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At least 307 records · Page 17Linked to original sources

Mantle peridotite in newly discovered far-inland subduction complex, southwest Arizona: Initial report

The latest Cretaceous to early Palaeogene Orocopia Schist and related units are generally considered a low-angle subduction complex that underlies much of southern California and Arizona. A recently discovered exposure of Orocopia Schist at Cemetery Ridge west of Phoenix, Arizona, lies exceptionally far inland from the continental margin. Unexpectedly, this body of Orocopia Schist contains numerous blocks, as large as ~300 m, of variably serpentinized mantle peridotite. These are unique; elsewhere in the Orocopia and related schists, peridotite is rare and completely serpentinized. Peridotite and metaperidotite at Cemetery Ridge are of three principal types: (1) serpentinite and tremolite serpentinite, derived from dunite; (2) partially serpentinized harzburgite and olivine orthopyroxenite (collectively, harzburgite); and (3) granoblastic or schistose metasomatic rocks, derived from serpentinite, made largely of actinolite, calcic plagioclase, hercynite, and chlorite. In the serpentinite, paucity of relict olivine, relatively abundant magnetite (5%), and elevated Fe 3 + /Fe indicate advanced serpentinization. Harzburgite contains abundant orthopyroxene, only slightly serpentinized, and minor to moderate (1–15%) relict olivine. Mantle tectonite fabric is locally preserved. Several petrographic and geochemical characteristics of the peridotite at Cemetery Ridge are ambiguously similar to either abyssal or mantle-wedge (suprasubduction) peridotites and serpentinites. Least ambiguous are orthopyroxene compositions. Orthopyroxene is distinctively depleted in Al 2 O 3 , Cr 2 O 3 , and CaO, indicating mantle-wedge affinities. Initial interpretation of field and petrologic data suggests that the peridotite blocks in the Orocopia Schist subduction complex at Cemetery Ridge may be derived from the leading corner or edge of a mantle wedge, presumably in (pre-San Andreas fault) southwest California. However, derivation from a subducting plate is not precluded.

Arizona

Simulating the water budget of a Prairie Potholes complex from LiDAR and hydrological models in North Dakota, USA

Hydrological processes of the wetland complex in the Prairie Pothole Region (PPR) are difficult to model, partly due to a lack of wetland morphology data. We used Light Detection And Ranging (LiDAR) data sets to derive wetland features; we then modelled rainfall, snowfall, snowmelt, runoff, evaporation, the “fill-and-spill” mechanism, shallow groundwater loss, and the effect of wet and dry conditions. For large wetlands with a volume greater than thousands of cubic metres (e.g. about 3000 m 3 ), the modelled water volume agreed fairly well with observations; however, it did not succeed for small wetlands (e.g. volume less than 450 m 3 ). Despite the failure for small wetlands, the modelled water area of the wetland complex coincided well with interpretation of aerial photographs, showing a linear regression with R 2 of around 0.80 and a mean average error of around 0.55 km 2 . The next step is to improve the water budget modelling for small wetlands.

North Dakota

Characterization of branch complexity by fractal analyses and detect plant functional adaptations

The comparison between complexity in the sense of space occupancy (box-counting fractal dimension Dc and information dimension DI ) and heterogeneity in the sense of space distribution (average evenness index and evenness variation coefficient JCV) were investigated in mathematical fractal objects and natural branch ¯ J structures. In general, increased fractal dimension was paired with low heterogeneity. Comparisons between branch architecture in Anthyllis cytisoides under different slope exposure and grazing impact revealed that branches were more complex and more homogeneously distributed for plants on northern exposures than southern, while grazing had no impact during a wet year. Developmental instability was also investigated by the statistical noise of the allometric relation between internode length and node order. In conclusion, our study demonstrated that fractal dimension of branch structure can be used to analyze the structural organization of plants, especially if we consider not only fractal dimension but also shoot distribution within the canopy (lacunarity). These indexes together with developmental instability analyses are good indicators of growth responses to the environment.

