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Reproductive isolation and the expansion of an invasive hybrid swarm

Biological invasions involving hybridization proceed according to prezygotic and postzygotic reproductive isolating mechanisms. Yet few comparisons of reproductive isolation have been carried out to understand how different mechanisms prevent or promote invasions involving hybridization. Here we present a study of prezygotic and postzygotic isolation between non-native red shiner ( Cyprinella lutrensis ) and native blacktail shiner ( C. venusta stigmatura ) from the Coosa River basin (USA) to better understand the formation and expansion of invasive hybrid swarms. We conducted spawning trials to measure mating preferences and raised broods from crosses to assay hybrid viability through early juvenile development. Females of both species were more responsive to conspecific mates, although blacktail shiner females responded more often to heterospecific mates than did red shiner females. Fecundity of red shiner females was also higher than blacktail shiner females. Heterospecific crosses resulted in lower fertilization and egg hatching rates, but we found no other evidence of inviability. Rather, we found comparatively low larval mortality of F1 hybrids, which is suggestive of heterosis. These findings support prior inferences of assortative mating from genetic descriptions of hybridization, and that the invasion in the Coosa River is likely proceeding due to interspecific competition and intrinsic hybrid viability.

Biological Invasions

Assessing the impact of charr research past, present, and future

The 9th International Charr Symposium convened on 18–21 June 2018, in Duluth, Minnesota, USA to gather scientists with an interest in charr biology and management from the entire geographical range of the genus Salvelinus. The symposium was attended by 169 individuals from six countries, and included 99 oral and 32 poster presentations, 28 of which were published in the ensuing proceedings. Topic areas of presentations and publications included: (1) cultural anthropology; (2) genetics, evolution, taxonomy and systematics; (3) behavior, movement and habitat; (4) reproductive ecology, developmental ontogeny and physiology; (5) population ecology, dynamics and life history; (6) trophic ecology, parasites, predators, toxicology and pollution; and (7) management. Research reported in these proceedings has built upon the history of earlier symposiums and will be continued with the 10th symposium scheduled to convene in Nikko City, Tochigi Prefecture, Japan, in 2021.

Hydrobiologia

Dissolution of aragonite-strontianite solid solutions in nonstoichiometric Sr (HCO3)2-Ca (HCO3)2-CO2-H2O solutions

Synthetic strontianite-aragonite solid-solution minerals were dissolved in CO 2 -saturated non-stoichiometric solutions of Sr(HCO 3 ) 2 and Ca(HCO 3 ) 2 at 25°C. The results show that none of the dissolution reactions reach thermodynamic equilibrium. Congruent dissolution in Ca(HCO 3 ) 2 solutions either attains or closely approaches stoichiometric saturation with respect to the dissolving solid. In Sr(HCO 3 ) 2 solutions the reactions usually become incongruent, precipitating a Sr-rich phase before reaching stoichiometric saturation. Dissolution of mechanical mixtures of solids approaches stoichiometric saturation with respect to the least stable solid in the mixture. Surface uptake from subsaturated bulk solutions was observed in the initial minutes of dissolution. This surficial phase is 0–10 atomic layers thick in Sr(HCO 3 ) 2 solutions and 0–4 layers thick in Ca(HCO 3 ) 2 solutions, and subsequently dissolves and/or recrystallizes, usually within 6 min of reaction. The initial transient surface precipitation (recrystallization) process is followed by congruent dissolution of the original solid which proceeds to stoichiometric saturation, or until the precipitation of a more stable Sr-rich solid. The compositions of secondary precipitates do not correspond to thermodynamic equilibrium or stoichiometric saturation states. X-ray photoelectron spectroscopy (XPS) measurements indicate the formation of solid solutions on surfaces of aragonite and strontianite single crystals immersed in Sr(HCO 3 ) 2 and Ca(HCO 3 ) 2 solutions, respectively. In Sr(HCO 3 ) 2 solutions, the XPS signal from the outer ~ 60 Å on aragonite indicates a composition of 16 mol% SrCO 3 after only 2 min of contact, and 14–18 mol% SrCO 3 after 3 weeks of contact. The strontianite surface averages approximately 22 mol% CaCO 3 after 2 min of contact with Ca(HCO 3 ) 2 solution, and is 34–39 mol% CaCO 3 after 3 weeks of contact. XPS analysis suggests the surface composition is zoned with somewhat greater enrichment in the outer ~25 Å (as much as 26 mol% SrCO 3 on aragonite and 44 mol% CaCO 3 on strontianite). The results indicate rapid formation of a solid-solution surface phase from subsaturated aqueous solutions. The surface phase continually adjusts in composition in response to changes in composition of the bulk fluid as net dissolution proceeds. Dissolution rates of the endmembers are greatly reduced in nonstoichiometric solutions relative to dissolution rates observed in stoichiometric solutions. All solids dissolve more slowly in solutions spiked with the least soluble component ((Sr(HCO 3 ) 2 )) than in solutions spiked with the more soluble component (Ca(HCO 3 ) 2 ), an effect that becomes increasingly significant as stoichiometric saturation is approached. It is proposed that the formation of a non-stoichiometric surface reactive zone significantly decreases dissolution rates.

