Search USGSSearch

SEARCH · Search USGS

Results for “Evolution”

Search indexed USGS publications on groundwater, aquifers, geologic maps, mineral resources and earthquakes. Explore source records by subject and place.

Quote a phrase for an exact phrase match. Source license links do not imply unrestricted reuse.

At least 289 records · Page 16Linked to original sources

Role of a polymorphism in a Hox/Pax-responsive enhancer in the evolution of the vertebrate spine

Patterning of the vertebrate skeleton requires the coordinated activity of Hox genes. In particular, Hox10 proteins are essential to set the transition from thoracic to lumbar vertebrae because of their rib-repressing activity. In snakes, however, the thoracic region extends well into Hox10-expressing areas of the embryo, suggesting that these proteins are unable to block rib formation. Here, we show that this is not a result of the loss of rib-repressing properties by the snake proteins, but rather to a single base pair change in a Hox/Paired box (Pax)-responsive enhancer, which prevents the binding of Hox proteins. This polymorphism is also found in Paenungulata, such as elephants and manatees, which have extended rib cages. In vivo, this modified enhancer failed to respond to Hox10 activity, supporting its role in the extension of rib cages. In contrast, the enhancer could still interact with Hoxb6 and Pax3 to promote rib formation. These results suggest that a polymorphism in the Hox/Pax-responsive enhancer may have played a role in the evolution of the vertebrate spine by differently modulating its response to rib-suppressing and rib-promoting Hox proteins.

Proceedings of the National Academy of Sciences

State shifts in the deep Critical Zone drive landscape evolution in volcanic terrains

Understanding the near-surface environment where atmospheric and solid earth processes interact, often termed the “Critical Zone,” is important for assessing resources and building resilient societies. Here, we examine a volcanic landscape in the Oregon Cascade Range, an understudied Critical Zone setting that is host to major regional water resources, pervasive silicate weathering, and significant geohazards. We leverage a bedrock age chronosequence to show that the volcanic Critical Zone undergoes a structural shift, from depth extents of >1 km to meters, over timescales of ~1 My. We map an active groundwater volume comparable to major continental lakes, stored at the Cascade Range crest. This state shift makes volcanic landscape evolution a unique probe of deep coupling between Earth systems.

PNAS

Biostratigraphy and evolution of the Momipites‐Caryapollenites lineage in the early tertiary in the Wind River Basin, Wyoming

Sequential morphologic changes in pollen of the Momipites‐Caryapollenites lineage from Paleocene strata in the Wind River Basin of Wyoming appear to reflect evolution within the family Juglandaceae. The stratigraphic occurrence of the species within this complex permits the establishment of six biostratigraphic zones that can be correlated between outcropping and subsurface sequences within the basin and to surrounding areas. Trends involving changes in shape and size of the pollen and in structure of exine at the poles proceed from a basic form of Momipites to six other distinguishable species. The genus Caryapollenites appears to have been derived from the basic form of Momipites by changes in size and development of heteropolarity in pore position. Species of Momipites described as new are M. wyomingensis, M. waltmanensis, M. ventifluminis, M. actinus, M. anellus, and M, leffingwellii. The genus Caryapollenites is redescribed and C. prodromus, C. imparalis, C. inelegans, and C. wodehousei are described as new.

Wyoming

New Washakiin primates (Omomyidae) from the Eocene of Wyoming and Colorado, and comments on the evolution of the Washakiini

Two new species of washakiin omomyids occur in deposits of early Bridgerian age. Shoshonius bowni , sp. nov., from the Aycross Formation, Absaroka Range, Wyoming, differs from S. cooperi in having enlarged conules on the upper molars and a second metaconule, features convergent with Washakius insignis. Washakius izetti , sp. nov., from the Green River Formation, Piceance Creek Basin, Colorado, is the most primitive known species of Washakius , showing incipient development of features present in the later W. insignis and W. woodringi. Washakius , cf. W. izetti occurs in the early Bridgerian of the Huerfano Basin. W. izetti is closely related to Utahia kayi , a. washakiin possibly related to Stockia. Hemiacodon , sometimes included in the Washakiini, is probably more closely related to the Omomyini. Stockia is distinct from Omomys and is questionably included in the Washakiini, of which Loveina is the stem taxon. More advanced washakiins form two groups between which there was significant parallel evolution in dental morphology. One group includes Washakius, Dyseolemur, Utahia , and possibly Stockia , and is characterized by development of an open talonid notch before the consistent appearance of metastylids. The other group consists of Shoshonius , where the establishment of metastylids preceded the full opening of the talonid basins.