International Journal of Plant Sciences

Tradeoffs with growth limit host range in complex life-cycle helminths

Parasitic worms with complex life cycles have several developmental stages, with each stage creating opportunities to infect additional host species. Using a dataset for 973 species of trophically transmitted acanthocephalans, cestodes, and nematodes, we confirmed that worms with longer life cycles (i.e. more successive hosts) infect a greater diversity of host species and taxa (after controlling for study effort). Generalism at the stage level was highest for ‘middle’ life stages, the second and third intermediate hosts of long life cycles. By simulating life cycles in real food webs, we found that middle stages had more potential host species to infect, suggesting that opportunity constrains generalism. However, parasites usually infected fewer host species than expected from simulated cycles, suggesting generalism also has costs. There was no tradeoff in generalism from one stage to the next, but worms spent less time growing and developing in stages where they infected more taxonomically diverse hosts. Our results demonstrate that life cycle complexity favors high generalism, and host use across life stages is determined by both ecological opportunity and life history tradeoffs.

The American Naturalist

Speciation genomics in the tiger whiptail lizards ( Aspidoscelis tigris complex)

The transition from small genetic to genome-scale datasets for studying biodiversity has revealed that genetic exchange through introgressive hybridization is a widespread phenomenon in nature. Despite this, a lack of high-quality reference genomes for most non-model species limits our understanding of the impact of this process for many taxonomic groups. This restricts the range of insights that genomic tools can provide for conservation biologists, who often hope to employ genomic datasets to accurately identify historically isolated lineages to protect and to predict their evolutionary fate in the face of environmental change. Tiger whiptail lizards ( Aspidoscelis tigris complex) are an abundant and important ecological component of ecosystems across the southwestern United States. In this study, we assembled and annotated a chromosome-level reference genome for A. t. stejnegeri from coastal California. We then used this reference genome to reconstruct patterns of speciation and admixture within the larger species complex, finding evidence that gene flow is widespread both geographically and across the genome.

northern Mexico, southwestern United States

Stress- and structure-controlled anisotropy in a region of complex faulting—Yuha Desert, California

We examine shear velocity anisotropy in the Yuha Desert, California using aftershocks of the 2010 M7.2 El Mayor-Cucapah earthquake. The Yuha Desert is underlain by a complex network of right- and left-lateral conjugate faults, some of which experienced triggered slip during the El Mayor-Cucapah earthquake. An automated method that implements multiple measurement windows and a range of bandpass filters is used to estimate the fast direction ( ϕ ) and delay time ( δt ) of the split shear waves. We find an average ϕ oriented approximately north–south suggesting it is primarily controlled by the regional maximum compressive stress direction. However, the spatial variability in ϕ reveals that the fault structures that underlie the Yuha Desert also influence the measured splitting parameters. We infer that the northeast- and northwest-oriented ϕ reflect shear fabric subparallel to the conjugate fault structures. We do not observe a simple correlation between δt and hypocentral distance. Instead, the observed spatial variation in δt suggests that near-source variation in anisotropic strength may be equal to or more important than effects local to the station. No temporal variation in splitting parameters is observed during the 70-day period following the main shock. In this region of complex faulting, we observe a spatially variable pattern of anisotropy that is both stress- and structure-controlled. This study suggests that shear fabric can form even along short, discontinuous fault strands with minimal offset.