Geochimica et Cosmochimica Acta

Interparticle migration of metal cations in stream sediments as a factor in toxics transport

Sorption of metal cations by stream sediments is an important process affecting the movement of released contaminants in the environment. The ability of cations to desorb from one sediment particle and subsequently sorb to another can greatly affect metal transport rates but rates for this process have not been reported. The objective of this study was to determine the rate at which sorbed metals can migrate from contaminated sediment particles to uncontaminated sediment particles as a function of the concentration of the contaminating solution and the duration of the contact with the contaminating solution. Samples of small sediment particles were exposed to solutions containing cobalt, after which they were rinsed and combined with larger uncontaminated sediment particles in the presence of stream water. Initial concentrations of the contaminating solution ranged from 1ng/l to 1000mg/l and exposures to the contaminating solution ranged from 6h to 14 days. The rate of the migration increased with increasing concentrations in the contaminating solution and with decreasing times of exposure to the contaminating solution. Under the conditions of these experiments, the time required for the migration to reach equilibrium was on the order of months or longer. In separate experiments, the kinetics of adsorption and desorption of cobalt were measured as a function of concentration of the contaminating solution. The time required to reach adsorption equilibrium increased with increasing concentration in the contaminating solution. Times to sorption equilibrium were on the order of months. Desorption was much slower than adsorption and, together with intraparticle diffusion, probably controls the rate of migration from contaminated to uncontaminated sediment. The results of this study show that interparticle migration of metal cations can proceed at significant rates that are strongly influenced by the length of time that the metal has been in contact with the sediment. Copyright ?? 2001 Elsevier Science B.V.Sorption of metal cations by stream sediments is an important process affecting the movement of released contaminants in the environment. The ability of cations to desorb from one sediment particle and subsequently sorb to another can greatly affect metal transport rates but rates for this process have not been reported. The objective of this study was to determine the rate at which sorbed metals can migrate from contaminated sediment particles to uncontaminated sediment particles as a function of the concentration of the contaminating solution and the duration of the contact with the contaminating solution. Samples of small sediment particles were exposed to solutions containing cobalt, after which they were rinsed and combined with larger uncontaminated sediment particles in the presence of stream water. Initial concentrations of the contaminating solution ranged from 1 ng/l to 1000 mg/l and exposures to the contaminating solution ranged from 6 h to 14 days. The rate of the migration increased with increasing concentrations in the contaminating solution and with decreasing times of exposure to the contaminating solution. Under the conditions of these experiments, the time required for the migration to reach equilibrium was on the order of months or longer. In separate experiments, the kinetics of adsorption and desorption of cobalt were measured as a function of concentration of the contaminating solution. The time required to reach adsorption equilibrium increased with increasing concentration in the contaminating solution. Times to sorption equilibrium were on the order of months. Desorption was much slower than adsorption and, together with intraparticle diffusion, probably controls the rate of migration from contaminated to uncontaminated sediment. The results of this study show that interparticle migration of metal cations can proceed at significant rates that are strongly influenced by the length of time that the metal has

Journal of Hazardous Materials

Effects of light intensity and temperature on Cryptomonas ovata (Cryptophyceae) growth and nutrient uptake rates