Journal of Vertebrate Paleontology

Evolution of trophic transmission in parasites: Why add intermediate hosts?

Although multihost complex life cycles (CLCs) are common in several distantly related groups of parasites, their evolution remains poorly understood. In this article, we argue that under particular circumstances, adding a second host to a single-host life cycle is likely to enhance transmission (i.e., reaching the target host). For instance, in several situations, the propagules of a parasite exploiting a predator species will achieve a higher host-finding success by encysting in a prey of the target predator than by other dispersal modes. In such a case, selection should favor the transition from a singleto a two-host life cycle that includes the prey species as an intermediate host. We use an optimality model to explore this idea, and we discuss it in relation to dispersal strategies known among free-living species, especially animal dispersal. The model found that selection favored a complex life cycle only if intermediate hosts were more abundant than definitive hosts. The selective value of a complex life cycle increased with predation rates by definitive hosts on intermediate hosts. In exploring trade-offs between transmission strategies, we found that more costly trade-offs made it more difficult to evolve a CLC while less costly trade-offs between traits could favor a mixed strategy.

American Naturalist

Eruption and deposition of the Fisher Tuff (Alaska)--Evidence for the evolution of pyroclastic flows

Recognition that the Fisher Tuff (Unimak Island, Alaska) was deposited on the leeside of an ∼500–700‐m‐high mountain range (Tugamak Range) more than 10 km away from its source played a major role in defining pyroclastic flows as momentum‐driven currents. We reexamined the Fisher Tuff to evaluate whether deposition from expanded turbulent clouds can better explain its depositional features. We studied the tuff at 89 sites and sieved bulk samples from 27 of those sites. We find that the tuff consists of a complex sequence of deposits that record the evolution of the eruption from a buoyant plume (22 km) that deposited ∼0.2 km 3 of dacite magma as a pyroclastic fall layer to erupting ∼10–100 km 3 of andesitic magma as Scoria‐rich pyroclastic falls and flows that were mainly deposited to the north and northwest of the caldera, including those in valleys within the Tugamak Range. The distribution of the flow deposits and their welding, internal stratification, and the occurrence of lithic breccia all suggest that the pyroclastic flows were fed from a fountaining column that vented from an inclined conduit, the first time such a conduit has been recognized during a large‐volume caldera eruption. Pyroclastic flow deposits before and after the mountain range and thin veneer deposits high in the range are best explained by a flow that was stratified into a dense undercurrent and an overriding dilute turbulent cloud, from which deposition before the range was mainly from the undercurrent. When the flow ran into the mountain range, however, the undercurrent was blocked, but the turbulent cloud continued on. As the flow continued north, it restratified, forming another undercurrent. The Fisher Tuff thus records the passing of a flow that was significantly higher (800–1100 m thick) than the mountain range and thus did not require excessive momentum.

Alaska

Evolution of pathogen virulence across space during an epidemic

We explore pathogen virulence evolution during the spatial expansion of an infectious disease epidemic in the presence of a novel host movement trade-off, using a simple, spatially explicit mathematical model. This work is motivated by empirical observations of the Mycoplasma gallisepticum invasion into North American house finch ( Haemorhous mexicanus ) populations; however, our results likely have important applications to other emerging infectious diseases in mobile hosts. We assume that infection reduces host movement and survival and that across pathogen strains the severity of these reductions increases with pathogen infectiousness. Assuming these trade-offs between pathogen virulence (host mortality), pathogen transmission, and host movement, we find that pathogen virulence levels near the epidemic front (that maximize wave speed) are lower than those that have a short-term growth rate advantage or that ultimately prevail (i.e., are evolutionarily stable) near the epicenter and where infection becomes endemic (i.e., that maximize the pathogen basic reproductive ratio). We predict that, under these trade-offs, less virulent pathogen strains will dominate the periphery of an epidemic and that more virulent strains will increase in frequency after invasion where disease is endemic. These results have important implications for observing and interpreting spatiotemporal epidemic data and may help explain transient virulence dynamics of emerging infectious diseases.

