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At least 289 records · Page 16Linked to original sources

Persistence of native and exotic plants 10 years after prairie reconstruction

Prairie reconstructions are a critical component of preservation of the imperiled tallgrass prairie ecosystem in the Midwestern United States. Sustainability of this endeavor depends on establishment of persistent cover of planted native species and resistance to noxious weeds. The goal of this study was to understand the influence of early reconstruction practices on long-term outcomes. Twelve replicates of three planting methods (dormant-season broadcast, growing-season broadcast, and growing-season drill) and three seed mix richness levels (10, 20, or 34 species), fully crossed in a completely randomized design were planted in 2005 on nine former agricultural fields located in Iowa and Minnesota. Cover by species was estimated in 2005–2007, 2010, and 2015. In 2015, cover of planted species, native nonplanted species, and exotic species were similar to those recorded in 2010. Cover of the noxious weed Cirsium arvense had also declined by an average of 49% without herbicide from a peak in 2007 to low stable levels from 2010 to 2015. Richness of planted forbs, on the other hand, were still increasing in high-richness broadcast treatments (e.g. 17–59% increase 2010–1015 in Minnesota). Two results in 2015 are reasons for concern: cover of planted species is only slightly over 50% in both Minnesota and Iowa, though with forbs still increasing, this may improve; and the cool-season exotic grasses Poa pratensis and Bromus inermis are increasing at both Minnesota and Iowa sites. Control of these invasive grasses will be necessary, but care will be needed to avoid negative impacts of control methods on natives.

Restoration Ecology

Rapid colonization of a Hawaiian restoration forest by a diverse avian community

Deforestation of tropical forests has led to widespread loss and extirpation of forest bird species around the world, including the Hawaiian Islands which have experienced a dramatic loss of forests over the last 200–800 years. Given the important role birds play in forest ecosystem functions via seed dispersal and pollination, a bird community's response to forest restoration is an important measure of the success of such conservation actions. We evaluated the bird response to reforestation at an important bird sanctuary, Hakalau Forest National Wildlife Refuge, Hawai′i Island, using 26 years of bird count data. We show that most species from within the diverse avian community increased significantly, but species colonized the restoration forest at different rates. Distance from intact forest and time since restoration were both important predictors of colonization rate, interacting such that for most species it took more time to colonize areas farther from the intact forest. In addition, both forest cover and understory diversity helped to explain bird densities, but the effect varied among species, suggesting that different habitat requirements may help drive variation in colonization rates. This article provides the first detailed evaluation of how a diverse community of birds has responded to one of the largest, ongoing reforestation projects in Hawai′i.

Hawai′i

Role of a naturally varying flow regime in Everglades restoration

The Everglades is a low-gradient floodplain predominantly on organic soil that undergoes seasonally pulsing sheetflow through a network of deepwater sloughs separated by slightly higher elevation ridges. The seasonally pulsing flow permitted the coexistence of ridge and slough vegetation, including the persistence of productive, well-connected sloughs that seasonally concentrated prey and supported wading bird nesting success. Here we review factors contributing to the origin and to degradation of the ridge and slough ecosystem in an attempt to answer “How much flow is needed to restore functionality”? A key restoration objective is to increase sheetflow lost during the past century to reestablish interactions between flow, water depth, vegetation production and decomposition, and transport of flocculent organic sediment that build and maintain ridge and slough distinctions. Our review finds broad agreement that perturbations of water level depth and its fluctuations were primary in the degradation of landscape functions, with critical contributions from perturbed water quality, and flow velocity and direction. Whereas water levels are expected to be improved on average across a range of restoration scenarios that replace between 79 and 91% of predrainage flows, the diminished microtopography substantially decreases the probability of timely improvements in some areas whereas others that retain microtopographic differences are poised for restoration benefits. New advances in predicting restoration outcomes are coming from biophysical modeling of ridge–slough dynamics, system-wide measurements of landscape functionality, and large-scale flow restoration experiments, including active management techniques to kick-start slough regeneration.