Specific growth rate of Cryptomonas ovata var. palustris Pringsheim was measured in batch culture at 14 light-temperature combinations. Both the maximum growth rate (μ m ) and optimum light intensity (I opt ) fit an empirical function that increases exponentially with temperature up to an optimum (T opt ), then declines rapidly as temperature exceeds T opt . Incorporation of these functions into Steele's growth equation gives a good estimate of specific growth rate over a wide range of temperature and light intensity. Rates of phosphate, ammonium and nitrate uptake were measured separately at 16 combinations of irradiance and temperature and following a spike addition of all starved cells initially took up nutrient at a rapid rate. This transitory surge was followed by a period of steady, substrate-saturated uptake that persisted until external nutrient concentration fell. Substrate-saturated NO 3 − -uptake proceeded at very slow rates in the dark and was stimulated by both increased temperature and irradiance; NH 4 + -uptake apparently proceeded at a basal rate at 8 and l4 C and was also stimulated by increased temperature and irradiance. Rates of NH 4 − -uptake were much higher than NO 3 − -uptake at all light-temperature combinations. Below 20 C, PO 4 −3 -uptake was more rapid in dark than in light, but was light enhanced at 26 C.

Journal of Phycology

Rupture history of the 2008 M w 7.9 Wenchuan, China, earthquake: Evaluation of separate and joint inversions of geodetic, teleseismic, and strong-motion data

An extensive data set of teleseismic and strong-motion waveforms and geodetic offsets is used to study the rupture history of the 2008 Wenchuan, China, earthquake. A linear multiple-time-window approach is used to parameterize the rupture. Because of the complexity of the Wenchuan faulting, three separate planes are used to represent the rupturing surfaces. This earthquake clearly demonstrates the strengths and limitations of geodetic, teleseismic, and strong-motion data sets. Geodetic data (static offsets) are valuable for determining the distribution of shallower slip but are insensitive to deeper faulting and reveal nothing about the timing of slip. Teleseismic data in the distance range 30°–90° generally involve no modeling difficulties because of simple ray paths and can distinguish shallow from deep slip. Teleseismic data, however, cannot distinguish between different slip scenarios when multiple fault planes are involved because steep takeoff angles lead to ambiguity in timing. Local strong-motion data, on the other hand, are ideal for determining the direction of rupture from directivity but can easily be over modeled with inaccurate Green’s functions, leading to misinterpretation of the slip distribution. We show that all three data sets are required to give an accurate description of the Wenchuan rupture. The moment is estimated to be approximately 1.0 × 10 21 N · m with the slip characterized by multiple large patches with slips up to 10 m. Rupture initiates on the southern end of the Pengguan fault and proceeds unilaterally to the northeast. Upon reaching the cross-cutting Xiaoyudong fault, rupture of the adjacent Beichuan fault starts at this juncture and proceeds bilaterally to the northeast and southwest.

Wenchuan

Rupture history of the 2011 M 9 Tohoku Japan earthquake determined from strong‐motion and high‐rate GPS recordings: Subevents radiating energy in different frequency bands

Strong‐motion records from KiK‐net and K‐NET, along with 1 sample/s Global Positioning System (GPS) records from GEONET, were analyzed to determine the location, timing, and slip of subevents of the M 9 2011 Tohoku earthquake. Timing of arrivals on stations along the coast shows that the first subevent was located closer to the coast than subevent (2), which produced the largest slip. A waveform inversion of data from 0 to 0.2 Hz indicates that the first subevent primarily ruptured down‐dip and north of the hypocenter and had an M of 8.5. The areas of this subevent that generated the low (<0.2 Hz) and high (>0.2 Hz) frequency energy are located in the same vicinity. The inversion result for the second subevent ( M 9.0) has large slip on the shallow part of the fault with peak slip of about 65 m above about 25 km depth. This slip generated the tsunami. The preferred inversion has initiation of subevent 2 on the shallow portion of the fault so that rupture proceeded down‐dip and mainly to the south. Subevent 2 started about 35 s after subevent 1, which allows for the possibility of dynamic triggering from subevent 1. The slip model predicts displacements comparable to those found from ocean‐bottom transducers near the epicenter. At frequencies that most affect tall buildings (0.1–0.5 Hz), there is a strong pulse (subevent 3) in the strong‐motion records that arrives after the near‐field ramp from subevent 2. High‐frequency subevent 3 was located down‐dip and south of the high‐slip portion of subevent 2 and was initiated as rupture from subevent 2 proceeded down‐dip. The compact pulse for subevent 3 is modeled with an M 8.0 source in a 75 by 30 km area that ruptured down‐dip and to the south with a high slip velocity, indicating high stress drop.