California

Rare earth element evidence for the petrogenesis of the banded series of the Stillwater Complex, Montana, and its anorthosites

A rare earth element (REE) study was made by isotope-dilution mass spectrometry of plagioclase separates from a variety of cumulates stratigraphically spanning the Banded series of the Stillwater Complex, Montana. Evaluation of parent liquid REE patterns, calculated on the basis of published plagioclase-liquid partition coefficients, shows that the range of REE ratios is too large to be attributable to fractionation of a single magma type. At least two different parental melts were present throughout the Banded series. This finding supports hypotheses of previous workers that the Stillwater Complex formed from two different parent magma types, designated the anorthositic- or A-type liquid and the ultramafic- or U-type liquid. On the basis of our data, one melt has a REE pattern with a distinctive shallow slope and is represented by samples from the thick, massive Anorthosite zones I and II (AN I and AN II) of the Middle Banded series. Although samples from AN I and AN II are separated by as much as 1400 m stratigraphically, they have remarkably similar calculated parent liquid characteristics, with (Ce/Sm)n = 1.7–1.9, (Nd/Sm)n = 1.3–1.4 and (Ce/Yb)n = 2.9–4.6 (where n denotes chondrite-normalized). These calculated liquids are probably close to representing A-type magma. In addition, plagioclase-bronzite cumulates from Norite zones I and II (N I and N II), although thought to be U-type cumulates, contain plagioclase that has A-type REE characteristics, implying that A-type magmas were injected into the magma chamber during formation of those zones. In contrast, calculated parent liquids of cumulus augite-bearing rocks have REE patterns that display distinctly steeper slopes than the A-type REE pattern. The extreme is the calculated parent liquid of a plagioclase-bronzite-augite cumulated with (Ce/Sm)n = 2.9, (Nd/Sm)n = 1.7, and (Ce/Yb)n = 10.1.

Journal of Petrology

Fish composition in a complex freshwater estuary: Environmental DNA metabarcoding versus capture surveys

Objective The potential for environmental DNA (eDNA) to disperse widely from source organisms enables high detection efficiency but raises questions about eDNA's ability to differentiate fine-scale spatial patterns relative to conventional fish capture data. Methods We evaluate these questions in the St. Louis River estuary—a hydrologically and spatially complex coastal system within Lake Superior that supports a diverse assemblage of resident and migratory fish species—via comparison of eDNA metabarcoding (12S and 16S loci) to multigear capture survey data from 2 years and two seasons. Results The eDNA and capture surveys collectively yielded 68 fish species: 2 species detected only by capture, 27 detected only by eDNA, and 39 shared across both survey types but having generally higher occurrence frequencies with eDNA than capture. Six species detected only by eDNA were unexpected, having no prior records in the Lake Superior basin. Data from paired eDNA and capture stations showed little relationship between the two survey types, with capture yielding species at stations that eDNA did not, eDNA detecting species in different habitats and distant locations from any captures, and assemblage patterns homogenized in eDNA surveys relative to capture surveys. Conclusions Our study finds that eDNA is a sensitive tool for assessing species presence at the system scale but that capture surveys may better yield the fine-scale spatial distribution information of interest to fisheries and habitat managers, especially in spatially and hydrologically complex systems.

Transactions of the American Fisheries Society

How to predict community responses to perturbations in the face of imperfect knowledge and network complexity

Recent attempts to predict the response of large food webs to perturbations have revealed that in larger systems increasingly precise information on the elements of the system is required. Thus, the effort needed for good predictions grows quickly with the system's complexity. Here, we show that not all elements need to be measured equally well, suggesting that a more efficient allocation of effort is possible. We develop an iterative technique for determining an efficient measurement strategy. In model food webs, we find that it is most important to precisely measure the mortality and predation rates of long-lived, generalist, top predators. Prioritizing the study of such species will make it easier to understand the response of complex food webs to perturbations.

