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Results for “Journal of Vegetation Science”

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At least 37 records · Page 2Linked to original sources
USGS2016

Temperature is better than precipitation as a predictor of plant community assembly across a dryland region

Question How closely do plant communities track climate? Research suggests that plant species converge toward similar environmental tolerances relative to the environments that they experience. Whether these patterns apply to severe environments or scale up to plant community-level patterns of relative climatic tolerances is poorly understood. Using estimates of species' climatic tolerances acquired from occurrence records, we determined the contributions of individual species' climatic niche breadths and environmental filtering to the relationships between community-average climatic tolerances and the local climates experienced by those communities. Location Southwestern United States drylands. Methods Interspecific variation in niche breadth was assessed as a function of species' climatic optima (median climatic niche value). The relationships between climatic optima and tolerances were used as null expectations for the relationship between abundance-weighted mean climatic tolerances of communities and the local climate of that community. Deviations from this null expectation indicate that species with greater or lesser climatic tolerances are favoured relative to co-occurring species. The intensity of environmental filtering was estimated by comparing the range of climatic tolerances within each community to a null distribution generated from a random assembly algorithm. Results The temperature niches of species were consistently symmetrical and of similar breadths, regardless of their temperature optima. In contrast, precipitation niches were skewed toward wetter conditions, and niche breadth increased with increasing precipitation optima. At the community level, relationships with climate were much stronger for temperature than for precipitation. Furthermore, cold and heat were stronger assembly filters than drought or precipitation, with the intensity of environmental filtering increasing at both ends of climatic gradients. Community-average climatic tolerances did deviate significantly from null expectations, indicating that species with higher or lower relative climatic tolerances were favoured under certain conditions. Conclusions Despite strong water limitation of plant performance in dryland ecosystems, communities tracked variation in temperature much more closely, intimating strong responses to anticipated temperature increases. Furthermore, abundance distributions were biased toward species with higher or lower relative climatic tolerances under different climatic conditions, but predictably so, indicating the need for assembly models that include processes other than simple environmental filtering.

Journal of Vegetation Science↗
USGS2018

Individual species–area relationships in temperate coniferous forests

Questions What drives individual species–area relationships in temperate coniferous forests? Location Two 25.6‐ha forest plots on the Pacific Slope of North America, one in California, and one in Washington State. Methods We mapped all trees ≥1 cm in diameter and examined tree species diversity of their local neighbourhoods by calculating the individual species–area relationship for each species and for each of three diameter classes (saplings, mature trees and large‐diameter trees). Results In the California plot, small trees in four of the five major species occurred in neighbourhoods with higher levels of diversity than would be expected at random. In the Washington plot, small trees for four of five abundant species had neighbourhoods with lower than expected diversity at distances ≤5 m for small trees. However, at distances >5 m, all five species showed higher than expected diversity in their neighbourhoods. Larger trees at both plots tended to occur in neighbourhoods with lower than expected diversity, and no large‐diameter focal species had neighbourhoods with higher than expected diversity. Conclusion Diversity and co‐existence in temperate conifer‐dominated forests do not appear to be the result of random processes. Competitive interactions appear to dominate for the largest trees of most species, resulting in neighbourhoods with lower diversity. For smaller trees, we suggest that a positive response to environmental heterogeneity is the likely driver of neighbourhoods with higher than expected diversity, although we cannot rule out the possibility that facilitation or conspecific negative density dependence (CNDD) also play a role.

Journal of Vegetation Science↗
USGS2019

Regeneration of Metrosideros polymorpha forests in Hawaii after landscape‐level canopy dieback