Bulletin of the Seismological Society of America

Predicting vertically-nonsequential wetting patterns with a source-responsive model

Water infiltrating into soil of natural structure often causes wetting patterns that do not develop in an orderly sequence. Because traditional unsaturated flow models represent a water advance that proceeds sequentially, they fail to predict irregular development of water distribution. In the source-responsive model, a diffuse domain (D) represents flow within soil matrix material following traditional formulations, and a source-responsive domain (S), characterized in terms of the capacity for preferential flow and its degree of activation, represents preferential flow as it responds to changing water-source conditions. In this paper we assume water undergoing rapid source-responsive transport at any particular time is of negligibly small volume; it becomes sensible at the time and depth where domain transfer occurs. A first-order transfer term represents abstraction from the S to the D domain which renders the water sensible. In tests with lab and field data, for some cases the model shows good quantitative agreement, and in all cases it captures the characteristic patterns of wetting that proceed nonsequentially in the vertical direction. In these tests we determined the values of the essential characterizing functions by inverse modeling. These functions relate directly to observable soil characteristics, rendering them amenable to evaluation and improvement through hydropedologic development.

Vadose Zone Journal

Preliminary report upon a reconnaissance through southern and southeastern Nevada, made in 1869

By authority from headquarters Military Division of the Pacific, Lieut. George M. Wheeler, United States Engineers, will proceed with his civil assistants and three enlisted men to either Camps Halleck or Ruby, Nevada, and having been joined by Lieut. D.W. Lockwood, United States Engineers, now en route via Fort Churchill, will there organize a party, to consist of two-non-commissioned officers and twenty-three enlisted men, (cavalry, or infantry mounted,) such drivers, packers, and guides as may be required; equip them with the necessary, full, and complete outfit, as far as the resources of the posts will enable him so to do; after which he will proceed, via the White Pine district, to make a thorough and careful reconnaissance of the district of country to the south and east of White Pine, extending thereto from the White Pine or Grant district, of obtaining correct data for a military map of the country, and for the selection of the site or sites for such military post or posts to cover the mining country south and east of White Pine from hostile Indians, as may be required. Such explorations and examinations as may will be made in reference to the physical geography of the country, its resources in wood, water, agricultural or mineral productions. The character, habits, and numbers of Indian tribes, and their dispositions toward settlers and miners, will be subjects for investigations.

Nevada

Environmental Assessment for a Marine Geophysical Survey of Parts of the Arctic Ocean, August-September 2010