Proceedings of the Royal Society B: Biological Sci

Comparative phylogenomics reveal complex evolution of life history strategies in a clade of bivalves with parasitic larvae (Bivalvia: Unionoida: Ambleminae)

Freshwater mussels are a species‐rich group with biodiversity patterns strongly shaped by a life history strategy that includes an obligate parasitic larval stage. In this study, we set out to reconstruct the life history evolution and systematics in a clade of freshwater mussels adapted to parasitizing a molluscivorous host fish. Anchored hybrid enrichment and ancestral character reconstruction revealed a complex pattern of life history evolution with host switching and multiple instances of convergence, including reduction in size of larvae, increased fecundity, and growth during encapsulation. Our phylogenomic analyses also recovered non‐monophyly of taxa exhibiting multiple traits used as the basis for previous taxonomic hypotheses. Taxa with axe‐head shaped glochidia were resolved as paraphyletic, but our results strongly suggest the complex morphology is an adaptation to reduce larval size, with reduction in size further accentuated in taxa previously assigned to Leptodea . To more accurately reflect the evolutionary history of this group, we make multiple systematic changes, including the description of a new genus, Atlanticoncha gen. nov. , and the synonymy of the genus Leptodea under Potamilus . Our findings contribute to the growing body of literature showing that cladistic hypotheses based solely on morphological characters, including larval morphology, can be flawed in freshwater mussels.

Cladistics

The complimentary role of lentic and lotic habitats for Arctic grayling in a complex stream-lake network in Arctic Alaska

Lakes can be important to stream dwelling fishes, yet how individuals exploit habitat heterogeneity across complex stream-lake networks is poorly understood. Furthermore, despite growing awareness that intermittent streams are widely used by fish, studies documenting use of seasonally accessible lakes remain scarce. We studied Arctic grayling (Thymallus arcticus) in a small seasonally flowing (June – October) stream-lake network in Alaska using PIT telemetry. Overall, 70% of fish visited two lakes, 8% used a single lake, and 22% used only stream reaches. We identified four distinct behavioral patterns that differed in dominant macrohabitat used (deep-lake, shallow-lake, or stream reaches), entry-day into the network, and mobility. Some juvenile fish spent the entire summer in a shallow seasonally frozen lake (average 71 days), whereas others demonstrated prospecting behavior and only entered the stream channel briefly in September. Another group included adult and juvenile fish that were highly mobile, moving up to 27 km while in the 3-km stream-lake network, and used stream reaches extensively (average 59 days). Lentic and lotic habitats served differing roles for individuals, some fish occupied stream reaches as summer foraging habitat and other individuals used them as migration corridors to access lakes. Our study emphasizes the importance of considering stream-lake connectivity in stream fish assessments, even to shallow seasonally frozen habitats not widely recognized as important. Furthermore, we demonstrate that individuals may use temporary aquatic habitats in complex and changing ways across ontogeny that are not captured by typical classifications of fish movement behavior.

Alaska

Evidence from lava flows for complex polarity transitions: The new composite Steens Mountain reversal record

Geomagnetic polarity transitions may be significantly more complex than are currently depicted in many sedimentary and lava-flow records. By splicing together paleomagnetic results from earlier studies at Steens Mountain with those from three newly studied sections of Oregon Plateau flood basalts at Catlow Peak and Poker Jim Ridge 70–90 km to the southeast and west, respectively, we provide support for this interpretation with the most detailed account of a magnetic field reversal yet observed in volcanic rocks. Forty-five new distinguishable transitional (T) directions together with 30 earlier ones reveal a much more complex and detailed record of the 16.7 Ma reversed (R)-to-normal (N) polarity transition that marks the end of Chron C5Cr. Compared to the earlier R-T-N-T-N reversal record, the new record can be described as R-T-N-T-N-T-R-T-N. The composite record confirms earlier features, adds new west and up directions and an entire large N-T-R-T segment to the path, and fills in directions on the path between earlier directional jumps. Persistent virtual geomagnetic pole (VGP) clusters and separate VGPs have a preference for previously described longitudinal bands from transition study compilations, which suggests the presence of features at the core–mantle boundary that influence the flow of core fluid and distribution of magnetic flux. Overall the record is consistent with the generalization that VGP paths vary greatly from reversal to reversal and depend on the location of the observer. Rates of secular variation confirm that the flows comprising these sections were erupted rapidly, with maximum rates estimated to be 85–120 m ka −1 at Catlow and 130–195 m ka −1 at Poker Jim South. Paleomagnetic poles from other studies are combined with 32 non-transitional poles found here to give a clockwise rotation of the Oregon Plateau of 11.4°± 5.6° with respect to the younger Columbia River Basalt Group flows to the north and 14.5°± 4.6° with respect to cratonic North America (95 per cent confidence interval).