Questions (a) Have Metrosideros polymorpha trees become re‐established in Hawaiian forests previously impacted by canopy dieback in the 1970s? (b) Has canopy dieback expanded since the 1970s? (c) Can spatial patterns from this dieback be correlated with habitat factors to model future dieback in this area? Study Site An 83,603 ha study area on the eastern slopes of Mauna Loa and Mauna Kea volcanoes on the island of Hawaii, USA. Methods We analyzed very‐high‐resolution imagery to assess status of Metrosideros polymorpha forests across the eastern side of the island of Hawaii. We generated 1,170 virtual vegetation plots with a 100‐m radius; 541 plots in areas mapped in 1977 with trees dead or mostly defoliated (dieback), and 629 plots in adjacent wet forest habitat, previously mapped as non‐dieback condition. In each plot we estimated the frequency of M. polymorpha trees that were dead or mostly defoliated, and the frequency of trees with healthy crowns. These results were combined with habitat data to produce a spatial model depicting probability of canopy dieback within the study area. Results Seventy‐nine percent of plots mapped in 1977 in dieback condition recovered their canopy and were now considered in non‐dieback condition. Ninety‐one percent of plots in previous non‐dieback areas were found to still have a healthy M. polymorpha canopy in 2015. A spatial model allowed us to identify areas within the study area with high, medium, and low probability of experiencing this same type of canopy dieback in the future. Conclusions Most former dieback areas mapped within the study area in 1977 now show recovery of the tree canopy through growth of new cohorts of young M. polymorpha trees. This suggests these forest communities are resilient to this type of canopy loss and tree death so long as other factors do not disrupt the natural regeneration process.

Hawaii↗
USGS2019

Fire severity and changing composition of forest understory plant communities

Questions Gradients of fire severity in dry conifer forests can be associated with variation in understory floristic composition. Recent work in dry conifer forests in California, USA, has suggested that more severely burned stands contain more thermophilic taxa (those associated with warmer and drier conditions), and that forest disturbance may therefore accelerate floristic shifts already underway due to climate change. However, it remains unknown how rapidly thermophilic taxa shifts occur following disturbance, how long such shifts are likely to persist, and how different thermophilic post‐disturbance communities are from pre‐disturbance communities. Location Colorado Front Range, USA. Methods We investigated these questions using a unique 15‐year vegetation plot dataset that captures pre‐ and post‐fire understory community composition across a gradient of fire severity in dry conifer forests, classifying taxa using the biogeographic affinity concept. Results Thermophilization (defined here as a decrease in the ratio of cool‐mesic taxa to warm‐xeric taxa, based on biogeographic affinity of paleobotanical lineages) was observed as early as one year post‐fire for all fire severity classes, but was stronger at sites that burned at higher severity. The ratio of cool‐mesic to warm‐xeric taxa recovered to pre‐fire levels within 10 years in stands that burned at low severity, but not in stands that burned at moderate or high severity. The process of thermophilization after high‐severity fire appears to be driven primarily by the gain of warm‐xeric taxa that were absent before the fire, but losses of cool‐mesic taxa, which did not return during the duration of the study, also played a role. Conclusions Decreases in canopy cover appear to be a main contributor to understory thermophilization. Fine‐scale heterogeneity in post‐fire forest structure is likely an important driver of floristic diversity, creating the microclimatic variation necessary to maintain floristic refugia for species mal‐adapted to increasingly warm and dry conditions.

Colorado↗
USGS2020

Soil water availability shapes species richness in mid-latitude shrub steppe plant communities

Questions Ecological communities are controlled by multiple, interacting abiotic and biotic factors that influence the distribution, abundance, and diversity of species. These processes jointly determine resource availability, resource competition, and ultimately species richness. For many terrestrial ecosystems in dryland climates, soil water availability is the most frequent limiting resource for plant species. We used field sampling coupled with process‐based soil water balance modeling to explore the relative importance of multiple macroclimatic, ecohydrological, and biotic variables on plant species and functional type richness at the landscape scale in dryland plant communities. Location Dryland plant communities dominated by big sagebrush ( Artemisia tridentata ) that span climatic and elevational gradients in Wyoming, USA. Methods We quantified species richness at 1,000 m 2 and used multiple regression to determine whether mean climatic conditions, multiple metrics of soil moisture from a soil water balance model (SOILWAT2), soil physical and chemical properties, and shrub stand structure (biotic) variables were related to species and functional type richness. Results Species richness varied between 16 and 54 across sites. We found that species and functional type richness were related to both macroclimate and ecohydrology, but ecohydrology explained slightly more variation than climate. Biotic variables were always secondary to macroclimate and ecohydrology in our models. Variance partitioning revealed that large portions of variability in species (~54%), forb (~47%), and grass (~40%) richness were explained by ecohydrological variables. Conclusions Our results highlight the importance of the spatial and temporal distribution of soil water for dryland plant species richness and suggest that documenting the ways in which climate, vegetation, and soil properties interact to determine soil water availability is critical for understanding biodiversity patterns in dryland plant communities. This work has relevance for other mid‐latitude, shrub‐dominated dryland plant communities where soil water availability strongly influences ecosystem structure and function.

