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Low flows from drought and water use reduced total dissolved solids fluxes in the Lower Colorado River Basin between 1976 to 2008

Study area: We evaluated the Virgin, Verde, Salt, and Gila Rivers in the Lower Colorado River Basin. The watersheds have extents in Arizona, Utah, New Mexico, US and Sonora, MX. Study focus: We calculated trends in total dissolved solids (TDS) concentrations and fluxes with the Weighted Regressions on Time, Discharge and Season model. The modeling framework leverages daily streamflow and discrete water quality observations at specific monitoring sites. We evaluated trends for a common period (1976–2008) and the whole period of record at each monitoring site in terms of climate and anthropogenic controls. New hydrologic insights for the region: Three rivers had persistent TDS concentrations exceeding the EPA secondary drinking water standard. All were associated with a geologic source of TDS. We observed increases and decreases in TDS concentrations at our monitoring sites, contrasting with global freshwater salinization and declining TDS concentrations in the Upper Colorado River Basin (UCRB). We attributed concentration variability to wintertime hydroclimatic forcing, with secondary influences of human water use. Reservoirs may decrease TDS concentrations by 50%. Efficiency improvements in irrigation and mining water uses may decrease TDS concentrations, while municipal growth increases TDS concentrations. We observed TDS flux declines at most monitoring sites. We attributed up to 85% of the TDS flux trend to changes in streamflow arising from drought and groundwater use. This study informs salinity dynamics in arid and aridifying locations, including the UCRB.

Lower Colorado River Basin

Water quality and quantity of selected springs and seeps along the Colorado River corridor, Utah and Arizona: Arches National Park, Canyonlands National Park, Glen Canyon National Recreation Area, and Grand Canyon National Park, 1997-98

The U.S. Geological Survey, in cooperation with the National Park Service conducted an intensive assessment of selected springs along the Colorado River Corridor in Arches National Park, Canyonlands National Park, Glen Canyon National Recreation Area, and Grand Canyon National Park in 1997 and 1998, for the purpose of measuring and evaluating the water quality and quantity of the resource. This study was conducted to establish baseline data for the future evaluation of possible effects from recreational use and climate change. Selected springs and seeps were visited over a study period from 1997 to 1998, during which, discharge and on-site chemical measurements were made at selected springs and seeps, and samples were collected for subsequent chemical laboratory analysis. This interdisciplinary study also includes simultaneous studies of flora and fauna, measured and sampled coincidently at the same sites. Samples collected during this study were transported to U.S. Geological Survey laboratories in Boulder, Colorado, where analyses were performed using state-of-the-art laboratory technology. The location of the selected springs and seeps, elevation, geology, aspect, and onsite measurements including temperature, discharge, dissolved oxygen, pH, and specific conductance, were recorded. Laboratory analyses include determinations for alkalinity, aluminum, ammonium (nitrogen), antimony, arsenic, barium, beryllium, bismuth, boron, bromide, cadmium, calcium, cerium, cesium, chloride, chromium, cobalt, copper, dissolved inorganic carbon, dissolved organic carbon, dysprosium, erbium, europium, fluoride, gadolinium, holmium, iodine, iron, lanthanum, lead, lithium, lutetium, magnesium, manganese, mercury, molybdenum, neodymium, nickel, nitrate (nitrogen), nitrite (nitrogen), phosphate, phosphorus, potassium, praseodymium, rhenium, rubidium, samarium, selenium, silica, silver, sodium, strontium, sulfate, tellurium, terbium, thallium, thorium, thulium, tin, titanium, tungsten, uranium, vanadium, yttrium, ytterbium, zinc, and zirconium in these springs and seeps. Biological observations include physical setting, vegetation, invertebrate habitats, and invertebrate microhabitats.