American Naturalist

The ecology and evolution of synchronized reproduction in long-lived plants

Populations of many long-lived plants exhibit spatially synchronized seed production that varies extensively over time, so that seed production in some years is much higher than on average, while in others, it is much lower or absent. This phenomenon termed masting or mast seeding has important consequences for plant reproductive success, ecosystem dynamics and plant–human interactions. Inspired by recent advances in the field, this special issue presents a series of articles that advance the current understanding of the ecology and evolution of masting. To provide a broad overview, we reflect on the state-of-the-art of masting research in terms of underlying proximate mechanisms, ontogeny, adaptations, phylogeny and applications to conservation. While the mechanistic drivers and fitness consequences of masting have received most attention, the evolutionary history, ontogenetic trajectory and applications to plant–human interactions are poorly understood. With increased availability of long-term datasets across broader geographical and taxonomic scales, as well as advances in molecular approaches, we expect that many mysteries of masting will be solved soon. The increased understanding of this global phenomenon will provide the foundation for predictive modelling of seed crops, which will improve our ability to manage forests and agricultural fruit and nut crops in the Anthropocene.

Philosophical Transactions of the Royal Society B:

A transdisciplinary approach to growing an applied science of cultural evolution for a sustainable future

Addressing sustainability challenges requires an integrative approach that bridges scientific research with practical application. The field of cultural evolution (CE) offers a perspective that may guide transitions and cultural transformations for a sustainable future. However, there have been few efforts to apply this field to sustainability challenges. This study explores how CE can inform sustainability practices through mechanisms of social learning, processes of cultural adaptation and conditions that promote cooperative governance. Through a two-phase research design—comprising online discussions with professionals from six domains, and an international workshop—we examined the perceived benefits, challenges and opportunities for applying CE in real-world contexts. Our findings indicate that professionals recognize CE’s potential to enhance knowledge dissemination, foster adaptation to social–ecological changes, improve governance structures and generate cooperation. However, barriers such as complex terminology, unfamiliarity with CE, the lack of clear, context-specific evidence of added value and competition with existing frameworks hinder its application. To overcome these challenges, we propose simplifying CE concepts, demonstrating its unique contributions, and continuing to foster co-production with practitioners to refine its applicability. This study is an initial step in building an applied science of CE that can support sustainability transformations across diverse domains.

Philosophical Transactions of the Royal Society B

Understanding the evolution of groundwater-contaminant plume chemistry emanating from legacy contaminant sources: An example from a long-term crude oil spill

Understanding the evolution of plumes emanating from residual hydrocarbon contaminant sources requires evaluating how changes in source compositions over time cause changes in dissolved plume chemistry as residual sources age. This study investigates such changes at the site of a 1979 crude-oil pipeline spill and is the first comprehensive look at groundwater chemistry associated with a residual hydrocarbon source zones in different stages of aging. The data show a direct relationship between concentrations of benzene and naphthalene in the residual oil and those measured in water samples collected below the oil. Groundwater associated with oil near the spill site had different chemical composition compared with water associated with oil that had spread downgradient from the spill zone, indicating a shift in biodegradation reactions. These results emphasize that source zone processes are spatially and temporally heterogeneous and should be accounted for in natural attenuation studies where residual source zones persist.

Minnesota

Influence of volcanic activity on the population genetic structure of Hawaiian Tetragnatha spiders: Fragmentation, rapid population growth and the potential for accelerated evolution

Volcanic activity on the island of Hawaii results in a cyclical pattern of habitat destruction and fragmentation by lava, followed by habitat regeneration on newly formed substrates. While this pattern has been hypothesized to promote the diversification of Hawaiian lineages, there have been few attempts to link geological processes to measurable changes in population structure. We investigated the genetic structure of three species of Hawaiian spiders in forests fragmented by a 150-year-old lava flow on Mauna Loa Volcano, island of Hawaii: Tetragnatha quasimodo (forest and lava flow generalist), T. anuenue and T. brevignatha (forest specialists). To estimate fragmentation effects on population subdivision in each species, we examined variation in mitochondrial and nuclear genomes (DNA sequences and allozymes, respectively). Population subdivision was higher for forest specialists than for the generalist in fragments separated by lava. Patterns of mtDNA sequence evolution also revealed that forest specialists have undergone rapid expansion, while the generalist has experienced more gradual population growth. Results confirm that patterns of neutral genetic variation reflect patterns of volcanic activity in some Tetragnatha species. Our study further suggests that population subdivision and expansion can occur across small spatial and temporal scales, which may facilitate the rapid spread of new character states, leading to speciation as hypothesized by H. L. Carson 30 years ago.

