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25 records · Page 2Linked to original sources

Natural history of a northern population of twin-spotted rattlesnakes, Crotalus pricei

The twin-spotted rattlesnake ( Crotalus pricei ) is a small-bodied pitviper that has received little attention in the literature to date. The species reaches the northern limit of its range in southeastern Arizona, where it inhabits higher elevations than any of the state's 10 other rattlesnake species. During 1997–2000, we captured, measured, and marked 127 C. pricei in Arizona's Chiricahua Mountains between 2530 and 2900 m elevation. We also used radiotelemetry to track the movements of 16 C. pricei in the study area during 1997–1998. Mean (± SE) snout–vent length of C. pricei was 387.8 ± 8.3 mm (range = 168–572), and mean mass was 53.5 ± 3.3 g (range = 3.6–188.5). Based on fecal analyses, lizards constituted the bulk of prey (74%), but the diet of C. pricei also included mammals, birds, and a conspecific. Mating was concentrated in August and early September and parturition took place during late July and August. Mean number of embryos was 3.94 ± 0.34 (range = 1–6) and female reproduction appeared biennial or less frequent. Based on shed and growth rates, female C. pricei develop embryos at 4–5 years of age. Gravid females maintained warmer body temperatures relative to substrate temperature than nongravid females or males, presumably by spending more time basking than other snakes. Radiotelemetry revealed that movement patterns varied from year to year, as males moved over six times farther per week during the 1998 monsoon season (July to September) than during the 1997 monsoon season. Additionally, use of talus slopes by males decreased during 1998. During dry years, such as 1998, males may be forced off talus into cooler microclimates where resources are less concentrated than on talus.

Journal of Herpetology↗

Elevational gradient in clutch size of Red-faced Warblers

Our understanding of life history evolution has benefited from debates regarding the underlying causes, and geographic ubiquity, of spatial patterns in avian clutch sizes. Past studies have revealed that birds lay smaller clutch sizes at higher elevation. However, in most previous studies, investigators have failed to adequately control for elevational differences in breeding phenology. To better understand the elevational gradient in avian clutch size, we need to know how clutch size changes across the entire elevational breeding range of a species (i.e., the shape of the relationship between elevation and clutch size), and whether the elevational gradient in clutch size is merely an artifact of elevational gradients in breeding phenology or breeding season length. We examined the relationship between breeding elevation and clutch size of Red-faced Warblers ( Cardellina rubrifrons ) along a 1000-m elevational gradient in Arizona. Our objectives were to determine how clutch size changed with elevation, and if the relationship between clutch size and elevation merely reflected elevational changes in breeding season length or phenology. The proportion of 5-egg clutches decreased and the proportion of 3- and 4-egg clutches increased non-linearly with increasing elevation, even after controlling for the elevational gradient in nest initiation date. Thus, average clutch size declined across the elevational breeding range of Red-faced Warblers, but this decline was not due to elevational variation in breeding phenology. Timing of breeding changed, but the duration of the breeding season did not change appreciably across the elevational gradient. Hence, elevational differences in breeding season length or breeding phenology cannot explain why Red-faced Warblers (and perhaps other birds) breeding at higher elevations have smaller clutches.

Journal of Field Ornithology↗

Stratigraphy and Mesozoic–Cenozoic tectonic history of northern Sierra Los Ajos and adjacent areas, Sonora, Mexico

Geologic mapping in the northern Sierra Los Ajos reveals new stratigraphic and structural data relevant to deciphering the Mesozoic–Cenozoic tectonic evolution of the range. The northern Sierra Los Ajos is cored by Proterozoic, Cambrian, Devonian, Mississippian, and Pennsylvanian strata, equivalent respectively to the Pinal Schist, Bolsa Quartzite and Abrigo Limestone, Martin Formation, Escabrosa Limestone, and Horquilla Limestone. The Proterozoic–Paleozoic sequence is mantled by Upper Cretaceous rocks partly equivalent to the Fort Crittenden and Salero Formations in Arizona, and the Cabullona Group in Sonora, Mexico. Absence of the Upper Jurassic–Lower Cretaceous Bisbee Group below the Upper Cretaceous rocks and above the Proterozoic–Paleozoic rocks indicates that the Sierra Los Ajos was part of the Cananea high, a topographic highland during the Late Jurassic and Early Cretaceous. Deposition of Upper Cretaceous rocks directly on Paleozoic and Proterozoic rocks indicates that the Sierra Los Ajos area had subsided as part of the Laramide Cabullona basin during Late Cretaceous time. Basal beds of the Upper Cretaceous sequence are clast-supported conglomerate composed locally of basement (Paleozoic) clasts. The conglomerate represents erosion of Paleozoic basement in the Sierra Los Ajos area coincident with development of the Cabullona basin. The present-day Sierra Los Ajos reaches elevations of greater than 2600 m, and was uplifted during Tertiary basin-and-range extension. Upper Cretaceous rocks are exposed at higher elevations in the northern Sierra Los Ajos and represent an uplifted part of the inverted Cabullona basin. Tertiary uplift of the Sierra Los Ajos was largely accommodated by vertical movement along the north-to-northwest-striking Sierra Los Ajos fault zone flanking the west side of the range. This fault zone structurally controls the configuration of the headwaters of the San Pedro River basin, an important bi-national water resource in the US-Mexico border region.