According to the United Nations Convention on the Law of the Sea (UNCLOS), individual nations? sovereign rights extend to 200 nautical miles (n.mi.) (370 km) offshore or to a maritime boundary in an area called the continental shelf. These rights include jurisdiction over all resources in the water column and on and beneath the seabed. Article 76 of UNCLOS also establishes the criteria to determine areas beyond the 200 n.mi. (370 km) limit that could be defined as ?extended continental shelf,? where a nation could extend its sovereign rights over the seafloor and sub-seafloor (As used in UNCLOS, ?continental shelf? refers to a legally defined region of the sea floor rather than a morphological shallow-water area adjacent to continents commonly used by geologists and hydrographers.). This jurisdiction provided in Article 76 includes resources on and below the seafloor but not in the water column. The United States has been acquiring data to determine the outer limits of its extended continental shelf in the Arctic and has a vested interest in declaring and receiving international recognition of the reach of its extended continental shelf. The U.S. collaborated with Canada in 2008 and 2009 on extended continental shelf studies in the Arctic Ocean. The U.S. Coast Guard (USCG) Cutter Healy worked with the Canadian Coast Guard ship Louis S. St. Laurent to map the continental shelf beyond 200 n.mi. (370 km) in the Arctic. Each icebreaking vessel contributed different capabilities in order to collect data needed by both nations more efficiently in order to save money, avoid redundancy, and foster cooperation. Generally, the Healy collects bathymetric (sea-floor topography) data and the Louis S. St. Laurent collects seismic reflection profile data. The vessels work in concert when ice conditions are heavy, with one vessel breaking ice for the ship collecting data. The Canadian Environmental Assessments for these projects are available on line at http://www.ceaa.gc.ca/052/details-eng.cfm?pid=38185 (2008) and http://www.ceaa.gc.ca/052/details-eng.cfm?pid=46518 (2009). The U.S. Geological Survey (USGS) and Geological Survey of Canada (GSC) are undertaking a similar partnership again for 2010 in a limited area of U.S. waters during the period between ~10 and 16 August. The survey vessels will then proceed to international or Canadian waters where surveying will proceed until ~3 September, when the two icebreakers will separate to conduct independent work. The survey area of the joint work will be bounded approximately by 145? to 158? W longitude and 71? to 84? N latitude in water depths ranging from ~2,000 to 4,000 m (fig. 1). Ice conditions are expected to range from open water to 10/10 ice cover. The Louis S. St. Laurent will join accompanying vessel Healy in or near the survey area around 10 August to begin the joint survey work. As its energy source, the seismic system aboard Louis S. St. Laurent will employ a 3-airgun array consisting of three Sercel G-airguns. Two guns will have a discharge volume of 500 in3 and the third a discharge volume of 150 in3 for a total array discharge volume of 1,150 in3. The seismic survey will take place in water depths 2,000?4,000 m. This airgun array is identical to the system used in the 2008 and 2009 field programs by the Geological Survey of Canada. The USGS requested that the National Marine Fisheries Service (NMFS) issue an Incidental Harassment Authorization (IHA) to authorize the incidental, that is, not intentional, harassment of small numbers of cetaceans and seals should this occur during the seismic survey in U.S. waters. USGS is also consulting with the U.S. Fish and Wildlife Service (USFWS) regarding concerns about disturbance to walruses and polar bears. Through informal consultation with the Office of Protected Resources with the National Oceanic and Atmospheric Administration (NOAA), USGS proposes that no ESA-listed marine species?bowhead, fin, humpback or sperm whale?w

Open-File Report

Case studies of riparian and watershed restoration in the southwestern United States—Principles, challenges, and successes

Globally, rivers and streams are highly altered by impoundments, diversions, and stream channelization associated with agricultural and water delivery needs. Climate change imposes additional challenges by further reducing discharge, introducing variability in seasonal precipitation patterns, and increasing temperatures. Collectively, these changes in a river or stream’s annual hydrology affects surface and groundwater dynamics, fluvial processes, and the linked aquatic and riparian responses, particularly in arid regions. Recognizing the inherent ecosystem services that riparian and aquatic habitats provide, society increasingly supports restoring the functionality of riparian and aquatic ecosystems. Given the wide range in types and scales of riparian impacts, approaches to riparian restoration can range from tactical, short-term, and site-specific efforts to strategic projects and long-term collaborations best pursued at the watershed scale. In the spirit of sharing information, the U.S. Geological Survey’s Grand Canyon Monitoring and Research Center convened a workshop June 23-25, 2015, in Flagstaff, Ariz. for practitioners in restoration science to share general principles, successful restoration practices, and discuss the challenges that face those practicing riparian restoration in the southwestern United States. Presenters from the Colorado River and the Rio Grande basins, offered their perspectives and experiences in restoration at the local, reach and watershed scale. Outcomes of the workshop include this Proceedings volume, which is composed of extended abstracts of most of the presentations given at the workshop, and recommendations or information needs identified by participants. The organization of the Proceedings follows a general progression from local scale restoration to river and watershed scale approaches, and finishes with restoration assessments and monitoring.