Wyoming↗
USGS2020

Disentangling the effects of multiple fires on spatially interspersed sagebrush (Artemisia spp.) communities

Questions Relative to a landscape with a mosaic of two sagebrush community types and increasing fire frequency, we asked: (a) do vegetation characteristics vary significantly with number of times burned for each sagebrush community; (b) how do vegetation responses to different fire frequencies compare between the two sagebrush communities? Location Columbia Plateau Ecoregion, Washington, USA. Methods We sampled vegetation across a landscape that burned three times over a 10‐year period in two sagebrush community types that are interspersed on unique land forms: big sagebrush ( Artemisia tridentata ) communities that occur on small “mounds” and scabland sagebrush ( A. rigida ) communities that occur on surrounding “flats.” Spatially overlapping fires permitted a balanced sampling design to assess unburned and once‐, twice‐, and thrice‐burned locations for each land form/community type. We utilized a suite of statistical analyses to determine differences among plant functional groups and biomass among unburned/burned strata by land form and compared results between land forms. Results Big sagebrush and scabland sagebrush communities responded uniquely to multiple fires, due to different fuel loadings, fire severities, succession and invasion dynamics. Big sagebrush experienced nearly complete shrub loss and conversion from exotic‐invaded shrubland to exotic annual grassland after only one fire. In contrast, scabland sagebrush retained a minor shrub component and higher relative cover of native herbaceous species, even after three fires. Both communities retained cover of native perennial grasses, including shallow‐ and deep‐rooted species, likely reflecting decreasing fire intensity with number of times burned. Conclusions Despite different community‐level responses, increasing fire frequency is transforming the entire landscape to a non‐native/native grassland mix. Quantifying unique ecosystem responses to altered wildfire regimes is critical to understanding the relative resilience of communities to disturbance and their resistance to exotic species invasion (and community type conversion). Management actions may help to maintain spatial heterogeneity of ecosystems and fire‐tolerant native species.

Washington↗
USGS2023

Vegetation change over 140 years in a sagebrush landscape of the Rio Grande del Norte National Monument, New Mexico, USA

Questions Big sagebrush ( Artemisia tridentata ) ecosystems across the western United States have experienced many changes in ecosystem dynamics and vegetation composition over the last century due to livestock grazing, non-native species, and changing climate and fire regimes. We conducted the first systematic investigation of historical vegetation composition and vegetation change in a sagebrush landscape in the southwestern United States, asking whether sagebrush or grass dominated the landscape historically? Location The Rio Grande del Norte National Monument (RGDN), northern New Mexico, USA. Methods We combined General Land Office (GLO) surveys from 1881 with modern vegetation maps, field vegetation surveys, and sagebrush ages from growth ring analysis to test for changes in vegetation in the RGDN over the last 140 years. Results We found that big sagebrush presence across the study area increased significantly, from being present on 16% of section lines in 1881 to 79% in 2019, and only three section lines lost sagebrush presence during that period. Concurrently, the number of section lines with low grass index more than doubled since 1881, while moderate and high grass index declined. Grass declined equally in areas where sagebrush increased and areas with no change in sagebrush, suggesting that changes in both vegetation types were catalyzed by external factors, likely including overgrazing. The growth ring analysis of 93 sagebrush revealed a maximum age of 87 years and establishment in every decade since the 1930s, consistent with the GLO results. Conclusions The significant vegetation changes in the RGDN over the last century, including an increase of sagebrush, provide important context about the shifting mosaic of grasslands and shrublands relevant to current and future management and ecosystem dynamics.

New Mexico↗
USGS2024

Trends in plant cover derived from vegetation plot data using ordinal zero-augmented beta regression

