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USGS2025

One hundred ninety-nine dead birds: Review of the scientific basis of ecological incident reporting requirements for pesticide registrants under Fifra § 6(A)(2)

The U.S. Environmental Protection Agency (EPA) regulates pesticide use in the United States. The EPA is charged by the Federal Insecticide, Fungicide, and Rodenticide Act (FIFRA) with ensuring that a pesticide will not cause unreasonable adverse effects on the environment. Incident reports (documentation of exposure and injury from pesticide applications) can serve as a reality check on the pesticide registration decisions made by the EPA scientists and risk managers. The EPA collects incident reports on human, domestic animal, and ecological injury. The FIFRA section 6(a)(2) rule requires the pesticide registrant (generally, the company or other entity that wishes to market the pesticide, hereafter registrant) to submit such data to the EPA. The EPA’s ecological incident category includes injuries to aquatic (fish), terrestrial (wildlife), other non-target organisms (ONT, e.g., invertebrates) and plants. Our document focuses on the fish and wildlife ecological incidents that are submitted by registrants. We critique the application of the FIFRA section 6(a)(2) rule that controls the quality and quantity of ecological incident data that the EPA receives from registrants. We conclude that the section 6(a)(2) provisions can impede the transfer of ecological incident data from registrant to the EPA. Consequently, detailed data for many fish and wildlife incidents may never reach the EPA, and policies and decisions may be formulated in the absence of these data.

Buffalo Environmental Law Journal↗

USGS2009

Use of land surface remotely sensed satellite and airborne data for environmental exposure assessment in cancer research

In recent years, geographic information systems (GIS) have increasingly been used for reconstructing individual-level exposures to environmental contaminants in epidemiological research. Remotely sensed data can be useful in creating space-time models of environmental measures. The primary advantage of using remotely sensed data is that it allows for study at the local scale (e.g., residential level) without requiring expensive, time-consuming monitoring campaigns. The purpose of our study was to identify how land surface remotely sensed data are currently being used to study the relationship between cancer and environmental contaminants, focusing primarily on agricultural chemical exposure assessment applications. We present the results of a comprehensive literature review of epidemiological research where remotely sensed imagery or land cover maps derived from remotely sensed imagery were applied. We also discuss the strengths and limitations of the most commonly used imagery data (aerial photographs and Landsat satellite imagery) and land cover maps.

Journal of Exposure Science and Environmental Epid↗

USGS1999

May the forethought (and studies) be with your campsite-protection planning!

Visitation has reached record levels along the Appalachian Trail, a 2000+ mile footpath extending from Maine to Georgia along the crest of the Appalachian Mountains. Camping impacts associated with this use have also expanded rapidly in recent years, particularly in popular National Parks and at attraction features such as lakes and ponds. This article reviews recreation ecology research on camping impacts and their relationship to amount of use and environmental attributes. Management options for responding to camping management problems are described, including the manipulation of use-related, environmental, and managerial factors.

USGS2013

Agricultural sources of contaminants of emerging concern and adverse health effects on freshwater fish

Agricultural contaminants of emerging concern (CECs) are generally thought of as certain classes of chemicals associated with animal feeding and production facilities. Veterinary pharmaceuticals used in animal food production systems represent one of the largest groups of CECs. In our review, we discuss the extensive increase in use of antibiotics in animal feeding operations (AFOs) around the world. AFOs are a major consumer of antibiotics and other veterinary pharmaceuticals and over the past decade there has been growing information on the occurrence, release, and fate of CECs from animal food production operations, including the application of pharmaceutical-containing manure to agricultural fields and releases from waste lagoons. Concentrations of CECs in surface and ground water in proximity to AFOs correspond to their presence in the AFO wastes. In many cases, the environmental concentrations of agriculturally-derived CECs are below toxicity thresholds. Hormones and hormone replacement compounds are a notable exception, where chemical concentrations near AFOs can exceed concentrations known to cause adverse effects on endocrine-related functions in fish. In addition, some agricultural pesticides, once thought to be safe to non-target organisms, have demonstrated endocrine-related effects that may pose threats to fish populations in agricultural regions. That is, we have pesticides with emerging concerns, thus, the concern is emerging and not necessarily the chemical. In this light, one must consider certain agricultural pesticides to be included in the list of CECs. Even though agricultural pesticides are routinely evaluated in regulatory testing schemes which have been used for decades, the potential hazards of some pesticides have only recently been emerging. Emerging concerns of pesticides in fish include interference with hormone signaling pathways; additive (or more than additive) effects from pesticide mixtures; and adverse population-level effects at concentrations below predicted toxicity thresholds. Consequently, there is a need to evaluate the environmental concerns related to pesticide exposures to fish populations based on current biological and toxicological techniques. This presentation reviews some of the agricultural chemicals that have emerged as contaminants of concern and potentially threaten fish populations in agricultural watersheds.