Open-File Report

Seismic hazard study of the western portion of the Garlock fault [California]

Investigations of the western segment of the Garlock fault were conducted at Castac Lake, Twin Lakes and Oak Creek Canyon. Studies were concentrated on the youngest fault trace as delineated by Clark (1973). Seismic refraction surveys, topographic surveys and geologic mapping provided positive evidence for fault offsets in Quaternary deposits at Twin Lakes and Oak Creek Canyon. Investigations at Castac Lake were concentrated on the fault segment north of the lake. Previous investigations south of the lake and across the dry lake bottom exposed unbroken strata dating back 8050 +300 years B.P. No evidence of offset of young deposits was found north of the lake from surface investigations or seismic refraction data. Trenching of sag pond deposits at Twin Lakes revealed fault traces representing 2 events confined to a zone 3 meters wide. The stratigraphy at Twin Lakes consists of alternating units of clay and sand allowing measurements of vertical offsets. Based on radiocarbon dates of detrital charcoal, the most recent movement occurred less than 890 +195 years B.P. and the proceeding event occurred greater than 2,800 +165 years B.P. Total vertical offset was 80 to 100 cm for the most recent movement and 55-60 cm for the proceeding event. Geomorphic analysis of offset alluvial stream channels at Oak Creek Canyon provided an estimate of 1.6 to 3.3 mm per year of left lateral movement during Holocene and late Pleistocene times. This estimate is based on correlation of soil profile development with dated soil stratigraphy in the San Joaquin Valley. Scarp heights and lack of definitive evidence for recent left-lateral offset in Oak Creek Canyon indicates a significant and perhaps predominant vertical component of movement with: 1) a comparatively long recurrence interval and/or 2) a small component of left-lateral movement. Arching of late Pleistocene deposits in Oak Creek Canyon is consistent with right-lateral rather than left-lateral wrench faulting, suggesting a fluctuation in stresses along this part of the fault since late Pleistocene time.

Open-File Report

Book review of Wildlife 2000: Modeling relationships of terrestrial vertebrates, edited by J. Verner, M.L. Morrison, and C.J. Ralph

"Wildlife 2000" is the proceedings of a conference held 7-11 October 1984, near Lake Tahoe, California, the objective of which was to present an up-to-date synthesis of models that predict the responses of wildlife to habitat change. This extremely attractive, well-produced volume has been well received by the wildlife management profession; the editors received an outstanding publication award from The Wildlife Society for this publication. The accolades are deserved. The symposium was purposely integrated in terms of research and management perspectives. Each of the six sections is summarized by both research and management points of view. A majority of the 60 papers presented deal with birds. Although many chapters require a strong quantitative background, especially in multivariate statistics, many others do not. When one compares this publication with previous habitat-modeling symposia proceedings, one realizes what a superior contribution "Wildlife 2000" is, and how incredible far wildlife-habitat modelers have come in a short time. There are very few redundant papers of "nonpapers" in this volume. The wide array of modeling procedures, statistical methods, and computer software developed and used by the authors is impressive; we have indeed learned how to build models. Whether or not we have learned how to build good models is another question.

The Wilson Bulletin

Opinion: A preferred approach for dealing with reproducibility and replicability in science

Science impacts our daily lives and guides national and international policies (1). Thus, results of scientific studies are of paramount importance; yet, there are concerns that many studies are not reproducible or replicable (2). To address these concerns, the National Research Council conducted a Consensus Study [NASEM 2019 (3)] that provides definitions of key concepts, discussions of problems, and recommendations for dealing with these problems. These recommendations are useful and well considered, but they do not go far enough in our opinion. The NASEM recommendations treat reproducibility and replicability as single-study issues, despite clear acknowledgement of the limitations of isolated studies and the need for research synthesis (3). We advocate a strategic approach to research, focusing on the accumulation of evidence via designed sequences of studies, as a means of dealing more effectively with reproducibility, replicability, and related problems. These sequences are designed to provide iterative tests based on comparison of data from empirical studies with predictions from competing hypotheses. Evidence is then formally accumulated based on the relative predictive abilities of the different hypotheses as the sequential studies proceed.