Questions Plant cover values in vegetation plot data are bounded between 0 and 1, and cover is typically recorded in discrete classes with non-equal intervals. Consequently, cover data are skewed and heteroskedastic, which hampers the application of conventional regression methods. Recently developed ordinal beta regression models consider these statistical difficulties. Our primary question is whether we can detect species trends in vegetation plot time series data with this modelling approach. A second question is whether trends in cover have additional value compared to trends in occurrence, which are easier to assess for practitioners. Location The Netherlands, Western Europe. Methods We used vegetation plot data collected from 10,000 fixed plots which were surveyed once every four years during 1999–2022. We used the ordinal zero-augmented beta regression (OZAB) model, a hierarchical model consisting of a logistic regression for presence and an ordinal beta regression for cover. We adapted the OZAB model for longitudinal data and produced estimates of cover and occurrence for each four-year period. Thereafter we assessed trends in cover and in occurrence across all periods. Results We found evidence of a trend in cover in 318 out of the 721 species (44%) with sufficient data. Most species showed similar directional trends in occurrence and percent cover. No trend in occurrence was detected for 64 species that had evidence of a trend in cover. Declining species had stronger relative changes in cover than in occurrence. Conclusions Our model enables researchers to detect trends in cover using longitudinal vegetation plot data. Cover trends often corroborated trends in occurrence, but we also regularly found trends in cover even in the absence of evidence for trends in occurrence. Our approach thus contributes to a more complete picture of (changes in) vegetation composition based on large monitoring data sets.

Journal of Vegetation Science↗
USGS2006

Sample size and the detection of a hump-shaped relationship between biomass and species richness in Mediterranean wetlands

Questions: What is the observed relationship between biomass and species richness across both spatial and temporal scales in communities of submerged annual macrophytes? Does the number of plots sampled affect detection of hump-shaped pattern? Location: Don??ana National Park, southwestern Spain. Methods: A total of 102 plots were sampled during four hydrological cycles. In each hydrological cycle, the plots were distributed randomly along an environmental flooding gradient in three contrasted microhabitats located in the transition zone just below the upper marsh. In each plot (0.5 m x 0.5 m), plant density and above- and below-ground biomass of submerged vegetation were measured. The hump-shaped model was tested by using a generalized linear model (GLM). A bootstrap procedure was used to test the effect of the number of plots on the ability to detect hump-shaped patterns. Result: The area exhibited low species density with a range of 1 - 9 species and low values of biomass with a range of 0.2 - 87.6 g-DW / 0.25 m2. When data from all years and all microhabitats were combined, the relationships between biomass and species richness showed a hump-shaped pattern. The number of plots was large enough to allow detection of the hump-shaped pattern across microhabitats but it was too small to confirm the hump-shaped pattern within each individual microhabitat. Conclusion: This study provides evidence of hump-shaped patterns across microhabitats when GLM analysis is used. In communities of submerged annual macrophytes in Mediterranean wetlands, the highest species density occurs in intermediate values of biomass. The bootstrap procedure indicates that the number of plots affects the detection of hump-shaped patterns. ?? IAVS; Opulus Press.

Journal of Vegetation Science↗
USGS2007

Species richness and soil properties in Pinus ponderosa forests: A structural equation modeling analysis

Question: How are the effects of mineral soil properties on understory plant species richness propagated through a network of processes involving the forest overstory, soil organic matter, soil nitrogen, and understory plant abundance? Location: North-central Arizona, USA. Methods: We sampled 75 0.05-ha plots across a broad soil gradient in a Pinus ponderosa (ponderosa pine) forest ecosystem. We evaluated multivariate models of plant species richness using structural equation modeling. Results: Richness was highest at intermediate levels of understory plant cover, suggesting that both colonization success and competitive exclusion can limit richness in this system. We did not detect a reciprocal positive effect of richness on plant cover. Richness was strongly related to soil nitrogen in the model, with evidence for both a direct negative effect and an indirect non-linear relationship mediated through understory plant cover. Soil organic matter appeared to have a positive influence on understory richness that was independent of soil nitrogen. Richness was lowest where the forest overstory was densest, which can be explained through indirect effects on soil organic matter, soil nitrogen and understory cover. Finally, model results suggest a variety of direct and indirect processes whereby mineral soil properties can influence richness. Conclusions: Understory plant species richness and plant cover in P. ponderosa forests appear to be significantly influenced by soil organic matter and nitrogen, which are, in turn, related to overstory density and composition and mineral soil properties. Thus, soil properties can impose direct and indirect constraints on local species diversity in ponderosa pine forests. ?? IAVS; Opulus Press.

Journal of Vegetation Science↗
USGS2009

Distribution and conservation status of coastal sage scrub in southwestern California

A landscape-based characterization of vegetation has been developed for southwestern California using satellite imagery, air photos, existing vegetation maps, and field data. Distribution maps of nine dominant coastal scrub species and 13 species assemblages that were identified by divisive information analysis have been analyzed to quantify spatial patterns of species co-occurrence. Three general distribution patterns are identified that suggest the Diegan, Ventaran and Riversidian Associations identified by other workers. Vegetation data have also been related to land ownership and management to assess the conservation status of upland plant communities. A large proportion of the mapped distribution of species and vegetation types is on private land, and several taxa show less than 4 % of mapped distribution in nature reserves. The analysis highlights the need to extend current conservation planning efforts into the northern part of the region to encompass areas where Salvia leucophylla is a frequent community dominant.