Arizona, Utah

Holocene paleohydrology from alpine lake sediment, Emerald Lake, Wasatch Plateau of central Utah, USA

Holocene sediments at Emerald Lake in central Utah (3090 m asl) document the paleohydroclimatic history of the western Upper Colorado River headwater region. Multi-proxy analyses of sediment composition, mineralogy, and stable isotopes of carbonate (δ 18 O and δ 13 C) show changes in effective moisture for the past ca. 10,000 years at millennial to decadal timescales. Emerald Lake originated as a shallow, closed-basin cirque pond during the Early Holocene. By ca. 7000 cal yr BP, higher lake levels and carbonate δ 18 O values indicate rising effective moisture and higher proportions of summer precipitation continued at least until ca. 5500 cal yr BP when a landslide entered the lake margin. Between ca. 4500 and 2400 cal yr BP dry conditions at Emerald Lake envelop the timing of the ‘Late Holocene Dry Period’ identified at lower elevations. For the past ca. 2500 years, Emerald Lake δ 18 O values were relatively low, indicating wetter conditions and higher snow input (compared to rain), except for dry periods at ca. 2000 cal yr BP and during the Medieval Climate Anomaly at ca. 1000 and ca. 500 cal yr BP. Results provide a long-term perspective on precipitation extremes that influence regional water supplies from a snow-dominated catchment typical of the predominant source region for the Upper Colorado River.

Utah

Mass balance of two perennial snowfields: Niwot Ridge, Colorado and the Ulaan Taiga, Mongolia.

Perennial snowfields are generally receding worldwide, though the precise mechanisms causing recessions are not always well understood. Here we apply a numerical snowpack model to identify the leading factors controlling the mass balance of two perennial snowfields that have significant human interest: Arapaho glacier, located at Niwot Ridge in the Colorado Rocky Mountains (United States), and a snowfield located in the Ulaan Taiga (Mongolia). The two locations were chosen because they differ in elevation, slope and aspect. However, both have sub-arctic climates and are located within semi-arid regions. We show that for these two locations the snowfield mass balance is primarily sensitive to air temperature and wind speed, followed by precipitation and dust deposition amounts. We find that the sensitivities are similar for the center of the snowfield as well as the margins.

Colorado

Spatial reconstructions and comparisons of historic snow avalanche frequency and extent using tree rings in Glacier National Park, Montana, U.S.A.

Natural snow avalanches have periodically damaged infrastructure and disrupted railroad and highway traffic at the southwestern corner of Glacier National Park, Montana. The 94-year history of these disruptions constitutes an uncommon record of natural avalanches spanning over nine decades and presents a unique opportunity to examine how natural avalanche frequency and minimum extent have varied over time due to climatic or biophysical changes. This study compared the historic record of natural avalanches in one avalanche path with tree-ring evidence of avalanches from 109 cross sections and increment cores collected in the same path. Results from combined historic and tree-ring records yielded 27 avalanche years in the 1910-2003 chronology, with the historic record alone underestimating avalanche years by half. Mean return period was 3.2 years. Interpolated maps allowed for more spatially precise estimates of return periods throughout the runout zone than previous studies. The maps show return periods increase rapidly downslope from 2.3 to 25 years. Avalanche years were associated with positive Snow Water Equivalent anomalies at a nearby snow course. Minimum avalanche extent was highly variable but not associated with snowpack anomalies. Most avalanche years coincided with years in which the mean January-February Pacific Decadal Oscillation (PDO) and El Nin??o-Southern Oscillation (ENSO) 3.4 indices were neutral. The findings suggest that changes in Pacific climate patterns that influence snowfall could also alter the frequency of natural snow avalanches and could thus change disturbance patterns in the montane forests of the canyon. ?? 2008 Regents of the University of Colorado.