Sierra Los Ajos↗

Geographical and climatic limits of needle types of one- and two-needled pinyon pines

Aim: The geographical extent and climatic tolerances of one- and two-needled pinyon pines (Pinus subsect. Cembroides) are the focus of questions in taxonomy, palaeoclimatology and modelling of future distributions. The identification of these pines, traditionally classified by one- versus two-needled fascicles, is complicated by populations with both one- and two-needled fascicles on the same tree, and the description of two more recently described one-needled varieties: the fallax-type and californiarum-type. Because previous studies have suggested correlations between needle anatomy and climate, including anatomical plasticity reflecting annual precipitation, we approached this study at the level of the anatomy of individual pine needles rather than species. Location: Western North America. Methods: We synthesized available and new data from field and herbarium collections of needles to compile maps of their current distributions across western North America. Annual frequencies of needle types were compared with local precipitation histories for some stands. Historical North American climates were modelled on a c. 1-km grid using monthly temperature and precipitation values. A geospatial model (ClimLim), which analyses the effect of climate-modulated physiological and ecosystem processes, was used to rank the importance of seasonal climate variables in limiting the distributions of anatomical needle types. Results: The pinyon needles were classified into four distinct types based upon the number of needles per fascicle, needle thickness and the number of stomatal rows and resin canals. The individual needles fit well into four categories of needle types, whereas some trees exhibit a mixture of two needle types. Trees from central Arizona containing a mixture of Pinus edulis and fallax-type needles increased their percentage of fallax-type needles following dry years. All four needle types occupy broader geographical regions with distinctive precipitation regimes. Pinus monophylla and californiarum-type needles occur in regions with high winter precipitation. Pinus edulis and fallax-type needles are found in regions with high monsoon precipitation. Areas supporting californiarum-type and fallax-type needle distributions are additionally characterized by a more extreme May-June drought. Main conclusions: These pinyon needle types seem to reflect the amount and seasonality of precipitation. The single needle fascicle characterizing the fallax type may be an adaptation to early summer or periodic drought, while the single needle of Pinus monophylla may be an adaptation to summer-autumn drought. Although the needles fit into four distinct categories, the parent trees are sometimes less easily classified, especially near their ancestral Pleistocene ranges in the Mojave and northern Sonoran deserts. The abundance of trees with both one- and two-needled fascicles in the zones between P. monophylla, P. edulis and fallax-type populations suggest that needle fascicle number is an unreliable characteristic for species classification. Disregarding needle fascicle number, the fallax-type needles are nearly identical to P. edulis, supporting Little's (1968) initial classification of these trees as P. edulis var. fallax, while the californiarum-type needles have a distinctive morphology supporting Bailey's (1987) classification of this tree as Pinus californiarum.

Journal of Biogeography↗

Middle Tertiary extension recorded by lacustrine fan-delta deposits, Plush Ranch Basin, western Transverse Ranges, California

The Plush Ranch Formation (upper Oligocene and lower Miocene) consists of more than 1800 m of nonmarine sedimentary and volcanic rocks that record the history of an extensional basin referred to here as the Plush Ranch basin. Distinctive depositional facies, provenance, and sediment transport directions along each basin margin suggest an asymmetric basin shape that is consistent with a half-graben origin. The northern basin margin consists of sandstone-dominated alluvial-plain deposits (0.1-1.5 m thick, normally graded, lenticular sandstone beds). Small deltaic sequences 1-2 m thick were formed where these alluvial systems flowed southward into a lake. Lenses of massive, boulder-rich granitic breccia that represent rockslide deposits derived from a nearby northern granitic provenance interfinger with the alluvial-plain facies. In contrast to the northern margin, the southern basin margin is represented by coarse-grained fan-delta deposits. Matrix- and clast-supported lenticular conglomerate beds 0.2-5 m thick with interbedded trough-cross-bedded pebbly sandstone represent braided-stream and flood-flow and/or noncohesive debris-flow deposits of alluvial fans that drained a highland area to the south. The alluvial-fan deposits interfinger to the north with several types of subaqueous sediment-gravity-flow facies including turbidite sandstone beds and matrix-supported debris-flow conglomerate. Each of the basin-margin depositional systems grades basinward and to the east into lacustrine deposits that include organic-rich dark shale, evaporite, and limestone. The lacustrine deposits represent the central and eastern parts of the Plush Ranch basin, which received little coarse siliciclastic sediment. Basalt deposits that are at least 50 m thick in the west and thicken eastward are interbedded mainly with the lacustrine facies. The southern margin of the Plush Ranch basin formed along a north-dipping, normal-slip fault along which dip separation increased toward the southwest; the northern margin developed on the tilted hanging-wall block of this fault. This fault was later reactivated in post-middle Miocene time as the present left-lateral strike-slip Big Pine fault. The Plush Ranch is one of several extensional and transtensional basins that formed in southern California and western Arizona about 25-20 Ma as a response to the change from a convergent to a strikeslip tectonic regime along western North America.

