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The Rocky Mountain population of the western Canada goose: Its distribution, habitats, and management

The western Canada goose ( Branta canadensis moffitti ) was divided into a Rocky Mountain population (RMP) and a Pacific population (PP) on the basis of band recovery patterns examined in this study and recovery data from other investigators. Habitat information obtained from nine cooperating wildlife agencies within the RMP's range provided a base line for evaluating future changes in nesting, molting, and wintering areas. The habitat inventory indicated that none of the seasonal habitats were currently limiting the size of the RMP. The RMP's range is divided into 15 reference areas and these are briefly described. Past studies of Canada geese in the Intermountain Region are reviewed. Topics covered in the discussion of breeding biology are nesting chronology, spring population composition, breeding age, clutch size, nesting success. artificial nesting structures, and gosling survival. Much of the mortality of Canada geese occurs before the birds are fledged. Man-made nesting structures reduce losses during incubation. but research is needed on the relations between brooding sites and gosling survival. Some western Canada geese, mainly prebreeders and unsuccessful nesters, make molt migrations to and from molting areas during and after the brood-rearing season. More than half of these molt-migrants are yearlings too young to nest; there are indications that even some successful nesters leave nesting areas to molt before the fledging of their offspring. Geese 2 years old or older may serve as guides to traditional molting areas for the first-time migrants (i.e., yearlings). Lack of disturbance appears to influence selection of specific molting areas within the nesting range of moffitti , whereas movements of molters out of the Intermountain Region may be related to the evolution of this subspecies. Apparently. molters of both the PP and RMP that leave the Region go to the Northwest Territories of Canada. Although the taxonomic status of moffitti as related to the giant Canada goose ( B. c. maxima ) is unclear. these two subspecies are closely related. as evidenced by similar molt migrations to subarctic Canada. similar blood serum proteins. and only dinal differences in body size ,!!nd color. Mean annual survival rates for birds banded on nesting areas averaged 53 ± 2% ( X̿ ± SE) for immatures and 64 ± 1 % for adults. Mean annual survival rates of adults captured on molting areas averaged 70 ± 1 %. Sport hunting accounts for more than 86% of the mortality of fledged Rocky Mountain geese. and hunting may limit the population's growth. Because the number of waterfowl hunters in the Rocky Mountain West is increasing, the continued expansion or future maintenance of the RMP may require more restrictive hunting regulations. Other management recommendations include the refinement and standardization of spring and winter aerial surveys, and more accurate age and sex determinations when geese are banded and color-marked.

Alberta, Arizona, California, Colorado, Idaho, Mon

Effects of leg flags on nest survival of four species of Arctic‐breeding shorebirds

Marking wild birds is an integral part of many field studies. However, if marks affect the vital rates or behavior of marked individuals, any conclusions reached by a study might be biased relative to the general population. Leg bands have rarely been found to have negative effects on birds and are frequently used to mark individuals. Leg flags, which are larger, heavier, and might produce more drag than bands, are commonly used on shorebirds and can help improve resighting rates. However, no one to date has assessed the possible effects of leg flags on the demographic performance of shorebirds. At seven sites in Arctic Alaska and western Canada, we marked individuals and monitored nest survival of four species of Arctic‐breeding shorebirds, including Semipalmated Sandpipers ( Calidris pusilla ), Western Sandpipers ( C. mauri ), Red‐necked Phalaropes ( Phalaropus lobatus ), and Red Phalaropes ( P. fulicarius ). We used a daily nest survival model in a Bayesian framework to test for effects of leg flags, relative to birds with only bands, on daily survival rates of 1952 nests. We found no evidence of a difference in nest survival between birds with flags and those with only bands. Our results suggest, therefore, that leg flags have little effect on the nest success of Arctic‐breeding sandpipers and phalaropes. Additional studies are needed, however, to evaluate the possible effects of flags on shorebirds that use other habitats and on survival rates of adults and chicks.