Geophysical Journal International

Refugial isolation and divergence in the Narrowheaded Gartersnake species complex (Thamnophis rufipunctatus) as revealed by multilocus DNA sequence data

Glacial–interglacial cycles of the Pleistocene are hypothesized as one of the foremost contributors to biological diversification. This is especially true for cold‐adapted montane species, where range shifts have had a pronounced effect on population‐level divergence. Gartersnakes of the Thamnophis rufipunctatus species complex are restricted to cold headwater streams in the highlands of the Sierra Madre Occidental and southwestern USA. We used coalescent and multilocus phylogenetic approaches to test whether genetic diversification of this montane‐restricted species complex is consistent with two prevailing models of range fluctuation for species affected by Pleistocene climate changes. Our concatenated nuDNA and multilocus species analyses recovered evidence for the persistence of multiple lineages that are restricted geographically, despite a mtDNA signature consistent with either more recent connectivity (and introgression) or recent expansion (and incomplete lineage sorting). Divergence times estimated using a relaxed molecular clock and fossil calibrations fall within the Late Pleistocene, and zero gene flow scenarios among current geographically isolated lineages could not be rejected. These results suggest that increased climate shifts in the Late Pleistocene have driven diversification and current range retraction patterns and that the differences between markers reflect the stochasticity of gene lineages (i.e. ancestral polymorphism) rather than gene flow and introgression. These results have important implications for the conservation of T. rufipunctatus (sensu novo), which is restricted to two drainage systems in the southwestern US and has undergone a recent and dramatic decline.

Molecular Ecology

Constraints on the depth of generation and emplacement of a magmatic epidote-bearing quartz diorite pluton in the Coast Plutonic Complex, British Columbia

Petrology and P–T estimates indicate that a magmatic epidote-bearing quartz diorite pluton from Mt. Gamsby, Coast Plutonic Complex, British Columbia, was sourced at pressures below ∼1.4 GPa and cooled nearly isobarically at ∼0.9 GPa. The P–T path indicates that the magma was within the stability field of magmatic epidote early and remained there upon final crystallization. The pluton formed and crystallized at depths greater than ∼30 km. REE data indicate that garnet was absent in the melting region and did not fractionate during crystallization. This suggests that the crust was less than or equal to ∼55 km thick at 188 Ma during the early phases of magmatism in the Coast Plutonic Complex. Late Cretaceous contractional deformation and early Tertiary extension exhumed the rocks to upper crustal levels. Textures of magmatic epidote and other magmatic phases, combined with REE data, can be important for constraining the P–T path followed by magmas.

British Columbia

Stage structure alters how complexity affects stability of ecological networks

Resolving how complexity affects stability of natural communities is of key importance for predicting the consequences of biodiversity loss. Central to previous stability analysis has been the assumption that the resources of a consumer are substitutable. However, during their development, most species change diets; for instance, adults often use different resources than larvae or juveniles. Here, we show that such ontogenetic niche shifts are common in real ecological networks and that consideration of these shifts can alter which species are predicted to be at risk of extinction. Furthermore, niche shifts reduce and can even reverse the otherwise stabilizing effect of complexity. This pattern arises because species with several specialized life stages appear to be generalists at the species level but act as sequential specialists that are hypersensitive to resource loss. These results suggest that natural communities are more vulnerable to biodiversity loss than indicated by previous analyses.

