of females with cubs generally agrees with the model calculations but does not adequately reflect mortality from 1970 to 1974. The model may be useful in developing a composite index of population trend.

Idaho;Montana;Wyoming↗

USGS1997

Spring feeding on ungulate carcasses by grizzly bears in Yellowstone National Park

We studied the spring use of ungulate carcasses by grizzly bears ( Ursus arctos horribilis ) on ungulate winter ranges in Yellowstone National Park. We observed carcasses and bear tracks on survey routes that were travelled biweekly during spring of 1985-90 in the Firehole-Gibbon winter range and spring of 1987-90 in the Northern winter range. The probability that grizzly bears used a carcass was positively related to elevation and was lower within 400 m of a road, or within 5 km of a major recreational development compared to elsewhere. Carcass use peaked in April, coincident with peak ungulate deaths. Grizzly bears also were more likely to use carcasses in the Firehole-Gibbon compared to Northern Range study area. We attributed the effects of study area and elevation to the fact that grizzly bears den and are first active in the spring at high elevations and to differences in densities of competing scavengers. Probability of grizzly bear use was strongly related to body mass of carcasses on the Northern Range where densities of coyotes ( Canis latrans ) and black bears ( U. americanus ) appeared to be much higher than in the Firehole-Gibbon study area. We suggest that additional restrictions on human activity in ungulate winter ranges or movement of carcasses to remote areas could increase grizzly bear use of carrion. Fewer competing scavengers and greater numbers of adult ungulates vulnerable to winter mortality could have the same effect.

Wyoming↗

USGS1997

Grizzly bear predation rates on caribou calves in northeastern Alaska

During June 1993 and 1994, 11 radiocollared and 7 unmarked grizzly bears (Ursus arctos) were monitored visually (observation) from fixed-wing aircraft to document predation on calves of the Porcupine Caribou (Rangifer tarandus) Herd (PCH) in northeastern Alaska. Twenty-six (72%) grizzly bear observations were completed (???60 min) successfully (median duration = 180 min; ??95% CI = 136-181 min; range = 67-189 min) and 10 were discontinued (duration ???24 min) due to disturbance to the bear, or unfavorable weather conditions. Of the 26 successfully completed observations, 15 (58%) included predatory activity (encounter) directed at caribou calves and 8 (31%) included kills. Of 32 encounters, 9 resulted in kills, for a success rate of 28%. The median duration of encounters was 1 minute (??95% CI = 1-2 min; range = 1-6 min; n = 32;), and the median time spent at a kill was 14 minutes (??95% CI = 9-23 min; range = 6-56 min; n = 9). Sows with young (n = 4) killed more frequently (75%; P = 0.0178) than barren sows, boars, and consorting pairs combined (17%; n = 18). Estimated kill rate was highest for sows with young (6.3 kills/bear/day; n = 4), followed by barren sows (4.6 kills/bear/day; n = 5), boars (1.9 kills/bear/day; n = 5), and, finally, consorting pairs (1.0 kills/bear/day; n = 8). Estimated kill rate obtained via conventional radiotracking point surveys (4.8 kills/bear/day) was higher than that obtained via concurrent bear observations (3.1 kills/bear/day). Our research provides baseline estimates of predation rates by grizzly bears on caribou calves that will enhance the capability of wildlife professionals in managing populations of both predators and their prey.

Journal of Wildlife Management↗

USGS1996

How many grizzlies in Yellowstone?

Trend data indicate that the Yellowstone grizzly bear ( Ursus arctos horribilis ) population has been increasing in recent years, after a decline induced by closure of open garbage dumps in 1970-71. Current population size appears to be approaching a level where management to curb further increases might be desirable, even though it will be highly controversial. Continual close monitoring is essential for managers to know how to safeguard the population. Estimating total population size of an endangered or threatened species should be secondary to measuring essential population parameters, but nonetheless may be necessary to avoid misunderstandings. Knowledge of survival and reproductive rates is essential if causes of a decline are to be detected and corrected.