Florida

Passive restoration following ungulate removal in a highly disturbed tropical wet forest devoid of native seed dispersers

Overabundant ungulate populations can alter forests. Concurrently, global declines of seed dispersers may threaten native forest structure and function. On an island largely devoid of native vertebrate seed dispersers, we monitored forest succession for 7 years following ungulate exclusion from a 5-ha area and adjacent plots with ungulates still present. We observed succession from open scrub to forest and understory cover by non-native plants declined. Two trees, native Hibiscus tiliaceus and non-native Leucaena leucocephala , accounted for most forest regeneration, with the latter dominant. Neither species is dependent on animal dispersers nor was there strong evidence that plants dependent on dispersers migrated into the 5-ha study area. Passive restoration following ungulate removal may facilitate restoration, but did not show promise for fully restoring native forest on Guam. Restoration of native forest plants in bird depopulated areas will likely require active outplanting of native seedlings, control of factors resulting in bird loss, and reintroduction of seed dispersers.

Guam

Long-term trends in restoration and associated land treatments in the southwestern United States

Restoration treatments, such as revegetation with seeding or invasive species removal, have been applied on U.S. public lands for decades. Temporal trends in these management actions have not been extensively summarized previously, particularly in the southwestern United States where invasive plant species, drought, and fire have altered dryland ecosystems. We assessed long-term (1940–2010) trends in restoration using approximately 4,000 vegetation treatments conducted on Bureau of Land Management lands across the southwestern United States. We found that since 1940, the proportions of seeding and vegetation/soil manipulation (e.g. vegetation removal or plowing) treatments have declined, while the proportions of prescribed burn and invasive species treatments have increased. Treatments in pinyon-juniper and big sagebrush communities declined in comparison to treatments in desert scrub, creosote bush, and riparian woodland communities. Restoration-focused treatment objectives increased relative to resource extraction objectives. Species richness and proportion of native species used in seeding treatments also increased. Inflation-adjusted costs per area rose 750% for vegetation/soil manipulation, 600% for seeding, and 400% for prescribed burn treatments in the decades from 1981 to 2010. Seeding treatments were implemented in warmer and drier years when compared to the climate conditions of the entire study period and warmer and wetter years relative to several years before and after the treatment. These results suggest that treatments over a 70-year period on public lands in the southwestern United States are shifting toward restoration practices that are increasingly large, expensive, and related to fire and invasive species control.

Restoration Ecology

Northern tamarisk beetle (Diorhabda carinulata) and tamarisk (Tamarix spp.) interactions in the Colorado River basin

Northern tamarisk beetles (Diorhabda carinulata) were released in the Upper Colorado River Basin in the United States in 2004–2007 to defoliate introduced tamarisk shrubs (Tamarix spp.) in the region’s riparian zones. The primary purpose was to control the invasive shrub and reduce evapotranspiration (ET) by tamarisk in an attempt to increase stream flows. We evaluated beetle–tamarisk interactions with MODIS and Landsat imagery on 13 river systems, with vegetation indices used as indicators of the extent of defoliation and ET. Beetles are widespread and exhibit a pattern of colonize–defoliate–emigrate, so that riparian zones contain a mosaic of completely defoliated, partially defoliated, and refoliated tamarisk stands. Based on satellite data and ET algorithms, mean ET before beetle release (2000–2006) was 416 mm/year compared to postrelease (2007–2015) ET of 355 mm/year (p<0.05) for a net reduction of 61 mm/year. This is lower than initial literature projections that ET would be reduced by 300–460 mm/year. Reasons for the lower-than-expected ET reductions are because baseline ET rates are lower than initially projected, and percentage ET reduction is low because tamarisk stands tend to regrow new leaves after defoliation and other plants help maintain canopy cover. Overall reductions in tamarisk green foliage during the study are 21%. However, ET in the Upper Basin has shown a steady decline since 2007 and equilibrium has not yet been reached. Defoliation is now proceeding from the Upper Basin into the Lower Basin at a rate of 40 km/year, much faster than initially projected.