Journal of Field Ornithology

Wintering site fidelity and movement patterns of Western Sandpipers Calidris mauri in the San Francisco Bay estuary

Western Sandpipers Calidris mauri are the most numerous shorebird species in the San Francisco Bay estuary during winter. A sample of 106 Western Sandpipers was captured in mist nets and radio-marked with 1-g transmitters to examine their wintering site fidelity and movements. Differences in distances moved, home range extent and core area size were examined by age, sex, season, site, time of day and tide. All birds remained in the south San Francisco Bay region during winter and exhibited strong site fidelity, with a mean home range of 22.0 km 2 or only 8% of the study area. First-year birds had larger home ranges (26.6 ± 3.6 km 2 ) than adults (17.2 ± 2.5 km 2 ) in winter, but home range sizes of males and females were not significantly different in any period. Home range sizes were similar between seasons, but core areas were smaller in spring (6.3 ± 1.2 km 2 ) than in early (9.6 ± 4.0 km 2 ) or late (11.6 ± 1.6 km 2 ) winter. Movements and home range size were similar for radio-marked birds located during day and night. The high degree of regional and local site fidelity demonstrated that the mixture of natural mud fiats and artificial salt ponds in southern San Francisco Bay provided sufficient resources for large wintering populations of Western Sandpipers.

Ibis

Reoccurrence of 'Ōma'o in leeward woodland habitat and their distribution in alpine habitat on Hawai'i island

The endemic solitaire, 'Oma'o (Myadestes obscurus), is common in windward forests of Hawai'i Island, but has been historically extirpated from leeward forests. The last detections of 'Oma'o on the leeward side of the island were in woodland habitat on the western flank of Mauna Loa in 1978. 'Oma'o were detected in woodland habitat in relatively low densities during a 2010 forest bird survey of Hawai'i Volcanoes National Park. The source of the population is unknown. It is probable they originated from a documented but unsurveyed population of 'Oma'o in scrub alpine lava. Alternatively, the birds may have persisted undetected for nearly 35 years, or expanded from windward mesic forests on southeast Mauna Loa. There is no evidence 'Oma'o recolonized the wet mesic forests of leeward Mauna Loa. The 'Oma'o can occupy diverse native habitats compared to other species in the Hawai'i Myadestes genus, of which most species are now extinct. The connectivity of each population is not understood but we assume there are significant geographic, physiological, and behavioral barriers for scrub alpine and wet mesic forest populations. The expansion of 'Oma'o to leeward woodlands is encouraging as the species is Hawai'i Island's last native frugivore capable of dispersing small and medium sized seeds of rare angiosperms, and could have an important role in re-establishing ecosystem function.

Hawaii

Notes on longevity and flightlessness in bristle-thighed curlews

Bristle-thighed Curlews ( Numenius tahitiensis ) are among the least-studied shorebirds in the world. They have a highly restricted breeding range in western Alaska and winter exclusively on remote islands in the central and south Pacific (AOU 1983, Kessel 1989). The status of these birds is not known, but they are considered to be rare throughout their range (Johnsgard 1981). We describe two unusual traits of Bristle-thighed Curlews - exceptional longevity and flightlessness during molt-and discuss their significance to the species' migration and nonbreeding ecology.

The Auk

Grassland bird communtiy response to large wildfires

We studied breeding season communities of grassland birds on short-grass and mixed-grass prairie sites during the second and third breeding seasons following two large wildfires in March 2006 in the Texas panhandle, USA. There was an apparent temporary shift in avian community composition following the fires due to species-specific shifts associated with life-history traits and vegetation preferences. Species that prefer sparse vegetation and bare ground on short-grass sites, such as Horned Lark ( Eremophila alpestris ), benefited from wildfires, while others, such as Western Meadowlark ( Sturnella neglecta ), that prefer more dense vegetation, were negatively impacted. Mixed-grass sites had species-specific shifts in 2007, two breeding seasons after the fires; grassland bird communities on burned plots were similar by 2008 to those on unburned plots. Avian communities appeared to return to pre-burn levels within 3 years following wildfires. Many of the responses in our study of wildfire were similar to those reported following prescribed fires elsewhere. Prescribed fires appear to have similar effects on the avian community despite differences in intensity and environmental conditions during wildfires.

Wilson Journal of Ornithology

Point counts of landbirds in bottomland hardwood forests of the Mississippi Alluvial Valley: How long and how many?