Molecular Ecology

Evolution of a Holocene delta driven by episodic sediment delivery and coseismic deformation, Puget Sound, Washington, USA

Episodic, large-volume pulses of volcaniclastic sediment and coseismic subsidence of the coast have influenced the development of a late Holocene delta at southern Puget Sound. Multibeam bathymetry, ground-penetrating radar (GPR) and vibracores were used to investigate the morphologic and stratigraphic evolution of the Nisqually River delta. Two fluvial–deltaic facies are recognized on the basis of GPR data and sedimentary characteristics in cores, which suggest partial emplacement from sediment-rich floods that originated on Mount Rainier. Facies S consists of stacked, sheet-like deposits of andesitic sand up to 4 m thick that are continuous across the entire width of the delta. Flat-lying, highly reflective surfaces separate the sand sheets and comprise important facies boundaries. Beds of massive, pumice- and charcoal-rich sand overlie one of the buried surfaces. Organic-rich material from that surface, beneath the massive sand, yielded a radiocarbon age that is time-correlative with a series of known eruptive events that generated lahars in the upper Nisqually River valley. Facies CF consists of linear sandbodies or palaeochannels incised into facies S on the lower delta plain. Radiocarbon ages of wood fragments in the sandy channel-fill deposits also correlate in time to lahar deposits in upstream areas. Intrusive, sand-filled dikes and sills indicate liquefaction caused by post-depositional ground shaking related to earthquakes. Continued progradation of the delta into Puget Sound is currently balanced by tidal-current reworking, which redistributes sediment into large fields of ebb- and flood-oriented bedforms.

Washington

Perspectives on the future of ecology, evolution, and biodiversity from the Council on Microbial Sciences of the American Society for Microbiology

The field of microbial ecology, evolution, and biodiversity (EEB) is at the leading edge of understanding how microbes shape our biosphere and influence the well-being of humankind. To that end, EEB is developing new tools to analyze the massive, complex, transdisciplinary datasets that result from such studies. The American Society for Microbiology’s Council on Microbial Sciences hosted a virtual retreat in 2023 to identify the trajectory of EEB both within the Society and microbiology at large. Retreat presentations and discussions collectively emphasized the interconnectedness of microbes and their outsized global influence on environmental and host health. The retreat repeatedly emphasized that the future of science is through transdisciplinary collaborations that promote integration of diverse datasets and advanced technologies, while also genuinely fostering inclusivity, global collaborations, and public engagement. The American Society for Microbiology and EEB stand poised to lead a paradigm shift that will result in a new era of collaboration, innovation, and societal relevance for microbiology.

mSphere

Impact of the dimethyl sulfoxide reductase superfamily on the evolution of biogeochemical cycles

The dimethyl sulfoxide reductase (or MopB) family is a diverse assemblage of enzymes found throughout Bacteria and Archaea . Many of these enzymes are believed to have been present in the last universal common ancestor (LUCA) of all cellular lineages. However, gaps in knowledge remain about how MopB enzymes evolved and how this diversification of functions impacted global biogeochemical cycles through geologic time. In this study, we perform maximum likelihood phylogenetic analyses on manually curated comparative genomic and metagenomic data sets containing over 47,000 distinct MopB homologs. We demonstrate that these enzymes constitute a catalytically and mechanistically diverse superfamily defined not by the molybdopterin- or tungstopterin-containing [molybdopterin or tungstopterin bis (pyranopterin guanine dinucleotide) (Mo/W- bis PGD)] cofactor but rather by the structural fold that binds it in the protein. Our results suggest that major metabolic innovations were the result of the loss of the metal cofactor or the gain or loss of protein domains. Phylogenetic analyses also demonstrated that formate oxidation and CO 2 reduction were the ancestral functions of the superfamily, traits that have been vertically inherited from the LUCA. Nearly all of the other families, which drive all other biogeochemical cycles mediated by this superfamily, originated in the bacterial domain. Thus, organisms from Bacteria have been the key drivers of catalytic and biogeochemical innovations within the superfamily. The relative ordination of MopB families and their associated catalytic activities emphasize fundamental mechanisms of evolution in this superfamily. Furthermore, it underscores the importance of prokaryotic adaptability in response to the transition from an anoxic to an oxidized atmosphere.