Wyoming↗

USGS1998

Grizzly bears and calving caribou: What is the relation with river corridors?

Researchers have debated the effect of the Trans-Alaska Pipeline (TAP) and associated developments to caribou ( Rangifer tarandus ) of the central Arctic herd (CAH) since the 1970s. Several studies have demonstrated that cows and calves of the CAH avoided the TAP corridor because of disturbance associated with the pipeline, whereas others have indicated that female caribou of the CAH avoided riparian habitats closely associated with the pipeline. This avoidance was explained as a predator-avoidance strategy. We investigated the relation between female caribou and grizzly bear ( Ursus arctos ) use of river corridors on the yet undisturbed calving grounds of the Porcupine caribou herd (PCH) in northeastern Alaska. On the coastal plain, caribou were closer to river corridors than expected (P = 0.038), but bear use of river corridors did not differ from expected (P = 0.740). In the foothills, caribou use of river corridors did not differ from expected (P = 0.520), but bears were farther from rivers than expected (P = 0.001). Our results did not suggest an avoidance of river corridors by calving caribou or a propensity for bears to be associated with riparian habitats, presumably for stalking or ambush cover. We propose that PCH caribou reduce the risks of predation to neonates by migrating to a common calving grounds, where predator swamping is the operational antipredator strategy. Consequently, we hypothesize that nutritional demands, not predator avoidance strategies, ultimately regulate habitat use patterns (e.g., use of river corridors) of calving PCH caribou.

Alaska↗

USGS1995

Biological consequences of relocating grizzly bears in the Yellowstone ecosystem

Relocating grizzly bears ( Ursus arctos ) from human/bear conflict situations has been a standard management procedure. Using data from Yellowstone National Park, we present components of situations that may affect the outcome of a relocation. Survival rates of transported bears were lower ( l x = 0.83) ( P = 0.001) than those not transported ( l x = 0.89). Survival was largely affected by whether the bear returned to the capture site ( P = 0.029). Return rate was most affected by distance transported ( P = 0.012) and age-sex group ( P = 0.014). Return rates decreased at distances -75 km, and subadult females returned least ( P = 0.050) often. Because of low survival and high return rates, transporting grizzly bears should be considered a final action to eliminate a conflict situation. However, transporting females must be considered a viable management technique because transports of some individuals have resulted in contributions to the population through successful reproduction.

Journal of Wildlife Management↗

USGS1995

Caribou calf mortality in Denali National Park, Alaska

Calf mortality is a major component of caribou (Rangifer tarandus) population dynamics, but little is known about the timing or causes of calf losses, or of characteristics that predispose calves to mortality. During 1984-87, we radiocollared 226 calves (≤3 days old) in the Denali Caribou Herd (DCH), an unhunted population utilized by a natural complement of predators, to determine the extent, timing, and causes of calf mortality and to evaluate influences of year, sex, birthdate, and birth mass on those losses. Overall, 39% of radio-collared calves died as neonates (≤15 days old), and 98% of those deaths were attributed to predation. Most neonatal deaths (85%) occurred within 8 days of birth. Few deaths occurred after the neonatal period (5, 10, and 0% of calves instrumented died during 16-30, 31-150, and >150 days of age, respectively). Survival of neonates was lower (P = 0.038) in 1985, following a severe winter, than during the other 3 years. In years other than 1985, calves born during the peak of calving (approx 50% of the total, born 5-8 days after calving onset) experienced higher (P < 0.001) neonatal survival than did other calves. Grizzly bears (Ursus arctos), wolves (Canis lupus), and unknown large predators (i.e., grizzly bears or wolves) accounted for 49, 29, and 16% of the neonatal deaths, respectively. The rate of bear-caused mortalities declined (P < 0.001) with calf age, and bears killed few calves >10 days old. Wolf predation was not related (P > 0.05) to calf age and peaked 10 days after onset of calving. Grizzly bear and wolf predation on neonates during the calving season was a limiting factor for the Denali Caribou Herd.