To quantify efficacy of point count sampling in bottomland hardwood forests, we examined the influence of point count duration on corresponding estimates of number of individuals and species recorded. To accomplish this we conducted a totalof 82 point counts 7 May-16 May 1992distributed among three habitats (Wet, Mesic, Dry) in each of three regions within the lower Mississippi Alluvial Valley (MAV). Each point count consisted of recording the number of individual birds (all species) seen or heard during the initial three minutes and per each minute thereafter for a period totaling ten minutes. In addition, we included 384 point counts recorded during an 8-week period in each of 3 years (1985-1987) among 56 randomly-selected forest patches within the bottomlands of western Tennessee. Each point count consisted of recording the number of individuals (excluding migrating species) during each of four, 5 minute intervals for a period totaling 20 minutes. To estimate minimum sample size, we determined sampling variation at each level (region, habitat, and locality) with the 82 point counts from the lower (MAV) and applied the procedures of Neter and Wasserman (1974:493; Applied linear statistical models). Neither the cumulative number of individuals nor number of species per sampling interval attained an asymptote after 10 or 20 minutes of sampling. For western Tennessee bottomlands, total individual and species counts relative to point count duration were similar among years and comparable to the pattern observed throughout the lower MAV. Across the MAV, we recorded a total of 1,62 1 birds distributed among 52 species with the majority (8721/1621) representing 8 species. More birds were recorded within 25-50 m than in either of the other distance categories. There was significant variation in numbers of individuals and species among point counts. For both, significant differences between region and patch (nested within region) occurred; neither habitat nor interaction between habitat and region was significant. For = 0.05 and L3 = 0.10, minimum sample size estimates (per factor level) varied by orders of magnitude depending upon the observed or specified range of desired detectable difference. For observed regional variation, 20 and 40 point counts were required to accommodate variability in total birds (MSE = 9.28) and species (MSE = 3.79), respectively; 25 percent of the mean could be achieved with 5 counts per factor level. Corresponding sample sizes required to detect differences of rarer species (e.g., Wood Thrush) were 500; for common species (e.g., Northern Cardinal) this same level of precision could be achieved with 100 counts.

Book chapter

Marine birds and mammals of the Pacific Subarctic Gyres

The importance of the subarctic gyres of the North Pacific Ocean to marine birds and mammals is poorly known because of a paucity of data spanning appropriate scales of time and space. The little information that is available indicates the western subarctic gyre (WSAG) is more productive than the eastern subarctic gyre (ESAG). In summer the WSAG supports a greater density and higher biomass of seabirds than the ESAG, including at least two species that are more abundant at nesting colonies in the eastern subarctic. Perhaps most revealing of the seabird distributions in this regard is that of southern hemisphere shearwaters (Puffinus spp.) that overwinter in the North Pacific. Their biomass is an order of magnitude greater than that of any northern hemisphere species and is three-fold greater in the WSAG than in the ESAG. Several species of cetaceans also appear to be, or to have been prior to commercial depletions, more abundant in the WSA. Among the many prey species consumed by marine birds and mammals, squids and fishes in the family Myctophidae predominate overall. Other forage species, notably euphausiids, Pacific saury (Cololabis saira) and Atka mackerel (Pleurogrammus monopterygius) are important at times to certain species. The principal exceptions to this generalization are baleen whales and small seabirds that consume zooplankton. Interannual and decadal-scale variability in the physical environment and food web production affect seabirds and marine mammals at sea and at coastal breeding locations around the margins of the gyres.

Progress in Oceanography

Bird community response to field-level integration of prairie strips

Grassland birds are under threat worldwide due to loss of habitat to agriculture. Prairie strips are a new agricultural conservation practice composed of linear strips of reconstructed diverse, native, herbaceous, perennial vegetation designed to promote land sharing among agriculture and biodiversity, while also addressing soil and water conservation goals. We evaluated bird community response to establishment of prairie strips on commercial row-crop fields (corn [( Zea mays ] and soybean [ Glycine max ]) in Iowa, USA compared to controls fields without prairie strips, from 2015 to 2020. We found a 2.94-fold higher density of grassland birds on fields with prairie strips compared to control fields, and a 1.87-fold higher density of birds overall. Time since prairie strip establishment was a significant predictor of grassland bird density, with significant increases between years 1 and 2 and years 3 and 4. Species with the strongest positive response to prairie strips were Red-winged Blackbird ( Agelaius phoeniceus ), Common Yellowthroat ( Geothlypis trichas ), Western Meadowlark ( Sturnella neglecta ) and two species of greatest conservation need: Dickcissel ( Spiza americana ) and Eastern Meadowlark ( Sturnella magna ). Diversity measures (e.g., Shannon’s and Simpson’s indices) did not differ between fields with prairie strips versus those without. Prairie strips provide quality breeding habitat for a suite of species, including grassland species and those of conservation concern. While improving several bird community measures, prairie strips do not provide habitat for area-sensitive grassland birds. Larger grassland patches are needed, potentially managed as land-sparing reserves, to achieve overall biodiversity goals in agricultural landscapes.