Colorado River

Regeneration and invasion of cottonwood riparian forest following wildfire

Populus deltoides is considered to be a weak resprouter and highly susceptible to wildfire, but few post-wildfire studies have tracked P. deltoides response and resprouting within the Great Plains of North America. Following a wildfire in southwestern Kansas, U.S.A., we surveyed burned and unburned areas of a cottonwood riparian forest along the Cimarron River that included a major understory invader, tamarisk ( Tamarix ramosissima Ledeb.). We tested the following hypotheses, which are consistent with the current understanding of P. deltoides response to wildfire in the Great Plains: (1) regeneration of P. deltoides will be low in areas burned by the wildfire; (2) the number of dead P. deltoides individuals will be greater in the wildfire than unburned areas; and (3) tamarisk regeneration will be higher than P. deltoides regeneration in the wildfire areas because tamarisk is considered a stronger resprouter. We found evidence contrary to two of our hypotheses 3 years following the wildfire. (1) P. deltoides regeneration was high following the wildfire, averaging 692 individuals/ha. (2) The number of dead mature cottonwood trees was greater in wildfire plots than in unburned plots. (3) There was more P. deltoides regeneration than tamarisk regeneration following wildfire. These findings, which diverge from the majority of studies examining P. deltoides regeneration in the Great Plains, suggest that differing local environmental and forest stand conditions, coupled with the timing and intensity of the fire, could be important determinants of riparian forest species' responses to wildfire.

Kansas

Waterbird communities and seed biomass in managed and reference-restored wetlands in the Mississippi Alluvial Valley

The Natural Resources Conservation Service (NRCS) commenced the Migratory Bird Habitat Initiative (MBHI) in summer 2010 after the April 2010 Deepwater Horizon oil spill in the Gulf of Mexico. The MBHI enrolled in the program 193,000 ha of private wet- and cropland inland from potential oil-impaired wetlands. We evaluated waterfowl and other waterbird use and potential seed/tuber food resources in NRCS Wetland Reserve Program easement wetlands managed via MBHI funding and associated reference wetlands in the Mississippi Alluvial Valley of Arkansas, Louisiana, Mississippi, and Missouri. In Louisiana and Mississippi, nearly three times more dabbling ducks and all ducks combined were observed on managed than reference wetlands. Shorebirds and waterbirds other than waterfowl were nearly twice as abundant on managed than referenced wetlands. In Arkansas and Missouri, managed wetlands had over twice more dabbling ducks and nearly twice as many duck species than reference wetlands. Wetlands managed via MBHI in Mississippi and Louisiana contained ≥1.3 times more seed and tuber biomass known to be consumed by waterfowl than reference wetlands. Seed and tuber resources did not differ between wetlands in Arkansas and Missouri. While other studies have documented greater waterbird densities on actively than nonmanaged wetlands, our results highlighted the potential for initiatives focused on managing conservation easements to increase waterbird use and energetic carrying capacity of restored wetlands for waterbirds.

Arkansas, Louisiana, Mississippi, Missouri

Gauging resource exploitation by juvenile Chinook salmon (Oncorhynchus tshawytscha) in restoring estuarine habitat

In the context of delta restoration and its impact on salmonid rearing, success is best evaluated based on whether out-migrating juvenile salmon can access and benefit from suitable estuarine habitat. Here, we integrated 3 years of post-restoration monitoring data including habitat availability, invertebrate prey biomass, and juvenile Chinook salmon ( Oncorhynchus tshawytscha ) physiological condition to determine whether individuals profited from the addition of 364 ha of delta habitat in South Puget Sound, Washington, United States. Productivity in the restored mudflat was comparable to reference sites 3 years after dike removal, surpassing a mean total of 6 million kJ energy from invertebrate prey. This resulted from the development of a complex network of tidal channels and a resurgence in dipteran biomass that was unique to the restoration area. Consequently, a notable shift in invertebrate consumption occurred between 2010 and 2011, whereby individuals switched from eating primarily amphipods to dipteran flies; however, dietary similarity to the surrounding habitat did not change from year to year, suggesting that this shift was a result of a change in the surrounding prey communities. Growth rates did not differ between restored and reference sites, but catch weight was positively correlated with prey biomass, where greater prey productivity appeared to offset potential density-dependent effects. These results demonstrate how the realized function of restoring estuarine habitat is functionally dependent. High prey productivity in areas with greater connectivity may support healthy juvenile salmon that are more likely to reach the critical size class for offshore survival.

Washington

Enhanced invertebrate prey production following estuarine restoration supports foraging for multiple species of juvenile salmonids (Oncorhynchus spp.)