Arctic, Antarctic, and Alpine Research

Natural resource management confronts the growing scale and severity of ecosystem responses to drought and wildfire

Intensification of drought and wildfire associated with climate change has triggered widespread ecosystem stress and transformation. Natural resource managers are on the frontline of these changes, yet their perspectives on whether management actions match the scale and align with the severity of ecosystem responses to improve outcomes are not well understood. To provide new insight, a new conceptual framework that linked scale and severity was tested by conducting interviews and surveys of staff associated with natural resource management on the Colorado Plateau in the southwestern United States (U.S.), which contains the highest concentration of public lands in the contiguous U.S. Results indicate that drought was experienced more frequently than wildfire, and both stressors were happening at large scales and moderate to abrupt timeframes with a high degree of impact to ecosystems. Ecosystem responses were perceived to increase in severity under future climate change with limited capacity to recover, and a majority of resource managers expressed that they had low control to shape these trajectories. Although management strategies to address drought and wildfire were well recognized, adaptation-specific actions remained unclear or had limited financial and staffing resources to support implementation. Additional effort could help close a growing misalignment between management actions and natural resource responses, including effective science communication, refined information tailored to meet adaptation goals at management-relevant spatiotemporal scales, and opportunities for adaptive management that can proactively address intensification of drought and wildfire.

Ecology and Society

Effects of weather on survival in populations of boreal toads in Colorado

Understanding the relationships between animal population demography and the abiotic and biotic elements of the environments in which they live is a central objective in population ecology. For example, correlations between weather variables and the probability of survival in populations of temperate zone amphibians may be broadly applicable to several species if such correlations can be validated for multiple situations. This study focuses on the probability of survival and evaluates hypotheses based on six weather variables in three populations of Boreal Toads (Bufo boreas) from central Colorado over eight years. In addition to suggesting a relationship between some weather variables and survival probability in Boreal Toad populations, this study uses robust methods and highlights the need for demographic estimates that are precise and have minimal bias. Capture-recapture methods were used to collect the data, and the Cormack-Jolly-Seber model in program MARK was used for analysis. The top models included minimum daily winter air temperature, and the sum of the model weights for these models was 0.956. Weaker support was found for the importance of snow depth and the amount of environmental moisture in winter in modeling survival probability. Minimum daily winter air temperature was positively correlated with the probability of survival in Boreal Toads at other sites in Colorado and has been identified as an important covariate in studies in other parts of the world. If air temperatures are an important component of survival for Boreal Toads or other amphibians, changes in climate may have profound impacts on populations. Copyright 2008 Society for the Study of Amphibians and Reptiles.

Journal of Herpetology

Development of industrial minerals in Colorado

Technology and engineering have helped make mining safer and cleaner for both humans and the environment. Inevitably, mineral development entails costs as well as benefits. Developing a mine is an environmental, engineering, and planning challenge that must conform to many Federal, State, and local regulations. Community collaboration, creative design, and best management practices of sustainability and biodiversity can be positive indicators for the mining industry. A better understanding of aesthetics, culture, economics, geology, climate, vegetation and wildlife, topography, historical significance, and regional land planning is important in resolving land-use issues and managing mineral resources wisely. Ultimately, the consuming public makes choices about product use (including water, food, highways, housing, and thousands of other items) that influence operations of the mineral industry. Land planners, resource managers, earth scientists, designers, and public groups have a responsibility to consider sound scientific information, society's needs, and community appeals in making smart decisions concerning resource use and how complex landscapes should change. An effort to provide comprehensive geosciences data for land management agencies in central Colorado was undertaken in 2003 by scientists of the U.S. Geological Survey and the Colorado Geological Survey. This effort, the Central Colorado Assessment Project, addressed a variety of land-use issues: an understanding of the availability of industrial and metallic rocks and minerals, the geochemical and environmental effects of historic mining activity on surface water and groundwater, and the geologic controls on the availability and quality of groundwater. The USDA Forest Service and other land management agencies have the opportunity to contribute to the sustainable management of natural aggregate and other mineral resources through the identification and selective development of mineral resources and the reclamation of mines on lands that they administer. The information in this Circular will help them carry out that task.