Journal of Vegetation Science↗
USGS2000

Factors associated with plant species richness in a coastal tall-grass prairie

In this study we examine the factors associated with variations in species richness within a remnant tall-grass prairie in order to gain insight into the relative importance of controlling variables. The study area was a small, isolated prairie surrounded by wetlands and located within the coastal prairie region, which occurs along the northwestern Gulf of Mexico coastal plain. Samples were taken along three transects that spanned the prairie. Parameters measured included micro-elevation, soil characteristics, indications of recent disturbance, above-ground biomass (including litter), light penetration through the plant canopy, and species richness. Species richness was found to correlate with micro-elevation, certain soil parameters, and light penetration through the canopy, but not with above-ground biomass. Structural equation analysis was used to assess the direct and indirect effects of micro-elevation, soil properties, disturbance, and indicators of plant abundance on species richness. The results of this analysis showed that observed variations in species richness were primarily associated with variations in environmental effects (from soil and microtopography) and were largely unrelated to variations in measures of plant abundance (biomass and light penetration). These findings suggest that observed variations in species richness in this system primarily resulted from environmental effects on the species pool. These results fit with a growing body of information that suggests that environmental effects on species richness are of widespread importance.

Journal of Vegetation Science↗
USGS2001

Effect of flooding and draw-down disturbance on germination from a seashore meadow seed bank

The objective of this study was to investigate the effects of flooding and draw-down on the germination from the coastal grassland seed banks and to determine whether the effect of flooding varies between the delta and the seashore. Seed bank samples were collected from three shore transects in SW Finland, two on the shore of the Baltic Sea and one on the delta of River Kokemäenjoki. Samples were germinated in non-flooded and flooded conditions for over a month, after which both treatments were maintained in non-flooded conditions. A total of 9267 seedlings of 47 species germinated and mean density of seeds in the soil was ca. 84 000/m 2 . Most of the seedlings were monocots (98%) and perennials (98%). Ca. 30–40% of the species found in the above-ground vegetation had a seed bank including the majority of the most abundant species. The number of seeds and species richness increased as the organic layer became thicker. The organic layer was thicker in the seashore samples and the seed bank was significantly larger than in the delta. The flooding and draw-down treatment significantly increased the number of germinating seedlings in the seashore and also increased species richness in two transects, one in the delta and the other in the seashore. Two species, Schoenoplectus tabernaemontani and Typha latifolia , had significantly higher germination in the flooded treatment than in the non-flooded. Apparently, many species in these coastal grasslands have adapted to flood disturbance and for seeds of some species flooding may work as a positive signal, possibly breaking dormancy.

Pori↗
USGS2009

Stem demography and post-fire recruitment of a resprouting serotinous conifer

The contribution of resprouts and seedling recruitment to post-fire regeneration of the South African fynbos conifer Widdringtonia nodiflora was compared eight months after wildfires in 1990. Stems on all trees were killed by fire but resprouting success was > 90 % at all but one site. A demographic study of burned skeletons revealed that prior to these fires, nearly all plants were multi-stemmed (4–9 stems/plant) and multi-aged, indicating continuous sprout production between fires. All stems were killed by these 1990 fires and at most sites > 90 % of the stems were burned to ground level. All diameter stems were susceptible to such incineration as, at most sites, there was no difference in average diameter of stems burned to ground level and those left standing. Individual genets usually had all ramets incinerated to ground level or all ramets charred, but intact, suggesting certain micro-sites burned hotter, whereas other sites were somewhat protected. Although not true of the 1990 fires, there was evidence that occasionally Widdring-tonia stems may survive fire. At one site, four of the 16 plants sampled had a burned stem twice as old as the oldest burned stem on the other 12 plants at the site, suggesting some stems had survived the previous fire (ca. 1970) and this conclusion was supported by fire-scars on these four stems that dated to ca. 1970. Based on the highly significant correlation between stem diameter and cone density left standing after the 1990 fires, we calculated that for most sites > 80 % of the initial cone crop was incinerated by fire. This is important because we observed a strong relationship between size of the canopy cone crop surviving fire and post-fire seedling recruitment. Under these conditions we hypothesize that sprouting confers a selective advantage to genets when fires cause heavy losses of seed. The infrequent occurrence of sprouting in the Cupressaceae suggests the hypothesis that resprouting is an apomorphic or derived trait in Widdringtonia. Data from this study suggests resprouting provides a selective advantage under severe fynbos fires, which are not only 'stand-replacing fires,’but also are intense enough to incinerate cone-bearing stems.