Estuaries provide crucial foraging resources and nursery habitat for threatened populations of anadromous salmon. As such, there has been a global undertaking to restore habitat and tidal processes in modified estuaries. The foraging capacity of these ecosystems to support various species of out-migrating juvenile salmon can be quantified by monitoring benthic, terrestrial, and pelagic invertebrate prey communities. Here, we present notable trends in the availability of invertebrate prey at several sites within a restoring large river delta in Puget Sound, Washington, U.S.A. Three years after the system was returned to tidal influence, we observed substantial additions to amphipod, copepod, and cumacean abundances in newly accessible marsh channels (from 0 to roughly 5,000–75,000 individuals/m 2 ). In the restoration area, terrestrial invertebrate colonization was dependent upon vegetative cover, with dipteran and hymenopteran biomass increasing 3-fold between 1 and 3 years post-restoration. While the overall biodiversity within the restoration area was lower than in the reference marsh, estimated biomass was comparable to or greater than that found within the other study sites. This additional prey biomass likely provided foraging benefits for juvenile Chinook, chum, and coho salmon. Primary physical drivers differed for benthic, terrestrial, and pelagic invertebrates, and these invertebrate communities are expected to respond differentially depending on organic matter exchange and vegetative colonization. Restoring estuaries may take decades to meet certain success criteria, but our study demonstrates rapid enhancements in foraging resources understood to be used for estuary-dependent wildlife.

Washington

Amendments fail to hasten biocrust recovery or soil stability at a disturbed dryland sandy site

In most drylands, biological soil crusts (biocrusts), an assemblage of lichens, bryophytes, fungi, green algae, and cyanobacteria, are critical to healthy ecosystem function. However, they are extremely sensitive to disturbance and attempts to facilitate their recovery have had variable success. In this study, we applied soil amendments designed to improve soil surface stability and accelerate biocrust recovery on an area disturbed by oil/gas exploration vehicles. Treatments included: 1) Control (one time water only); 2) Biocrust‐only: biocrust inoculum + nutrients in water; 3) Polyacrylamide gels (PAM; which are known to stabilize soils) + biocrust inoculum + nutrients in water; 4) Gypsum + biocrust inoculum + nutrients in water; and 5) Saline (NaCl) solution + biocrust inoculum + nutrients in water. Only the NaCl treatment showed any effects on soil properties and these were only short‐term. These effects included an increase in soil strength and a reduction in soil aggregate stability, unsaturated hydraulic conductivity (K h ), and cyanobacterial biomass. The inoculated biocrust material failed to develop and even after 10 years, there was only a very low natural recolonization of the plots. These results show that inoculating soils or applying these levels of soil amendments does not guarantee recovery of soil stability or biocrust, and that some sites are unlikely to recover without assistance. Thus, there is a need for more research into ways to enhance soil stability and identify the factors limiting biocrust establishment.

Restoration Ecology

Long‐term plant community trajectories suggest divergent responses of native and non‐native perennials and annuals to vegetation removal and seeding treatments

Land managers frequently apply vegetation removal and seeding treatments to restore ecosystem function following woody plant encroachment, invasive species spread, and wildfire. However, the long‐term outcome of these treatments is unclear due to a lack of widespread monitoring. We quantified how vegetation removal (via wildfire or management) with or without seeding and environmental conditions related to plant community composition change over time in 491 sites across the intermountain western United States. Most community metrics took over 10 years to reach baseline conditions posttreatment, with the slowest recovery observed for native perennial cover. Total cover was initially higher in sites with seeding after vegetation removal than sites with vegetation removal alone, but increased faster in sites with vegetation removal only. Seeding after vegetation removal was associated with rapidly increasing non‐native perennial cover and decreasing non‐native annual cover. Native perennial cover increased in vegetation removal sites irrespective of seeding and was suppressed by increasing non‐native perennial cover. Seeding was associated with higher non‐native richness across the monitoring period as well as initially higher, then declining, total and native species richness. Several cover and richness recovery metrics were positively associated with mean annual precipitation and negatively associated with mean annual temperature, whereas relationships with weather extremes depended on the lag time and season. Our results suggest that key plant groups, such as native perennials and non‐native annuals, respond to restoration treatments at divergent timescales and with different sensitivities to climate and weather variation.