Colorado

Climate-driven sulfate export in alpine watersheds may stimulate methylmercury production

Climate change is increasing sulfate export and changing wetland extent in mountain regions. These changes may increase microbially mediated production of the neurotoxic substance methylmercury due to enhanced sulfate metabolism in mountain environments. Here, we assess methylmercury concentrations and formation rates across high-elevation wetlands in the Colorado Rocky Mountains. We also investigate sulfate controls on methylmercury production within subalpine peatlands by amending soils with sulfate to mimic increased stream export of sulfate from the alpine zone and measuring methylmercury formation rates for different sulfate treatments. We found that subalpine peatlands have statistically significant higher methylmercury concentrations and formation rates compared to alpine, mineral-soil wetlands. Methylmercury production in subalpine peatlands also increased significantly ( p < 0.05) following sulfate additions; the highest rates occurred in sediments with intermediate extractable sulfate concentrations (∼0.60–1.4 mg sulfate g −1 dry soil). Our study is the first to identify soil sulfate-related thresholds for methylmercury production and sulfate-limitation of methylmercury production in subalpine peatlands. These findings highlight important linkages between climate-driven mineral weathering and mercury cycling in mountain regions globally.

Colorado

Proceedings of the 12th Biennial Conference of research on the Colorado Plateau

Introduction The Colorado Plateau is a physiographic region that encompasses 330,000 square kilometers in parts of four states in the southwestern United States (Colorado, Utah, New Mexico, and Arizona). Known for its high deserts, the Colorado Plateau also includes isolated mountains, high plateaus, and rugged canyons. Not only is the region topographically diverse, but geologically, biologically, and culturally diverse as well. The landscape is managed by Federal entities including the Bureau of Land Management, the National Park Service, and the U.S. Forest Service; Tribal nations including the Navajo Nation, Kaibab Paiute, Mountain Ute, Southern Ute, Hopi, Zuni, Hualapai, Havasupai, and White Mountain Apache Tribes; State land and wildlife management agencies; and privately owned holdings, creating complex interactions and management challenges. Population growth, increased tourism to Federal and State lands, and energy development have increased water demands and altered land-use patterns, and these changes have emerged as management challenges facing the people working and living in the region. Climate change, particularly the ongoing drought, has exacerbated the effects of population growth, land-use change, and other stressors such as invasive species. As managers seek solutions to the challenges facing the region&rsquo;s natural and cultural resources, the Biennial Conference of Science and Management of the Colorado Plateau has become an important venue for exchanging information about emerging management concerns and recent scientific research. Each biennial conference has sought to promote discussion, information sharing, and productive communication among the managers, scientists, students, administrators, tribal representatives, and others who attend the conference with the goal of enhancing the use of the best available science to manage the region&rsquo;s incomparable natural and cultural resources. The publication and dissemination of a conference proceedings series expands the reach of the conference beyond those people in attendance and creates a record on the research presented. The idea of producing a conference proceedings, and its subsequent publication, first occurred in 1993 following the first biennial conference in 1991. A published volume of contributed papers has followed each subsequent biennial conference, including this volume. The venue for publishing proceedings has changed over the years and has included the National Park Service, the Government Printing Office, the U.S. Geological Survey, and University of Arizona Press. Recently, van Riper and others (2015) published a compilation of the abstracts from the 11 previous conference proceedings. Collectively, the proceedings highlight approximately 25 years of natural- and cultural-resources research, promoting the integration of research with resource management across the Colorado Plateau. This volume is freely downloadable by the public, thereby further expanding the influence of this conference beyond the Colorado Plateau. The 12th Biennial Conference held in Flagstaff, Arizona, from September 16 to 19, 2013, covered a range of topics in the physical, biological, and socio-cultural sciences. The conference was organized and hosted by Northern Arizona University&rsquo;s ( NAU) Merriam-Powell Center for Environmental Research, the Colorado Plateau Cooperative Ecosystem Studies Unit, and the U.S. Geological Survey Southwest Biological Science Center. Financial and in-kind support was provided by a wide range of organizations including the U.S. Forest Service, National Park Service, Bureau of Land Management, Grand Canyon Trust, Colorado Plateau Research Station, and various NAU entities. NAU sponsors include the Landscape Conservation Initiative, School of Forestry, School of Earth Science and Environmental Sustainability, Office of the Provost, and Office of the Vice President of Research. Contributors to these proceedings include researchers and managers from Federal, State, and Tribal governments, universities, private entities, and non-profit organizations. In this regard, this conference has wide-ranging support and participation among private and public entities involved in the science and management of natural resources on the Colorado Plateau.