Microbiology Spectrum

Hydrologic budget of the late Oligocene Lake Creede and the evolution of the upper Rio Grande drainage system

The filling history, hydrologic budget, and geomorphic development of ancient Lake Creede and its tributary basin are evaluated to determine the factors that controlled its character. The lake filled the Creede caldera that formed in the late Oligocene as a consequence of the eruption of the Snowshoe Mountain Tuff. The caldera's sedimentary fill accumlated to a depth of about 1.26 km and had a volume of about 89 km 3 . The highest lake level was ~3300 m (10,800 ft) present altitude before it drained eastward across a broad volcanic plateau as the ancestral Rio Grande. A tributary canyon several hundred meters deep was cut into hard rhyolite in the north wall of the caldera before the lake was more than half full; its presence demonstrates that ancient Lake Creede filled slowly and thus occupied a long-lived, closed basin. The slow filling rate is incompatible with the present water flux through the Creede caldera basin, because such a flow would fill the basin geologically instantaneously. This mismatch, together with the recognition that the Oligocene climate was similar to that of today, forces the reexamination of the hydrologic and geomorphic history of the caldera. That appraisal shows that the caldera cannot have resurged rapidly immediately after caldera collapse, and that ancient watershed must have been lass than half as large as the present upper Rio Grande basin. The ancient lake had a more or less constant surface area of about 200 km 2 that approximated a steady-state condition between inflow and evaporation. Although the lake level fluctuated with climatic variations, its surface elevation steadily climbed as sediment accumulated, accelerating as resurgance and dome growth usurped spacewithin the basin. It could have had one playa stage early in its development and another after the basin had nearly filled with sediment, but there is no direct evidence for either. At least the lower half of the sedimentary column (the part sampled by the scientific drilling) formed in an euxinic environment. This argues against a persistent early playa, although evaporative accumulation of brine was inevitable. When the rate of resurgance was rapid relative to sedimentary infilling, the lake would have been deep (i.e., bordered by bedrock rather than sedimentary fans). The geomorphic evolution of the Creede caldera and its watershed tracks a two-phase topographic history, the first the Oligocene through Miocene, and the second for Pliocene to the recent. In Oligocene time, the San Juan volcanic field was a hydrologically immature, gently undulating, and outward sloping, constructional volcanic plateau straddling the ancient Continental Divide. West of the Creede caldera, a dendritic drainage discharged northeastward into ancestral Cebolla Creek (a tributary of the ancestral Gunnison River) through an early stage of the Clear Creek graben in the vicinity of Spring Creek Pass. Miocene basalt choked, but did not reconstruct, the drainage. By the end of Miocene time a mature topography of moderate relief developed, exposing some of the higher ores in the Creede district to weathering. In the late Miocene-early Pliocene time the San Juan Mountains were uplifted and titled eastward; the ancestral Rio Grande was revitalized and cut deeply into the older terrain, excavating much of the accessible sediment from the moat of the Creede caldera and exposing successively lowe levels in the Creede district to oxidation. Simultaneously, the southeast end of the Clear Creek graben was reactivated and breached the southwest wall of the Creede caldera. The rejuvenated Rio Grande captured the formerly northeast-directed headwaters of ancestral Cebolla Creek, shifting more than 1000 km 2 from the Pacific-directed drainage to the Atlantic. The water budget for ancient Lake Creede was strictly limited by the early stages of the fist geomorphic cycle; the modern water budget is the product of the second cycle.

Colorado

Tectonic evolution of the southern Gulf of Mexico

A detailed magnetic survey in the southern Gulf of Mexico shows subdued irregular magnetic anomalies that are similar in wave length to those attributed to sea-floor spreading on present-day oceanic rises. The small amplitude of these anomalies, about 75γ, would be compatible with an oceanic basalt source at a depth of 10 km, and previous seismic refraction studies in this area have shown that layer 2 of the oceanic crust (presumed to be basalt) does indeed lie at approximately that depth. Palynomorphs in Deep Sea Drilling Project samples of cap-rock material from the Sigsbee Knolls have shown the associated salt to be Jurassic. A crustal model, based on seismic refraction evidence and on new gravity data, suggests that, whereas the salt-dome belt of the southern gulf is underlain by a thick layer with a density of 2.2 g per cm 3 (presumed to be halite rock), adjacent deeper areas of the basin seem to lack this low density layer. In northeastern Mexico, Triassic red beds fill grabens that are correlative with the Newark Group of the Atlantic Coast and suggest that the Gulf of Mexico originated at the time of the initial rifting of the North Atlantic. When the gulf was about half-opened during the Jurassic, oceanic circulation was restricted; and thick deep-basin evaporite deposits, analogous to those found in the Mediterranean Sea by the Deep Sea Drilling Project, were laid down. Further opening established normal salinity and led to the development of salt-free areas of oceanic crust that separated the Sigsbee Escarpment (together with adjacent ridges of offshore Mexico) from the Sigsbee Knolls and salt domes of Cuba. The subsequent structural evolution of the Gulf of Mexico basin is believed to have been mainly a result of interaction between it and tectonic plates of the Pacific area. After opening of the gulf, subduction began along the Cuban arc, where Atlantic rifting had created a nearly uninterrupted tract of oceanic crust extending from the Pacific between Yucatán and Colombia. Then, folding and gravity sliding associated with Laramide deformation on the west side of the gulf led to salt anticlines that underlie the ridges offshore from Mexico. Except for continued diapirism and subsidence associated with sediment loading, present tectonic activity is confined to the southwestern corner of the gulf, where volcanism and intermediate-focus earthquakes are a distant manifestation of subduction along the Middle America Trench on the Pacific side of Mexico.