California↗

Archaeological sites in Grand Canyon National Park along the Colorado River are eroding owing to six decades of Glen Canyon Dam operations

The archaeological record documenting human history in deserts is commonly concentrated along rivers in terraces or other landforms built by river sediment deposits. Today that record is at risk in many river valleys owing to human resource and infrastructure development activities, including the construction and operation of dams. We assessed the effects of the operations of Glen Canyon Dam – which, since its closure in 1963, has imposed drastic changes to flow, sediment supply and distribution, and riparian vegetation – on a population of 362 archaeological sites in the Colorado River corridor through Grand Canyon National Park, Arizona, USA. We leverage 50 years of evidence from aerial photographs and more than 30 years of field observations and measurements of archaeological-site topography and wind patterns to evaluate changes in the physical integrity of archaeological sites using two geomorphology-based site classification systems. We find that most archaeological sites are eroding; moreover, most are at increased risk of continuing to erode, due to six decades of operations of Glen Canyon Dam. Results show that the wind-driven (aeolian) supply of river-sourced sand, essential for covering archaeological sites and protecting them from erosion, has decreased for most sites since 1973 owing to effects of long-term dam operations on river sediment supply and riparian vegetation expansion on sandbars. Results show that the proportion of sites affected by erosion from gullies controlled by the local base-level of the Colorado River has increased since 2000. These changes to landscape processes affecting archaeological site integrity limit the ability of the National Park Service and Grand Canyon-affiliated Native American Tribes to achieve environmental management goals to maintain or improve site integrity in situ . We identify three environmental management opportunities that could be used to a greater extent to decrease the risk of erosion and increase the potential for in-situ preservation of archaeological sites. Environmental management opportunities are: 1) sediment-rich controlled river floods to increase the aeolian supply of river-sourced sand, 2) extended periods of low river flow to increase the aeolian supply of river-sourced sand, 3) the removal of riparian vegetation barriers to the aeolian transport of river-sourced sand.

Arizona↗

Implications of low-temperature cooling history on a transect across the Colorado Plateau-Basin and Range boundary, west central Arizona

Fission track ages of apatite and zircon from metamorphic, plutonic, and sedimentary rocks along a 80-km transect across the Colorado Plateau-Basin and Range boundary in west central Arizona show differences in the low-temperature cooling histories between the provinces. The transect extends from Cypress Mountain in the Colorado Plateau transition zone to the eastern Buckskin Mountains in the Basin and Range. Along the northeast margin of the Basin and Range province, metamorphic rocks exposed in the footwall of a major detachment fault system yield zircon and apatite fission track ages of 16–10 Ma. These ages are similar to K-Ar fusion ages of biotite and age minima of K-feldspar on 40 Ar/ 39 Ar age spectra and collectively indicate rapid cooling. One K-feldspar age spectrum has an age maximum of about 22.8 Ma, an age minimum of 11.8 Ma, and a spectrum whose shape is suggestive of reheating, possibly in middle Miocene time. The heating event was probably related to hydrothermal activity during emplacement of Cu and Mn deposits in and above the detachment fault zone. Effects of this heating are only locally detected in rocks above the detachment fault. In the Poachie Range fission track ages of apatite and zircon are 60–50 and 80–70 Ma, respectively. The disparity between the apatite and zircon ages indicates that the rocks cooled slowly in Late Cretaceous and early Tertiary time, probably due to gradual uplift and erosion. Total uplift and denudation in the area of the Poachie Range since Cretaceous time is 6 km or more. To the northeast of the range, fission track ages of apatite and zircon increase and diverge, indicating that apparent uplift decreased in that direction. The apatite ages from the Poachie Range are concordant with early Tertiary hornblende ages determined in other studies of lower plate rocks near the southwest end of the transect. The ages represent cooling of crystalline rocks after Cretaceous regional metamorphism and magmatism. Near Bagdad, 20 km northeast of the Poachie Range, 2 km or less of erosion has occurred since intrusion of high-level plutons and dikes and caldera formation in Late Cretaceous time. Remnants of an erosion surface that developed in middle Tertiary time are preserved in the transition zone. Volcanic and sedimentary rocks at least as old as early Miocene were deposited on the erosion surface and filled valleys cut into it. Dissection of these deposits began about 8 Ma. We interpret these data combined with those from other studies to indicate that in Cretaceous time southward thrusting and later extensive magmatism in the middle crust led to thickening and heating of the crust. The Cretaceous igneous rocks at Bagdad are high-level manifestations of this magmatism. Uplift and slow cooling occurred in Late Cretaceous and early Tertiary time. In late Oligocene and early Miocene time during northeast-southwest extension, middle crustal rocks moved southwest put from beneath the southwest margin of the transition zone. Tectonic denudation rapidly exposed the crust that had been brought up from a depth of 10 km or more and rapidly cooled in the eastern Buckskin and Harcuvar mountains. Middle Miocene reheating occurred locally in the lower plate, along the detachment, and in nearby parts of the upper plate.

Journal of Geophysical Research↗