Proceedings of the National Academy of Sciences of

Frequency-dependent effects of rupture for the 2004 Parkfield mainshock, results from UPSAR

The frequency-dependent effects of rupture propagation of the Parkfield, California earthquake (Sept. 28, 2004, M6) to the northwest along the San Andreas fault can be seen in acceleration records at UPSAR (USGS Parkfield Seismic Array) in at least two ways. First, we can see the effects of directivity in the acceleration traces at UPSAR, which is about 11.5 km from the epicenter. Directivity or the seismic equivalent of a Doppler shift has been documented in many cases by comparing short duration, high-amplitude pulses (P or S) in the forward direction with longer duration body waves in the backward direction. In this case we detect a change from a relatively large amplitude, coherent, high-frequency signal at the start of rupture to a low-amplitude, low-coherent, low-frequency signal at about the time the rupture front transfers from the forward azimuth to the back azimuth at about 34-36 s (time is UTC and are the seconds after day 272 and 17 hours and 15 minutes. S arrival is just after 30s) for rays leaving the fault and propagating to UPSAR. The frequency change is obvious in the band about 5 to 30 Hz, which is significantly above the corner frequency of the earthquake (about 0.11Hz). From kinematic source models, the duration of faulting is about 9.2 s and the change in frequency is during faulting as the rupture extends to the northwest. Understanding the systematic change in frequency and amplitude of seismic waves in relation to the propagation of the rupture front is important for predicting strong ground motion. Second, we can filter the acceleration records from the array to determine if the low frequency energy emerges from the same part of the fault as the high frequency signal (e.g. has the same back azimuth and apparent velocity at UPSAR) an important clue to the dynamics of rupture. Analysis of sources of strong motion (characterized by relatively high frequencies) compared to kinematic slip models (relatively low frequency) for the March 11, 2011 Tohoku earthquake as well as Maule (Feb. 27, 2010) and Chi-Chi (Sept. 20, 1999) earthquakes show that high- and low-frequency sources do not have the same locations on the fault. In this paper we filter the accelerograms from UPSAR for the 2004 mainshock in various passbands and then re-compute the cross correlations to determine the vector slowness of the incoming waves. At Parkfield, it appears that for seismic waves with frequencies above 1 Hz there is no discernible frequency-dependent difference in source position (up to 8 Hz) based on estimates of back azimuth and apparent velocity. However at lower frequencies, sources appear to be from shallower depths and trail the high frequencies as the rupture proceeds down the fault. This result is greater than one standard deviation of an estimate of error, based on a new method of estimating error that is a measure of how broad the peak in correlation is and an estimate of the variance of the correlation values. These observations can be understood in terms of a rupture front that is more energetic and coherent near the front of rupture (radiating higher frequencies) and less coherent and less energetic (radiating in a lower frequency band) behind the initial rupture front. This result is a qualitative assessment of changes in azimuth and apparent velocity with frequency and time and does not include corrections to find the source location on the fault.

California

Episodic Holocene eruption of the Salton Buttes rhyolites, California, from paleomagnetic, U-Th, and Ar/Ar dating

In the Salton Trough, CA, five rhyolite domes form the Salton Buttes: Mullet Island, Obsidian Butte, Rock Hill, North and South Red Hill, from oldest to youngest. Results presented here include 40 Ar/ 39 Ar anorthoclase ages, 238 U- 230 Th zircon crystallization ages, and comparison of remanent paleomagnetic directions with the secular variation curve, which indicate that all domes are Holocene. 238 U- 230 Th zircon crystallization ages are more precise than but within uncertainty of 40 Ar/ 39 Ar anorthoclase ages, suggesting that zircon crystallization proceeded until shortly before eruption in all cases except one. Remanent paleomagnetic directions require three eruption periods: (1) Mullet Island, (2) Obsidian Butte, and (3) Rock Hill, North Red Hill, and South Red Hill. Borehole cuttings logs document up to two shallow tephra layers. North and South Red Hills likely erupted within 100 years of each other, with a combined 238 U- 230 Th zircon isochron age of: 2.83 &plusmn; 0.60 ka (2 sigma); paleomagnetic evidence suggests this age predates eruption by hundreds of years (1800 cal BP). Rock Hill erupted closely in time to these eruptions. The Obsidian Butte 238 U- 230 Th isochron age (2.86 &plusmn; 0.96 ka) is nearly identical to the combined Red Hill age, but its Virtual Geomagnetic Pole position suggests a slightly older age. The age of aphyric Mullet Island dome is the least well constrained: zircon crystals are resorbed and the paleomagnetic direction is most distinct; possible Mullet Island ages include ca. 2300, 5900, 6900, and 7700 cal BP. Our results constrain the duration of Salton Buttes volcanism to between ca. 5900 and 500 years.