Iowa

Geographic variation in migration chronology and winter distribution of midcontinent greater white-fronted geese

We evaluated spatial and temporal differences in migratory behavior among different breeding groups of midcontinent greater white-fronted geese (Anser albifrons) using band-recovery data and observations of neck collared geese during migration and winter. Birds from different breeding areas were initially delineated by geographic distance into 6 banding reference areas (BRAs): 1) interior Alaska, 2) North Slope of Alaska, 3) western Northwest Territories (NWT), 4) western Nunavut, 5) central Nunavut, and 6) eastern Nunavut. The banding groups also differed by breeding habitat, with geese from interior Alaska nesting in the boreal forest (taiga), and all other groups breeding in tundra habitats. Geese from interior Alaska migrated earlier during autumn, and were more likely to winter farther south (in Mexico) than geese from other breeding areas. Geese banded in central and eastern Nunavut (Queen Maud Gulf and Inglis River) wintered farther east (in Louisiana) than geese from other breeding areas. Small-scale (within-state) geographic segregation of wintering flocks was evidenced by the recent (post-1990) nearly exclusive use of a new wintering area in north central Texas by geese from interior Alaska. Segregation among BRAs was also apparent in Mexico, where taiga geese were found predominantly in the central Highlands (states of Zacatecas and Durango), whereas tundra geese mostly used states along the Gulf Coast (primarily Tamaulipas). Interior Alaska birds initiated spring migration earlier than geese from other areas, and were more likely than others to stop in the Rainwater Basin of Nebraska, a region where cholera outbreaks periodically kill thousands of geese. Geese from interior Alaska were the first to arrive at spring staging areas in prairie Canada where BRAs exhibited spatial delineation (a longitudinal cline) in relation to breeding areas. Our results show significant geographic and temporal variation among taiga and tundra breeding cohorts during autumn, winter, and spring. Temporal and spatial differences in migratory behavior may allow management practices that accommodate potential demographic differences between taiga and tundra populations.

Journal of Wildlife Management

Status and nesting of the yellow-billed cuckoo in Puerto Rico

Knowledge of the status of the Yellow-billed Cuckoo ( Coccyzus americanus ) in the Greater Antilles has been clouded by the bird's elusive behavior and spotty distribution on the islands, and by a major movement of North American migrants through the area to and from their wintering grounds in South America. Although Bond (1956) states that they breed in the Greater Antilles, nests are known only from Cuba, St. Croix (Bond 1956), and Jamaica (Lack 1976), and Bond considered the bird rare in Puerto Rico. Other students of Puerto Rican ornithology have suggested that the species is "uncommon" (Leopold 1963), "rare" (McCandless 1958), or a regular migrant or resident species (Wetmore 1916). Although no nests have been reported, most authors suspect that it breeds, and agree that it frequents coastal areas in western or southwestern Puerto Rico, primarily from May to September (Wetmore 1916, Danforth 1936, McCandless 1958, Leopold 1963, Biaggi 1970). From 1968 to 1971 we conducted linear strip censuses along pre-selected routes throughout the island, counting all birds heard or seen (Kepler and Kepler 1970); data we obtained on the Yellow-billed Cuckoo helped clarify and amplify existing information on its distribution and status.