Arizona, Colorado, New Mexico, Utah

Increased temperature and altered summer precipitation have differential effects on biological soil crusts in a dryland ecosystem

Biological soil crusts (biocrusts) are common and ecologically important members of dryland ecosystems worldwide, where they stabilize soil surfaces and contribute newly fixed C and N to soils. To test the impacts of predicted climate change scenarios on biocrusts in a dryland ecosystem, the effects of a 2–3 °C increase in soil temperature and an increased frequency of smaller summer precipitation events were examined in a large, replicated field study conducted in the cold desert of the Colorado Plateau, USA. Surface soil biomass (DNA concentration), photosynthetically active cyanobacterial biomass (chlorophyll a concentration), cyanobacterial abundance (quantitative PCR assay), and bacterial community composition (16S rRNA gene sequencing) were monitored seasonally over 2 years. Soil microbial biomass and bacterial community composition were highly stratified between the 0–2 cm depth biocrusts and 5–10 cm depth soil beneath the biocrusts. The increase in temperature did not have a detectable effect on any of the measured parameters over 2 years. However, after the second summer of altered summer precipitation pattern, significant declines occurred in the surface soil biomass (avg. DNA concentration declined 38%), photosynthetic cyanobacterial biomass (avg. chlorophyll a concentration declined 78%), cyanobacterial abundance (avg. gene copies g −1 soil declined 95%), and proportion of Cyanobacteria in the biocrust bacterial community (avg. representation in sequence libraries declined 85%). Biocrusts are important contributors to soil stability, soil C and N stores, and plant performance, and the loss or reduction of biocrusts under an altered precipitation pattern associated with climate change could contribute significantly to lower soil fertility and increased erosion and dust production in dryland ecosystems at a regional scale.

Global Change Biology

Mercury cycling across a U.S. semi-arid mountain ecosystem elevation gradient

Mountains comprise ∼30% of the Earth's surface, but mercury (Hg) cycling in these regions remains understudied, particularly in the semi-arid western U.S. where strong climatic and ecological gradients in mountainous landscapes influence Hg deposition, retention, and bioaccumulation. In this study, we quantified growing season inputs, storage, and bioaccumulation of Hg along a ∼2,000 m elevation gradient in the Colorado Rocky Mountains, spanning the plains to the alpine. We measured Hg in atmospheric deposition, vegetation, soil, and 12-day-old chickadees. Accounting for percent canopy cover, open precipitation was the largest component of atmospheric deposition at all elevations, followed by throughfall and litterfall fluxes. Atmospheric Hg fluxes peaked at mid-elevations, likely due to cloud-cap dynamics and denser canopy cover. Total gaseous Hg and precipitation fluxes were highest at low elevations, likely reflecting local emissions and meteorological pooling. Surface soil Hg storage was more strongly predicted by organic matter content ( R 2 = 0.49; p < 0.01) and water retention ( R 2 = 0.45; p < 0.01) than by elevation ( R 2 = 0.21; p < 0.05). Alpine soils (66.3 ± 25.3 ng g −1 ) had significantly higher total Hg concentrations than lower elevations (<41.0 ± 12.7 ng g −1 ; p < 0.01), likely reflecting slower organic matter turnover. Soils on north-facing slopes also retained significantly higher pools of Hg in surface soils compared with south- and east-facing slopes. Vegetation Hg pools were greatest in the alpine region, likely due to long-lived plant species. Methylmercury (MeHg) concentrations in chickadee feathers peaked at mid-elevations (205 ± 155 ng g −1 ), corresponding to higher ecosystem Hg inputs via throughfall. Our results show that deposition, canopy cover, and meteorological conditions—not elevation alone—predict Hg retention and bioaccumulation.