Journal of Wildlife Management↗

USGS1997

Use of lodgepole pine cover types by Yellowstone grizzly bears

Lodgepole pine ( Pinus contorta ) forests are a large and dynamic part of grizzly bear ( Ursus arctos ) habitat in the Yellowstone ecosystem. Research in other areas suggests that grizzly bears select for young open forest stands, especially for grazing and feeding on berries. Management guidelines accordingly recommend timber harvest as a technique for improving habitat in areas potentially dominated by lodgepole pine. In this paper I examine grizzly bear use of lodgepole pine forests in the Yellowstone area, and test several hypotheses with relevance to a new generation of management guidelines. Differences in grizzly bear selection of lodgepole pine cover types (defined on the basis of stand age and structure) were not pronounced. Selection furthermore varied among years, areas, and individuals. Positive selection for any lodgepole pine type was uncommon. Estimates of selection took 5-11 years or 4-12 adult females to stabilize, depending upon the cover type. The variances of selection estimates tended to stabilize after 3-5 sample years, and were more-or-less stable to slightly increasing with progressively increased sample area. There was no conclusive evidence that Yellowstone's grizzlies favored young (<40 yr) stands in general or for their infrequent use of berries. On the other hand, these results corroborated previous observations that grizzlies favored open and/or young stands on wet and fertile sites for grazing. These results also supported the proposition that temporally and spatially robust inferences require extensive, long-duration studies, especially for wide-ranging vertebrates like grizzly bears.

Wyoming↗

USGS2000

Effect of hibernation and reproductive status on body mass and condition of coastal brown bears

We investigated the effect of hibernation and reproductive status on changes in body mass and composition of adult female brown bears (Ursus arctos) on the Kenai Peninsula, Alaska. This information is fundamental to understanding nutritional ecology of wild brown bear populations. Six adult females handled in the fall and following spring (paired samples) lost 73 ± 22 kg (x̄ ± SD; 32 ± 10%) of fall body mass over 208 ± 19 days. Of this mass loss, 56 ± 22% (55 ± 22 kg) was lipid and 44 ± 22% (43 ± 21 kg) was lean body mass. Catabolism of lipid stores accounted for 88.4 ± 8.1% of the body energy used to meet maintenance demands. Overwinter differences in body composition of adult females assessed only once in either the fall (n = 21) or spring (n = 32) were similar to those of paired samples. Relative fatness of bears entering the den was positively related to the contribution of fat (%) to body mass (P < 0.01) and body energy (P < 0.01) losses during hibernation. Thus, relative fatness at the onset of fasting influences the relative proportion of lipid stores and lean body mass catabolized to meet protein and energy demands during hibernation. In the spring, lone females had greater body and lean masses than females with cubs of the year or yearlings. Lipid content was greatest in lone females in the fall. Studies using body mass and composition as indices of population health should consider season or reproductive class.

Journal of Wildlife Management↗

USGS2001

Negative binomial models for abundance estimation of multiple closed populations

Counts of uniquely identified individuals in a population offer opportunities to estimate abundance. However, for various reasons such counts may be burdened by heterogeneity in the probability of being detected. Theoretical arguments and empirical evidence demonstrate that the negative binomial distribution (NBD) is a useful characterization for counts from biological populations with heterogeneity. We propose a method that focuses on estimating multiple populations by simultaneously using a suite of models derived from the NBD. We used this approach to estimate the number of female grizzly bears (Ursus arctos) with cubs-of-the-year in the Yellowstone ecosystem, for each year, 1986-1998. Akaike's Information Criteria (AIC) indicated that a negative binomial model with a constant level of heterogeneity across all years was best for characterizing the sighting frequencies of female grizzly bears. A lack-of-fit test indicated the model adequately described the collected data. Bootstrap techniques were used to estimate standard errors and 95% confidence intervals. We provide a Monte Carlo technique, which confirms that the Yellowstone ecosystem grizzly bear population increased during the period 1986-1998.