The Auk

Birds of the St. Croix River valley: Minnesota and Wisconsin

The St. Croix River Valley encompasses nearly 11,550 km2 in east-central Minnesota and northwestern Wisconsin. A wide range of habitats are available for birds including upland oak, lowland deciduous, maple-basswood, lowland and upland coniferous forests, natural basin wetlands, and grasslands. Situated in the north-central region of the United States, the valley is a biological 'crossroads' for many species. Because of the mixed affinities of plant communities, the valley includes the northern and southern range limits for a number of species. Also, because the valley lies near the forest-prairie transition zone, many typical western breeding species (e.g. pintail, western meadowlark, yellow-headed blackbird) breed in proximity to typical eastern species such as tufted titmouse, eastern meadowlark, and cardinal. From 1966 to 1980, I conducted extensive surveys of avian distribution and abundance in the St. Croix River Valley. I have supplemented the results of these surveys with published and unpublished observations contributed by many ornithologists. These additional data include compilations from Christmas Bird Counts sponsored by the National Audubon Society and from the Breeding Bird Survey coordinated by the U.S. Fish and Wildlife Service. Three hundred fourteen species have been recorded in the study area; data are presented on the migration period, nesting season distribution, winter distribution, relative abundance, and habitat use of each species. Recognizing the uniqueness of the area, and its importance not only to wildlife but also to man, the U.S. Congress designated the St. Croix a National Scenic Riverway. This action provided a considerable degree of protection to lands along and directly adjacent to the river. Unfortunately, no similar legal measure exists to protect lands away from the river. With the exception of the northern quarter of the St. Croix River Valley, agricultural interests have made significant inroads into the habitat base. The continuing expansion of the nearby Minneapolis-St. Paul metropolitan region has degraded or destroyed many woodlots, upland fields, and wetlands. In numerous instances, degradation of natural habitats has influenced the abundance and distribution of bird species. Because of these changes, both the Federal government and State Departments of Natural Resources have listed several species in various categories based on their current status. In the St. Croix River Valley, seven species are endangered, eight are threatened, and 29 are watch or priority status in either or both states. Data presented in this report are of value to land managers, land use specialists, and ornithologists, in assessing current and projected habitat alterations on the avifauna of this valley. The St. Croix River bisects a large region of western Wisconsin and east central Minnesota that exhibits a wide range of habitat types. This region supports not only birds, but many mammals, fishes, reptiles and amphibians, and several thousand species of vascular and nonvascular plants. The river itself is relatively clean through most of its course, and its natural flow is interrupted by only two small dams. Because the river lies within a 1-day drive of nearly 10 million people (Waters 1977), use of the area for recreational purposes is extremely heavy. Recreational pursuits include sunbathing, boating, and wild river kayaking in the summer, and ice fishing and cross-country skiing in the winter. The large number of unique and highly fragile habitats that exist there may never be compatible with the uses and abuses of the land that go with expanding human populations. Through the efforts of a number of citizens concerned with the quality of their environment and the foresightedness of several local, State, and Federal legislators, a portion of the upper St. Croix River Valley (hereafter termed 'the Valley') was established as a National Wild and Scenic River. Through establishment of t

North American Fauna

Mercury concentrations in breast feathers of three upper trophic level marine predators from the western Aleutian Islands, Alaska

Mercury (Hg) is a toxic element distributed globally through atmospheric transport. Agattu Island, located in the western Aleutian Islands, Alaska, has no history of point-sources of Hg contamination. We provide baseline levels of total mercury (THg) concentrations in breast feathers of three birds that breed on the island. Geometric mean THg concentrations in feathers of fork-tailed storm-petrels ( Oceanodroma furcata ; 6703 ± 1635, ng/g fresh weight [fw]) were higher than all other species, including snowy owl ( Bubo scandiacus ; 2105 ± 1631, ng/g fw), a raptor with a diet composed largely of storm-petrels at Agattu Island. There were no significant differences in mean THg concentrations of breast feathers among adult Kittlitz’s murrelet ( Brachyramphus brevirostris ; 1658 ± 1276, ng/g fw) and chicks (1475 ± 671, ng/g fw) and snowy owls. The observed THg concentrations in fork-tailed storm-petrel feathers emphasizes the need for further study of Hg pollution in the western Aleutian Islands.