Colorado

Fire severity and changing composition of forest understory plant communities

Questions Gradients of fire severity in dry conifer forests can be associated with variation in understory floristic composition. Recent work in dry conifer forests in California, USA, has suggested that more severely burned stands contain more thermophilic taxa (those associated with warmer and drier conditions), and that forest disturbance may therefore accelerate floristic shifts already underway due to climate change. However, it remains unknown how rapidly thermophilic taxa shifts occur following disturbance, how long such shifts are likely to persist, and how different thermophilic post‐disturbance communities are from pre‐disturbance communities. Location Colorado Front Range, USA. Methods We investigated these questions using a unique 15‐year vegetation plot dataset that captures pre‐ and post‐fire understory community composition across a gradient of fire severity in dry conifer forests, classifying taxa using the biogeographic affinity concept. Results Thermophilization (defined here as a decrease in the ratio of cool‐mesic taxa to warm‐xeric taxa, based on biogeographic affinity of paleobotanical lineages) was observed as early as one year post‐fire for all fire severity classes, but was stronger at sites that burned at higher severity. The ratio of cool‐mesic to warm‐xeric taxa recovered to pre‐fire levels within 10 years in stands that burned at low severity, but not in stands that burned at moderate or high severity. The process of thermophilization after high‐severity fire appears to be driven primarily by the gain of warm‐xeric taxa that were absent before the fire, but losses of cool‐mesic taxa, which did not return during the duration of the study, also played a role. Conclusions Decreases in canopy cover appear to be a main contributor to understory thermophilization. Fine‐scale heterogeneity in post‐fire forest structure is likely an important driver of floristic diversity, creating the microclimatic variation necessary to maintain floristic refugia for species mal‐adapted to increasingly warm and dry conditions.

Colorado

Meteorological Data near Rabbit Ears Pass, Colorado, U.S.A., 1984-2008

In 1983, a snowmelt energy budget study was initiated by the U.S. Geological Survey on a small watershed near Rabbit Ears Pass, Colorado, to better understand snowmelt processes. The study included data collection from hydrological and meteorological instrumentation. Interest in long term, high-altitude meteorological sites has increased recently due to the increased awareness of global climate change. The meteorological data collected near Rabbit Ears Pass may aid researchers involved in global climate change studies. Meteorological data from 1984 to 2008 are presented.

Data Series

Water-quality data for streams in the southern Yampa River basin, northwestern Colorado

Increases in coal mining in Colorado have raised questions regarding the impact of mining on the quality of surface water. An area where increases have caused particular concern is in northwestern Colorado, in the southern part of the Yampa River basin. In order to identify the changes produced by mining in the area it is necessary first to identify the water chemistry resulting from the geology, climate, and land use. To answer the above questions, a program for the synoptic collection of water-quality data was developed in which a network of sampling sites was established to determine the surface-water chemistry. This study was begun in April 1981. Water-quality samples were collected from sites on all continuously flowing streams in the southern part of the Yampa River basin from the Oak Creek drainage near Steamboat Springs on the east to the surface-water gaging station on the Yampa River near Maybell on the west. Each water-quality site was sampled repetitively as changes occurred in discharge and specific conductance. Water-quality data from surface-water gaging stations in the area for water years 1976 to 1981 are included in the report.

Open-File Report

Environmental Setting and Implications on Water Quality, Upper Colorado River Basin, Colorado and Utah

The Upper Colorado River Basin in Colorado and Utah is 1 of 60 study units selected for water-quality assessment as part of the U.S. Geological Survey's National Water-Quality Assessment program, which began full implementation in 1991. Understanding the environmental setting of the Upper Colorado River Basin study unit is important in evaluating water-quality issues in the basin. Natural and human factors that affect water quality in the basin are presented, including an overview of the physiography, climatic conditions, general geology and soils, ecoregions, population, land use, water management and use, hydrologic characteristics, and to the extent possible aquatic biology. These factors have substantial implications on water-quality conditions in the basin. For example, high concentrations of dissolved solids and selenium are present in the natural background water conditions of surface and ground water in parts ofthe basin. In addition, mining, urban, and agricultural land and water uses result in the presence of certain constituents in the surface and ground water of the basin that can detrimentally affect water quality. The environmental setting of the study unit provides a framework of the basin characteristics, which is important in the design of integrated studies of surface water, ground water, and biology.