Utah

Adaptive management assists reintroduction as higher tides threaten an endangered salt marsh plant

In theory, extirpated plant species can be reintroduced and managed to restore sustainable populations. However, few reintroduced plants are known to persist for more than a few years. Our adaptive‐management case study illustrates how we restored the endangered hemiparasitic annual plant, Chloropyron maritimum subsp. maritimum (salt marsh bird's beak), to Sweetwater Marsh, San Diego Bay National Wildlife Refuge, California, United States, and used monitoring and experimentation to identify the factors limiting the reintroduced population. After extirpation in 1988, reintroduction starting that year led to a resilient, genetically diverse population in 2016 (a “boom” of approximately 14,000) that rebounded from a “bust” (62 in 2014). Multiple regressions attributed 82% of the variation in population counts to tidal amplitude, rainfall, and temperature. Populations of salt marsh bird's beak crashed when the diurnal tide range peaked during the 18.6‐year lunar nodal cycle (a rarely considered factor that periodically added approximately 12 cm to tidal ranges). We explain booms as follows: During smaller tidal amplitudes , above‐average rainfall could desalinize upper intertidal soils and stimulate salt marsh bird's beak germination. Then, moderate temperature in May favors growth to reproduction in June. In addition, salt marsh bird's beak needs a short and open canopy of native perennial plants, with roots to parasitize (not non‐native annual grass pseudohosts) and nearby upland soil for a preferred pollinator, ground‐burrowing bees. Although our reintroduced salt marsh bird's beak population is an exceptional case of persistence, this rare species‐specific environmental and biological requirement makes it vulnerable to rising sea levels and global warming.

California

Landscape and organismal factors affecting sagebrush-seedling transplant survival after megafire restoration

Larger and more frequent disturbances are motivating efforts to accelerate recovery of foundational perennial species by focusing efforts into establishing island patches to sustain keystone species and facilitate recovery of the surrounding plant community. Evaluating the variability in abiotic and biotic factors that contribute to differences in survival and establishment can provide useful insight into the relative importance of these factors. In the western United States, severe degradation of the sagebrush steppe has motivated substantial efforts to restore native perennial cover, but success has been mixed. In this study, we evaluated survival of >3000 sagebrush seedlings transplanted on 12 patches totaling of 650 ha within a 113,000 ha burn area, and related the survival to organismal and sub‐taxonomic traits, and to landscape variables. Big sagebrush has high intraspecific diversity attributed to subspecies and cytotypes identifiable through UV‐induced fluorescence, length:width of leaves, or genome size (ploidy). Of these organismal traits, survival was related only to UV fluorescence, and then only so when landscape variables were excluded from analyses. The most significant landscape variable affecting survival was soil taxonomic subgroup, with much lower survival where buried restrictive layers reduce deep water infiltration. Survival also decreased with greater slope steepness, exotic annual grass cover, and burn severity. Survival was optimal where perennial bunchgrasses comprised 8‐14% of total cover. These soil, topographic, and community condition factors revealed through monitoring of landscape‐level treatments can be used to explain the success of plantings and to strategically plan future restoration projects.

Restoration Ecology

Carbon accumulation and vertical accretion in a restored vs. historic salt marsh in southern Puget Sound, Washington, United States

Few comparisons exist between vertical accretion (VA) and carbon accumulation rates (CARs), in restored vs. historic (i.e., reference) marshes. Here we compare these processes in a formerly diked, sparsely vegetated, restored salt marsh (Six Gill Slough, SG), whose surface is subsided relative to the tidal frame, to an adjacent, relatively pristine, historic salt marsh (Animal Slough, AS). Six sediment cores were collected at both AS and SG ~six years after restoration. Cores were analyzed for bulk density, % loss of ignition, % organic carbon, and 210Pb. We found that sharp changes in bulk density in surface layers of SG cores were highly reliable markers for the onset of restoration. The mean VA since restoration at SG (0.79 (sd=0.29) cm yr-1) was ~twice that of AS (0.41 (sd=0.16) cm yr-1). In comparison, the VA at AS over 50 years was 0.30 (sd=0.09) cm yr-1. VA consisted almost entirely of inorganic sediment at SG whereas at AS it was ~55%. Mean CARs at SG were somewhat greater than at AS, but the difference was not significant due to high variability (SG: 81 - 210 g C m-2 yr-1; AS: 115 - 168 g C m-2 yr-1). The mean CAR at AS over the past 50 years was 118 (sd=23) g C m-2 yr-1. This study demonstrates that a sparsely vegetated, restored salt marsh can quickly begin to accumulate carbon and that historic and restored marshes can have similar CARs despite highly divergent formation processes.