Gulf of Mexico

Evolution of the yardangs at Rogers Lake, California

Yardangs are streamlined, wind-eroded hills common to most deserts. Yardangs at Rogers Lake, Mojave Desert, California, have streamlined forms characteristic of objects eroded by moving fluids, a teardrop shape that approaches an ideal 1:4 width-to-length ratio. In wind-tunnel simulations, miniature forms of various shapes changed sequentially by (1) erosion of the windward corners, (2) erosion of the windward slope, (3) erosion of the leeward corners and flanks, and (4) erosion of the leeward slope. Prominent mechanisms in yardang evolution apparently are abrasion at the windward end and deflation and reverse air flow near the middle and at the downstream end. Width-to-length ratios of yardangs are grossly similar to those of some fluvial and glacial streamlined landforms. The low kinetic energy of wind relative to ice and water, the erosional resistance to wind of most rocks, the rarity of long-term, unidirectional winds, and the presence of running water, topographic roughness, and vegetation all limit the abundance of yardangs.

Geological Society of America Bulletin

The late Cenozoic evolution of the Tuolumne River, central Sierra Nevada, California

Erosional remnants of volcanic rock deposited in a lO-m.y.-old channel of the Tuolumne River permit its partial reconstruction. Projection of the reconstructed channel west to the Central Valley and east to the range crest, together with several assumptions about the position of the hinge line and changes in channel gradient, allows estimates of the amount of uplift at the range crest during the past 10 m.y. At Tioga Pass, this amounts to as much as 1,830 m, as compared to the 2,150 m estimated in an earlier study for Deadman Pass at the San Joaquin River 30 km to the south. Comparison of the geometry of these river systems leads to the conclusion that 10 m.y. ago an ancestral range of hills occupied the present site of the Sierran crest, and, although of relatively moderate relief, it was a barrier to westward drainage even before late Cenozoic uplift. At that time, the San Joaquin River was apparently the only river flowing westward across the range from well south of Mount Whitney north to Sonora Pass. The Tuolumne River evidently never extended east of this range. Comparison of the ancient channel with the modern channel of the Tuolumne River permits analysis of the later evolution of the river system and the development of Hetch Hetchy Valley and the Grand Canyon of the Tuolumne. At Rancheria Mountain, where the volcanic "dam" in the ancient channel was highest, the river was forced to shift laterally southward around the dam and adjacent to the volcanic infilling, and start its new channel in granitic bedrock. Near Rancheria Mountain, as much as 1,525 m of new channel incision has taken place in the past 10 m.y., and the modern channel is about 915 m lower than the abandoned channel. An undetermined amount of this downcutting was from glacial erosion. The Tuolumne river system provides no direct evidence for timing the onset of uplift, but the shape of the lO-m.y.-old channel at Rancheria Mountain suggests that uplift had been underway for some time before the volcanic infilling. This timing is compatible with evidence from the upper San Joaquin River. Hetch Hetchy Valley on the Tuolumne is a much "fresher" glaciated valley than is Yosemite Valley. Hetch Hetchy was filled to the brim with glacial ice as recently as 15,000-20,000 yr ago (Tioga glaciation), whereas Yosemite Valley probably has not been filled for 750,000 yr or more (Sherwin glaciation). Thus the upper reaches of Yosemite Valley cliffs have been shaped by spalling rather than by glacial scour and are much more irregular than those in Hetch Hetchy.

California