Alaska

Fitness consequences of catastrophic wildfire are mitigated by behavioral responses of an iconic bird

Background Drought, human disturbance, and invasive species are reshaping disturbance regimes and increasing the scale, severity, and frequency of wildfire in many ecosystems around the globe, including the sagebrush steppe of western North America. Recent studies suggested greater sage-grouse ( Centrocercus urophasianus ) adhere to strong site fidelity in the aftermath of wildfire, remaining inside fire perimeters for nesting and brood rearing despite negative consequences for survival and reproduction. Sage-grouse in Idaho exhibited context-dependent changes to space use after a large, high-severity fire that burned > 40,000 ha, yet the specific behavioral responses to fire and their fitness consequences remain unclear. We used data collected from 269 hens over a 6-year period under a multi-level before-after-control-impact design to test the hypothesis that sage-grouse mitigated fitness consequences of high-severity wildfire through adaptive behavioral responses and spatial redistribution. Results We tested predictions deduced from our hypothesis at the population and individual levels using behavioral, demographic, and life history data, including nesting metrics, brood-rearing metrics, hen survival, and body mass at capture. Fifteen of 16 predictions were supported, demonstrating that post-fire space use and avoidance of the burn was adaptive and helped mitigate fitness effects of the fire. Short-term consequences included elimination of nesting and brood rearing habitat and subsequent shifts to the distribution of usable space. Yet fitness consequences were minimal because of behavioral flexibility employed by hens during nesting and brood rearing. Conclusions Behavioral responses to wildfire by sage-grouse are more flexible than has been described, and sage-grouse demonstrated resilience by rapidly adapting space use to avoid short-term consequences of catastrophic fire when high-quality habitat remained adjacent to the burn and within their seasonal range. Our results imply behavioral and fitness consequences of fire are context-dependent and likely impacted by attributes of the fire and surrounding landscape after disturbance. Furthermore, among-study differences in behavioral and fitness outcomes of sage-grouse after fire supported underappreciated predictions from both fire ecology and site fidelity theory, and suggest conditions where behavioral flexibility should be expressed, and fidelity relaxed, based on severity of disturbance, landscape context, and species mobility.

Idaho

Distribution of two western clapper rail races as determined by responses to taped calls

During 1969 and 1970, surveys of the endangered Yuma Clapper Rail were conducted using taped calls to elicit responses from the birds. During the two summers, more than 158 Yuma clappers were located in cattailtule marshes along the Colorado River south of Needles, California, to the International Boundary, a distance of about 240 miles. Clappers (probably of the same race) were also found in estuarian marshes of the Colorado River Delta of Mexico; in the Salton Sea; in two freshwater marsh areas near Phoenix, Arizona; and in two freshwater marshes adjacent to the lower Gila River near Tacna, Arizona.....Populations of Sonora Clapper Rails were discovered as permanent residents in five separate mangrove swamps along the west coast of Mexico in the vicinity of Kino Bay, Sonora. These observations were farther north than any heretofore reported for the race R. l. rhizophorae, and the swamps also represent the extreme northward limit of mangroves in Sonora.....During the winter, Yuma clappers did not respond to taped calls north of the International Boundary, whereas clappers along the coast of Sonora readily answered the calls during the same period of time. We conclude that most Yuma Clapper Rails migrate from their summer habitat along the Colorado River in September and do not return to the breeding areas until late April.

Condor

Age-specific survival rates, causes of death, and allowable take of golden eagles in the western United States

In the United States, the Bald and Golden Eagle Protection Act prohibits take of golden eagles ( Aquila chrysaetos ) unless authorized by permit, and stipulates that all permitted take must be sustainable. Golden eagles are unintentionally killed in conjunction with many lawful activities (e.g., electrocution on power poles, collision with wind turbines). Managers who issue permits for incidental take of golden eagles must determine allowable take levels and manage permitted take accordingly. To aid managers in making these decisions in the western United States, we used an integrated population model to obtain estimates of golden eagle vital rates and population size, and then used those estimates in a prescribed take level (PTL) model to estimate the allowable take level. Estimated mean annual survival rates for golden eagles ranged from 0.70 (95% credible interval = 0.66–0.74) for first-year birds to 0.90 (0.88–0.91) for adults. Models suggested a high proportion of adult female golden eagles attempted to breed and breeding pairs fledged a mean of 0.53 (0.39–0.72) young annually. Population size in the coterminous western United States has averaged ~31,800 individuals for several decades, with λ = 1.0 (0.96–1.05). The PTL model estimated a median allowable take limit of ~2227 (708–4182) individuals annually given a management objective of maintaining a stable population. We estimate that take averaged 2572 out of 4373 (59%) deaths annually, based on a representative sample of transmitter-tagged golden eagles. For the subset of golden eagles that were recovered and a cause of death determined, anthropogenic mortality accounted for an average of 74% of deaths after their first year; leading forms of take over all age classes were shooting (~670 per year), collisions (~611), electrocutions (~506), and poisoning (~427). Although observed take overlapped the credible interval of our allowable take estimate and the population overall has been stable, our findings indicate that additional take, unless mitigated for, may not be sustainable. Our analysis demonstrates the utility of the joint application of integrated population and prescribed take level models to management of incidental take of a protected species.