Geochemistry, Geophysics, Geosystems

Occupancy modeling species–environment relationships with non‐ignorable survey designs

Statistical models supporting inferences about species occurrence patterns in relation to environmental gradients are fundamental to ecology and conservation biology. A common implicit assumption is that the sampling design is ignorable and does not need to be formally accounted for in analyses. The analyst assumes data are representative of the desired population and statistical modeling proceeds. However, if data sets from probability and non‐probability surveys are combined or unequal selection probabilities are used, the design may be non‐ignorable. We outline the use of pseudo‐maximum likelihood estimation for site‐occupancy models to account for such non‐ignorable survey designs. This estimation method accounts for the survey design by properly weighting the pseudo‐likelihood equation. In our empirical example, legacy and newer randomly selected locations were surveyed for bats to bridge a historic statewide effort with an ongoing nationwide program. We provide a worked example using bat acoustic detection/non‐detection data and show how analysts can diagnose whether their design is ignorable. Using simulations we assessed whether our approach is viable for modeling data sets composed of sites contributed outside of a probability design. Pseudo‐maximum likelihood estimates differed from the usual maximum likelihood occupancy estimates for some bat species. Using simulations we show the maximum likelihood estimator of species–environment relationships with non‐ignorable sampling designs was biased, whereas the pseudo‐likelihood estimator was design unbiased. However, in our simulation study the designs composed of a large proportion of legacy or non‐probability sites resulted in estimation issues for standard errors. These issues were likely a result of highly variable weights confounded by small sample sizes (5% or 10% sampling intensity and four revisits). Aggregating data sets from multiple sources logically supports larger sample sizes and potentially increases spatial extents for statistical inferences. Our results suggest that ignoring the mechanism for how locations were selected for data collection (e.g., the sampling design) could result in erroneous model‐based conclusions. Therefore, in order to ensure robust and defensible recommendations for evidence‐based conservation decision‐making, the survey design information in addition to the data themselves must be available for analysts. Details for constructing the weights used in estimation and code for implementation are provided.

Ecological Applications

Considerations for maximizing the adaptive potential of restored coral populations in the western Atlantic

Active coral restoration typically involves two interventions: crossing gametes to facilitate sexual larval propagation; and fragmenting, growing, and outplanting adult colonies to enhance asexual propagation. From an evolutionary perspective, the goal of these efforts is to establish self‐sustaining, sexually reproducing coral populations that have sufficient genetic and phenotypic variation to adapt to changing environments. Here, we provide concrete guidelines to help restoration practitioners meet this goal for most Caribbean species of interest. To enable the persistence of coral populations exposed to severe selection pressure from many stressors, a mixed provenance strategy is suggested: genetically unique colonies (genets) should be sourced both locally as well as from more distant, environmentally distinct sites. Sourcing three to four genets per reef along environmental gradients should be sufficient to capture a majority of intraspecies genetic diversity. It is best for practitioners to propagate genets with one or more phenotypic traits that are predicted to be valuable in the future, such as low partial mortality, high wound healing rate, high skeletal growth rate, bleaching resilience, infectious disease resilience, and high sexual reproductive output. Some effort should also be reserved for underperforming genets because colonies that grow poorly in nurseries sometimes thrive once returned to the reef and may harbor genetic variants with as yet unrecognized value. Outplants should be clustered in groups of four to six genets to enable successful fertilization upon maturation. Current evidence indicates that translocating genets among distant reefs is unlikely to be problematic from a population genetic perspective but will likely provide substantial adaptive benefits. Similarly, inbreeding depression is not a concern given that current practices only raise first‐generation offspring. Thus, proceeding with the proposed management strategies even in the absence of a detailed population genetic analysis of the focal species at sites targeted for restoration is the best course of action. These basic guidelines should help maximize the adaptive potential of reef‐building corals facing a rapidly changing environment.

Ecological Applications