Idaho, Montana, Wyoming↗

USGS1992

Yellowstone grizzly bear mortality, human habituation, and whitebark pine seed crops

The Yellowstone grizzly bear ( Ursus arctos horribilis ) population may be extirpated during the next 100-200 years unless mortality rates stabilize and remain at acceptable low levels. Consequently, we analyzed relationships between Yellowstone grizzly bear mortality and frequency of human habituation among bears and size of the whitebark pine ( Pinus albicaulis ) seed crop. During years of large seed crops, bears used areas within 5 km of roads and 8 km of developments half as intensively as during years of small seed crops because whitebark pine's high elevation distribution is typically remote from human facilities. On average, management trappings of bears were 6.2 times higher, mortality of adult females 2.3 times higher, and mortality of subadult males 3.3 times higher during years of small seed crops. We hypothesize that high mortality of adult females and subadult males during small seed crop years was a consequence of their tendency to range closest (of all sex-age cohorts) to human facilities; they also had a higher frequency of human habituation compared with adult males. We also hypothesize that low morality among subadult females during small seed crop years was a result of fewer energetic stressors compared with adult females and greater familiarity with their range compared with subadult males; mortality was low even though they ranged close to humans and exhibited a high frequency of human habituation. Human-habituated and food-conditioned bears were 2.9 times as likely to range within 4 km of developments and 3.1 times as often killed by humans compared with nonhabituated bears. We argue that destruction of habituated bears that use native foods near humans results in a decline in the overall ability of bears to use available habitat; and that the number and extent of human facilities in occupied grizzly bear habitat needs to be minimized unless habituated bears are preserved and successful ways to manage the associated risks to humans are developed.

Idaho, Montana, Wyoming↗

USGS1993

Multivariate model of female black bear habitat use for a Geographic Information System

Simple univariate statistical techniques may not adequately assess the multidimensional nature of habitats used by wildlife. Thus, we developed a multivariate method to model habitat-use potential using a set of female black bear ( Ursus americanus ) radio locations and habitat data consisting of forest cover type, elevation, slope, aspect, distance to roads, distance to streams, and forest cover type diversity score in the Ozark Mountains of Arkansas. The model is based on the Mahalanobis distance statistic coupled with Geographic Information System (GIS) technology. That statistic is a measure of dissimilarity and represents a standardized squared distance between a set of sample variates and an ideal based on the mean of variates associated with animal observations. Calculations were made with the GIS to produce a map containing Mahalanobis distance values within each cell on a 60- × 60-m grid. The model identified areas of high habitat use potential that could not otherwise be identified by independent perusal of any single map layer. This technique avoids many pitfalls that commonly affect typical multivariate analyses of habitat use and is a useful tool for habitat manipulation or mitigation to favor terrestrial vertebrates that use habitats on a landscape scale.

Arkansas↗

USGS1994

Polar bear maternity denning in the Beaufort Sea

The distribution of polar bears ( Ursus maritimus ) is circumpolar in the Northern Hemisphere, but known locations of maternal dens are concentrated in relatively few, widely scattered locations. Denning is either uncommon or unknown within gaps. To understand effects of industrial development and propose increases in hunting, the temporal and spatial distribution of denning in the Beaufort Sea must be known. We caputred and radiocollared polar bears between 1981 and 1991 and determined tht denning in the Beaufort Sea region was sufficient to account for the estimated population there. Of 90 dens, 48 were on drifting pack ice, 38 on land, and 4 on land-fast ice. The portions of dens on land was higher ( P = 0.029) in later compared with earlier years of the study. Bears denning on pack ice drifting as far as 997 km ( x = 385 km) while in dens. there was no difference in cub production by bears denning on land and pack ice ( P = 0.66). Mean entry and exit dates were 11 November and 5 April for land dens and 22 November and 26 March for pack-ice dens. Female polar bears captured in the Beaufort Sea appeared to be isolated from those caught east of Cape Bathurst in Canada. Of 35 polar bears that denned along the mainland coast of Alaska and Canada 80% denned between 137°00′W and 146°59′W . Bears followed to >1 den did not reuse sites and consecutive dens were 20-1,304 km apart. However radio-collared bears are largely faithful to substrate (pack-ice, land, and land-fast ice) and the general geographic area of previous dens. Bears denning on land may be vunerable to human activities such as hunting and industrial development. However, predictable denning chronology and lack of site fidelity indicate that many potential impacts on denning polar bears could be mitigated.