Journal of Vegetation Science↗
USGS2003

Temporal species richness-biomass relationships along successional gradients

Diversity-biomass relationships are frequently reported to be hump-shaped over space at a given time. However, it is not yet clear how diversity and biomass change simultaneously and how they are related to each other over time (e.g. in succession) at one locality. This study develops a temporal model based on the projected changes of various community variables in a generalized terrestrial environment after fire and uses post-fire succession data on Santa Monica Mountains of southern California and other published succession data to examine the temporal diversity-biomass relationships. The results indicate that in the early stages of succession, both diversity and biomass increase and a positive relationship appears, while in the late stages of succession, biomass continued to increase but diversity usually declines; thus a negative relationship may be observed. When the scales of measurement become sufficiently large so that the measured diversity and biomass cross various stages of succession, a 'hump-shaped' relationship can emerge. The diversity-biomass relationship appears to be concordant in space and time when appropriate scales are used. Formerly proposed explanations for spatial patterns may well apply to the temporal patterns (particularly colonization, facilitation and competitive exclusion).

Journal of Vegetation Science↗
USGS2003

Temporal species richness-biomass relationships along successional gradients

Diversity-biomass relationships are frequently reported to be hump-shaped over space at a given time. However, it is not yet clear how diversity and biomass change simultaneously and how they are related to each other overtime (e.g. in succession) at one locality. This study develops a temporal model based on the projected changes of various community variables in a generalized terrestrial environment after fire and uses post-fire succession data on Santa Monica Mountains of southern California and other published succession data to examine the temporal diversity-biomass relationships. The results indicate that in the early stages of succession, both diversity and biomass increase and a positive relationship appears, while in the late stages of succession, biomass continued to increase but diversity usually declines; thus a negative relationship may be observed. When the scales of measurement become sufficiently large so that the measured diversity and biomass cross various stages of succession, a 'hump-shaped' relationship can emerge. The diversity-biomass relationship appears to be concordant in space and time when appropriate scales are used. Formerly proposed explanations for spatial patterns may well apply to the temporal patterns (particularly colonization, facilitation and competitive exclusion).

Journal of Vegetation Science↗
USGS1996

Exchangeable hydrogen explains the pH of spodosol Oa horizons

The chemistry of extremely acid Oa horizons does not conform to traditional pH, Al, and base saturation relationships. Results from two separate studies of northeastern U.S. forested soils were used to investigate relationships between pH in water or dilute salt solutions and other soil characteristics. In Oa horizons with pH below 4, soil pH in dilute CaCl 2 solution was correlated with exchangeable H + measured either by titration ( r = −0.88, P = 0.0001, n = 142) or by electrode ( r = −0.89, P = 0.0001, n = 45). Exchangeable H + expressed as a percentage of the cation-exchange capacity (CEC) was linear with pH and showed similar slopes for data from both studies. For all samples, pHw = 4.21 − 1.80 × H + /CEC ( R 2 = 0.69, n = 194). The reciprocal of the H + /CEC ratio is base saturation with Al added to the bases. Because of the low pH, exchangeable Al does not appear to behave as an acid. Exchangeable H + remains an operationally defined quantity because of the difficulty in separating exchange and hydrolysis reactions. In a variety of neutral-salt extractants, concentrations of H + were correlated with 0.1 M BaCl 2 -exchangeable H + ( r > 0.91, P = 0.0001, n = 26) regardless of the strength of the extract. Nine successive extractions with 0.33 m M CaCl 2 removed more H + than was removed by single batch extractions with either 1 M KCl or 0.1 M BaCl 2 (average H + of 70, 43, and 49 mmol kg −1 , respectively for 26 samples). The data showed little difference in the chemical behavior of Oa horizons from a variety of geographical sites and vegetation types.

Maine, New Hampshire, New York, Vermont↗