Water-Resources Investigations Report

Responses of Englemann spruce forests to nitrogen fertilization in the Colorado Rocky Mountains

Two old-growth coniferous forests in Colorado with differing initial soil conditions responded differently to four years of low-level fertilization with ammonium nitrate. The site (Fraser) with an average initial organic horizon soil C:N ratio of 36 and nitrogen (N) pool of 605 kg/ha showed no significant increase in net N mineralization rates. At the Fraser site, foliar and organic horizon soil percentage N increased significantly. In contrast, N mineralization rates and inorganic soil N increased significantly at the site (Loch Vale) with greater soil N (C:N of 24, N pool of 991 kg/ha), while foliar N and soil percentage N in the organic layer did not change. We predict continued fertilization at Fraser will narrow the soil C:N ratio to a point where increases in biogeochemical N cycling and fluxes will be detected. Additional N inputs to the site with already low soil C:N ratios will enhance N mineralization rates and leaching losses. The coniferous forests at Fraser and Loch Vale are similar in species composition, stand age, substrate, aspect, and climate. The differences in soil conditions strong enough to cause contrasting responses to fertilization could be due to differences in atmospheric N deposition. Regardless of the reason, the size of the organic soil N pool and C:N ratio of mature coniferous forests in Colorado controls the responsiveness of N pools and fluxes to fertilization, and even low levels of fertilization are sufficient to initiate measurable biogeochemical changes.

Colorado

Assessment of riparian vegetation patterns and change downstream from Glen Canyon Dam from 2014 to 2019

Changes in riparian vegetation cover and composition occur in relation to flow regime, geomorphic template, and climate, and can have cascading effects on aquatic and terrestrial ecosystems. Tracking such changes over time is therefore an important part of monitoring the condition and trajectory of riparian ecosystems. Maintaining diverse, self-sustaining riparian vegetation comprised of mostly native species is identified in the Glen Canyon Dam Long-Term Experimental and Management Plan as a key resource objective for the section of the Colorado River between Glen Canyon Dam and Lake Mead. The U.S. Geological Survey Grand Canyon Monitoring and Research Center implemented an annual monitoring program in 2014 to assess the status and trends of riparian vegetation along this section of river, particularly as they relate to flow regime. In this report, we summarize plant species composition and cover data collected under the annual monitoring program from 2014 to 2019, with special consideration given to the hydrologic position, associated geomorphic feature class, local climate patterns, native and nonnative species, and floristic region for key vegetation metrics and species. We divided the study area into four river segments (referred to as Glen Canyon, Marble Canyon, eastern Grand Canyon, and western Grand Canyon) on the basis of geography and floristic composition and calculated each recorded plant species’ relative frequency and foliar cover by river segment. These data were then used to evaluate species composition relationships among river segments, hydrologic zones, geomorphic features, and sampling years through ordination analysis. Temporal trends in our focal resource objectives—species richness, total foliar cover, proportion of native to nonnative species richness, proportion of native to nonnative species cover, Tamarix cover, Pluchea sericea cover, and Baccharis species cover—were assessed using mixed-effects models. Four patterns related to species composition emerged: (1) species composition of fixed-site sandbars differed from that of randomly selected sites (including randomly selected sandbars), (2) species composition of Glen Canyon sites differed from that of other previously identified floristic regions, (3) species composition differed across hydrologic zones related to dam operations, and (4) species composition within river segments did not change across years. For temporal patterns, four main findings emerged: (1) trends differed between fixed-sites and randomly selected sites; (2) although few directional changes were observed from 2014 to 2019, Baccharis species cover increased at randomly selected sites in areas influenced by daily water fluctuations; (3) native species cover and richness were greater than nonnative species cover and richness across all hydrologic zones; and (4) the temporal trend metrics used here can be used across floristic groups, enabling assessment of the Colorado River ecosystem as a whole. In addition to these findings, lists of recorded plant species are included as appendixes. The variations and patterns in vegetation status and trends presented in this report can be used as a baseline against which future monitoring can be compared.

Arizona