Alaska↗

USGS1983

Grizzly bear: habitat relationships in the Yellowstone area

Habitat use by grizzly bears ( Ursus arctos ) was studied from 1977 through 1979 in a 20,000-km 2 area with Yellowstone National Park in the center. Of 1826 aerial radio locations of 46 instrumental grizzlies, 90% were in timber. Three-fourths of the locations were 100 m or less from an edge between timber and an opening. Timber over 3 m tall with a canopy cover of 26-75% accounted for 50% of all activity sites from March through November. The Abies lasiocarpa/Vaccinium scoparium community alone contained 23% of the total activity sites and 35% of the forested activity sites. Of 507 observations of feeding activity, 45% were recorded in timber over 3 m tall with a canopy cover of 26-100%, 34% in timber with a 0.1-25% canopy cover, 20% in open habitats, and 3% in timber less than 3 m tall. Ninety-nine percent of examined day beds were in forested communities.

Yellowstone National Park↗

USGS1987

Prospects for Yellowstone grizzly bears

Recent analyses of data on the grizzly bear (Ursus arctos horribilis) population of Yellowstone National Park and its environs suggest the likelihood of a continuing decline in numbers if losses of fully adult females are not reduced. The size of the population is not known, and a simple projection model has been used to identify some inconsistencies in the available index data. Population dynamics calculations, based on Lotka's equation or a stochastic model, indicate a continuing decrease in numbers, although continued observations through radio-telemetry are needed to verify these trends. The margin between stabilizing the population and a continued decrease appears to be roughly the loss of 2 fully adult female bears per year. At present, the risk of extirpation of this population over the next 30 years appears to be small. Continued monitoring of survivorship will be needed, particularly because "recovery" of the population may be mainly characterized by a shift in the pattern of mortality, from adults to subadults, and not necessarily a reduction in absolute number of losses.

Idaho, Montana, Wyoming↗

USGS1994

Brown bear-human interactions associated with deer hunting on Kodiak Island

I compared distribution and range of brown bears ( Ursus arctos middendorffi ) with temporal and spatial distribution of Sitka black-tailed deer ( Odocoileus hemionus sitkensis ) hunting activity on westside Kodiak Island, Alaska, to examine impacts of deer hunting on bears. Mean number of bears that annually ranged ≤5 km from the coast, >5 km inland from the coast, or in both areas was 10, 8, and 11, respectively. Bears that exclusively or seasonally occupied the coast zone were usually classed as having moderate or high potential to interact with hunters because most hunter access and effort (>95%) was via the coast. Bears that ranged exclusively inland were considered unlikely to encounter hunters. Animals that ranged in both zones often (39%) moved inland during fall (Oct-Dec) and most bears (70%) denned in the inland zone. Females that denned near the coast entered dens later (x̄ = 22 Nov) than females that denned inland (x̄ = 12 Nov). Two radio-collared bears were known to raid deer-hunting camps and 9 other marked bears were observed by hunters or were located <200 m from hunting camps. Deer-hunter surveys revealed that more than two-thirds of the deer harvest occurred during October-November. About half of the hunters observed at least 1 bear during their hunt. Seven to 21% of the respondents reported having a threatening encounter with a bear and 5-26% reported losing deer meat to bears. Human-induced mortality to radio-collared bears occurred more often near the coast (5) than inland (3); 7 bears were harvested by sport hunters and 1 was killed (nonsport) in a Native village. Deer hunters killed 2 unmarked females in defense of life or property situations in the study area. High bear densities and concentrated deer-hunting activity combine to make conflicts unavoidable. Adverse impacts to bears can be minimized by maintaining low levels of human activity in inland areas and improving hunter awareness of bear ecology and behavior.

Alaska↗