Ecology Letters

Hydrostratigraphic modeling of a complex, glacial-drift aquifer system for importation into MODFLOW

Deposition from at least three episodes of glaciation left a complex glacial-drift aquifer system in central Illinois. The deepest and largest of these aquifers, the Sankoty-Mahomet Aquifer, occupies the lower part of a buried bedrock valley and supplies water to communities throughout central Illinois. Thin, discontinuous aquifers are present within glacial drift overlying the Sankoty-Mahomet Aquifer. This study was commissioned by local governments to identify possible areas where a regional water supply could be obtained from the aquifer with minimal adverse impacts on existing users. Geologic information from more than 2200 existing water well logs was supplemented with new data from 28 test borings, water level measurements in 430 wells, and 35 km of surface geophysical profiles. A three-dimensional (3-D) hydrostratigraphic model was developed using a contouring software package, a geographic information system (GIS), and the 3-D geologic modeling package, EarthVision??. The hydrostratigraphy of the glacial-drift sequence was depicted as seven uneven and discontinuous layers, which could be viewed from an infinite number of horizontal and vertical slices and as solid models of any layer. Several iterations were required before the 3-D model presented a reasonable depiction of the aquifer system. Layers from the resultant hydrostratigraphic model were imported into MODFLOW, where they were modified into continuous layers. This approach of developing a 3-D hydrostratigraphic model can be applied to other areas where complex aquifer systems are to be modeled and is also useful in helping lay audiences visualize aquifer systems.

Ground Water

Gulf Coast vicariance shapes phylogeographic history of a North American freshwater mussel species complex

Aim Freshwater mussels share habitat and are parasites of freshwater fishes during the larval life stage. Therefore, models of fish biogeography may also explain the historical biogeography of freshwater mussels. We tested this assumption using predictions of three biogeographic models constructed for northern Gulf of Mexico drainages on a freshwater mussel species complex. Specifically, we tested (1) if speciation was due to vicariant events of fluctuating sea levels that separated lineages east‐west of the Mobile Basin (Central Gulf Coast speciation hypothesis), (2) if the timing of divergences occurred 8.5–3.5 MYA (Gulf Coast allopatric speciation model) and (3) if diversification in Mississippi River populations was recent and for evidence of population increase consistent with range expansion into northern deglaciated regions (Pleistocene glaciation model). Location Eastern North America. Taxon Freshwater mussels (Bivalvia: Unionidae), Lampsilis teres and L. floridensis . Methods We collected 249 specimens from 73 localities across the group's distribution. We used three molecular markers (COI, NDI & ITSI) to conduct time calibrated Bayesian phylogenetic analyses, phylogeographic analyses (AMOVA & SAMOVA) and demographic analyses including Bayesian skyline plots. Results Lampsilis teres and L. floridensis are allopatric species whose distributions meet at the eastern edge of the Mobile Basin. Speciation was estimated to occur in the late Miocene. Populations from isolated river systems surrounding the Gulf of Mexico are almost all monophyletic. Mississippi drainage samples formed a shallow clade with recent diversification and showed evidence of recent population expansion. Main conclusions The historical biogeography of the L. teres species complex is broadly consistent with tested ichthyofaunal models. The timing of speciation and intraspecific divergences correspond to low sea‐level events suggesting that Gulf Coast sea‐level fluctuations are responsible for dispersal (sea‐level recession) and subsequent cladogenesis (sea‐level inundation). Consistent with numerous other freshwater studies, we found the Mobile Basin to be a suture zone, which may be due to the narrow, offshore continental shelf.

Journal of Biogeography

Origin of the Sudbury Complex by meteoritic impact: Neodymium isotopic evidence

Samarium-neodymium isotopic data on whole rocks and minerals of the Sudbury Complex in Canada gave an igneous crystallization age of 1840 ± 21 × 10 6 years. The initial epsilon neodymium values for 15 whole rocks are similar to those for average upper continental crust, falling on the crustal trend of neodymium isotopic evolution as defined by shales. The rare earth element concentration patterns of Sudbury rocks are also similar to upper crustal averages. These data suggest that the Sudbury Complex formed from melts generated in the upper crust and are consistent with a meteoritic impact.

Ontario