western United States

The role of landscape and habitat characteristics in limiting abundance of grassland nesting songbirds in an urban open space

We examine the relationships between abundance of grassland nesting songbirds observed in the Boulder Open Space, CO, USA and parameters that described landscape and habitat characteristics, in order to provide information for Boulder Open Space planners and managers. Data sets included bird abundance and plant species composition, collected during three breeding seasons (1994–1996), and landscape composition and configuration measures from a satellite image-derived land-cover map. We used regression quantiles to estimate the limitations imposed on bird abundance by urban encroachment and decreasing areas of grassland cover-types on the landscape, and habitat characteristics within 200 m diameter sample plots. After accounting for the effect of landscape grassland composition on four species (Western Meadowlark ( Sturnella neglecta ), Vesper Sparrow ( Pooecetes gramineus ), Horned Lark ( Eremophila alpestris ), and Grasshopper Sparrow ( Ammodramus savannarum )), change in abundance with proportion of urban area in the landscape was consistent with the pattern expected for limiting factors that were the active constraint at some times and places. Area of preferred grassland cover-types on the landscape was important for all species, and this remained the case when habitat variables were included in combined landscape–habitat models, with one exception (Western Meadowlark). Analysis of habitat variables enabled identification of important features at the local scale (e.g. shale plant communities in Lark Sparrow ( Chondestes grammacus ) habitat) that were indistinguishable using landscape data alone. Consideration of changes in the landscape due to urbanization and loss of grassland habitat are crucial for open space planning, and habitat features associated with localized and clumped bird species distributions provide important additional information. Widening the management focus to include areas that are not part of the open space system will facilitate a more complete understanding of potential limiting factor processes.

Landscape and Urban Planning

Amphibian, reptilian, and avian remains from the Fox Hills Formation (Maastrichtian): Shoreline and estuarine deposits of the Pierre Sea in south-central North Dakota

Although vertebrate fossils, except for fish, are not common in the Maastrichtian Fox Hills Formation, amphibian, reptilian, and avian remains have been recovered at several localities in south-central North Dakota from shoreline facies of the retreating Pierre-Fox Hills seaway. This mixed fauna of aquatic, terrestrial, and marine taxa provides insight into the composition of coastal communities and habitats at the interface between the Hell Creek delta and the Western Interior Seaway. The delta-platform aquatic paleocommunity is represented by the efficient swimming salamanders Opistho- trition kayi and Lisserpeton bairdi, the carnivorous soft-shelled turtle "Aspideretes" sensu lato, the underwater piscivorous predator Champsosaurus laramiensis, and the large, predatory crocodile IBorealosuchus. Terrestrial areas were inhabited by the tortoise-like Basilemys and the predatory dinosaurs Tyrannosaurus and cf. Saurornit- holestes. Birds occupied niches in the warm-temperate to subtropical, forested delta platform and shoreline areas. These nonmarine taxa in the Fox Hills Formation indicate that the geographic range of these animals extended to shoreline areas of the Western Interior Seaway. The toxochelyid turtle Lophochelys and the ambush predators Mosasaurus dekayi and IPlioplatecarpus resided in the shallow marine and estuarine habitats. These taxa and marine fish taxa reported earlier indicate that normal marine conditions in south- central North Dakota persisted into the latest Late Cretaceous in comparison with coeval Hell Creek Formation sites more distal from the Western Interior Seaway. ?? 2007 The Geological Society of America. All rights reserved.

Special Paper of the Geological Society of America