Washington

Hydroseeding tackifiers and dryland moss restoration potential

Tackifiers are long‐chain carbon compounds used for soil stabilization and hydroseeding and could provide a vehicle for biological soil crust restoration. We examined the sensitivity of two dryland mosses, Bryum argenteum and Syntrichia ruralis, to three common tackifiers ‐ guar, psyllium, and polyacrylamide (PAM) ‐ at 0.5x, 1.0x, and 2.0x of recommended (x) concentrations for erosion control and revegetation. We measured moss shoot, gemma, and protonema production as well as moss organic matter and bound sand masses as indicators of growth and soil holding ability. We tested sand and tackifier chemistry to investigate potential nutrient and toxicant potential on moss growth. Groups of ten fragments from field‐collected mosses were grown on sand in open petri dishes arranged in a growth chamber in replicated blocks containing each tackifier and concentration combination plus a distilled water control. Bryum ( n =10) and Syntrichia ( n =9) growth were measured at the end of six and five weeks, respectively. Overall model tests yielded statistically significant results ( p <0.001) for every variable in each species. When compared to water, guar tended to decrease growth, psyllium tended to increase growth, and PAM's effects were generally neutral to positive. Within tackifier types, increasing concentrations of guar tended to decrease growth, while increasing concentrations of psyllium tended to increase growth. Changes in PAM concentrations had little effect on growth. Increases in guar and psyllium lowered pH and increased P and K. Psyllium and PAM yielded promising results as potential agents of dispersal and adherence of dryland mosses in field restoration.

Restoration Ecology

Passive restoration of vegetation and biological soil crusts following 80 years of exclusion from grazing across the Great Basin

Restoration targets for biological soil crusts are largely unknown. We surveyed seven 80‐year‐old grazing exclosures across northern Nevada for biocrusts to quantify reference conditions at relatively undisturbed sites. Exclosures were associated with the following plant communities: Wyoming big sagebrush, black sagebrush, and areas co‐dominated by winterfat and Wyoming big sagebrush. Cover of biocrusts and shrubs were generally higher than other plant groups at these sites, regardless of being inside or outside of the exclosures, suggesting these groups make up most of the native flora across the region. Important in forming soil structure, cyanobacteria of the order Oscillatoriales were less abundant and diverse in black sagebrush communities. Grazing had a negative effect on the abundance of Oscillatoriales but not the number of algal taxa, including cyanobacteria. Abundance of light algal crusts were not influenced by plant community or grazing. Dark algal crusts were generally less abundant on grazed sites. Influences of plant community and grazing were most apparent when accounting for reproductive rates of lichens and mosses based on establishment mechanisms. Abundance of shrubs, perennial grasses, Oscillatoriales, fast reproducing biocrusts and the number of algal and cyanobacterial taxa, varied by plant community, suggesting that restoration should be plant community specific. We demonstrate the affinity of rapidly reproducing biocrusts for winterfat‐Wyoming big sagebrush co‐dominated plant communities, regardless of grazing pressure. Across sites, the effects of grazing were most evident on the abundance of Oscillatoriales and slowly reproducing biocrusts following 80 years of cessation from grazing.

Great Basin

Species-specific responses to wetland mitigation among amphibians in the Greater Yellowstone Ecosystem

Habitat loss and degradation are leading causes of biodiversity declines, therefore assessing the capacity of created mitigation wetlands to replace habitat for wildlife has become a management priority. We used single season occupancy models to compare the occurrence of larvae of four species of pond‐breeding amphibians in wetlands created for mitigation, wetlands impacted by road construction, and unimpacted reference wetlands along a highway corridor in the Greater Yellowstone Ecosystem, United States. Created wetlands were shallow and had less aquatic vegetation and surface area than impacted and reference wetlands. Occupancy of barred tiger salamander ( Ambystoma mavortium ) and boreal chorus frog ( Pseudacris maculata ) larvae was similar across wetland types, whereas boreal toads ( Anaxyrus boreas ) occurred more often in created wetlands than reference and impacted wetlands. However, the majority of created wetlands (>80%) dried partially or completely before amphibian metamorphosis occurred in both years of our study, resulting in heavy mortality of larvae and, we suspect, little to no recruitment. Columbia spotted frogs ( Rana luteiventris ), which require emergent vegetation that is not common in newly created wetlands, occurred commonly in impacted and reference wetlands but were found in only one created wetland. Our results show that shallow created wetlands with little aquatic vegetation may be attractive breeding areas for some amphibians, but may result in high mortality and little recruitment if they fail to hold water for the entire larval period.

Yellowstone