Conference Paper↗

USGS2024

Hydrologic investigations and a preliminary conceptual model of the groundwater system at North Penn Area 1 Superfund Site, Souderton, Montgomery County, Pennsylvania

The U.S. Geological Survey (USGS) conducted hydrogeologic investigations, reviewed existing data, and developed a preliminary conceptual model of the groundwater system as part of technical support of the U.S. Environmental Protection Agency (EPA) at the North Penn Area 1 Superfund Site (hereafter, the NP1 Site) located within the Borough of Souderton in Montgomery County, Pennsylvania. Field work and monitoring took place during 2012–18. The area is underlain by sedimentary formations that form a fractured-rock aquifer used for drinking water and industrial supply. The EPA placed the Site on the National Priorities List in 1989, identifying tetrachloroethylene (PCE) and trichloroethylene (TCE) as contaminants of concern. During 2012–18, the USGS conducted field activities that included drilling an 82-foot (ft)-deep monitoring well (MG 2220) in 2016, reconstructing a 208-ft-deep former industrial production well (MG 668 [Granite Knitting Mill]), and collecting borehole geophysical and video logs and water levels from those and five additional wells, which ranged in depth from about 50 to 200 ft below land surface. Continuous water levels were collected during 2014–17, and a synoptic set of water levels were measured in April 2018 in the seven wells. The borehole geophysical logs (caliper, acoustic televiewer, natural gamma, single-point resistance, vertical flow, and fluid temperature and resistivity) and borehole video logs in the seven wells were evaluated to assess potential for lithologic correlation and to identify and describe water-bearing features, which included both low- and high-angle fractures and other openings oriented along dipping bedding planes, joints, or possible faults. Borehole geophysical logs collected by USGS in 1992 in a 300-ft-deep former production well near the Site were also evaluated. Few to no distinctive features were identified on geophysical logs (natural gamma and single-point resistance) that could be used for correlation, thus limiting this approach to determining local geologic structure. Extensive fracturing in the upper 62 ft of monitoring well MG 2220 indicates that the well was likely drilled through a zone of faulting, and other evidence of faulting is present in the area near the Site. Assessment of continuous water levels showed hydraulic connections among some wells as indicated by rising or falling water levels in response to changes in pumping rates at nearby wells. A map of water levels measured in April 2018 indicates potential for groundwater flow generally toward the stream to the south and southwest of the Site, but the limited water-level data are insufficient to describe vertical groundwater gradients or lateral gradients in any detail. Review of 1999–2022 volatile organic compound (VOC) monitoring data collected by the Pennsylvania Department of Environmental Protection for five monitoring wells indicates that the highest groundwater concentrations of PCE and TCE were found in samples from extraction well MG 2201 (S-1) downgradient from, and nearest to, the previously identified Site contaminant source area, and these concentrations fluctuated through time. PCE concentrations were higher than TCE concentrations in samples from all five monitoring wells and were much higher than TCE concentrations in samples from extraction well MG 2201 (S-1). Temporally variable recharge is a possible factor affecting observed fluctuations in PCE concentrations in groundwater samples from well extraction MG 2201 (S-1), as indicated by a general inverse relation between PCE concentrations and water levels in a nearby long-term observation well. The PCE concentration of 1,830 micrograms per liter (μg/L) in a May 2018 water sample from monitoring well MG 2220 was more than four times the PCE concentration of 444 μg/L in a December 2017 sample from the nearby extraction well MG 2201 (S-1), which is open to fewer fractures. Low concentrations of VOCs were measured in surface water at two stream sites downgradient from wells with the highest groundwater VOC concentrations at the Site, indicating that discharge of contaminated groundwater to the stream is likely. Development of a conceptual model of the groundwater system was constrained by limited data. In areas with no pumping, groundwater-flow directions generally are thought to be controlled by topography and geologic structure (bedding orientation) and likely to the south and southwest of the Site, with local flow directions affected by orientations of fractures, joints, and local faults. Additional investigations that could help improve the conceptual model of the groundwater system and help delineate the extent of groundwater contamination and its transport are discussed.

Pennsylvania↗

USGS1980

A summary report of the regional geology, environmental geology, OCS resource appraisal, petroleum potential, and operational considerations in the area of proposed lease sale 73, offshore California

This report reviews geological, geophysical and technological data pertinent to proposed Lease Sale 73. Under consideration is the entire continental margin of California that is bounded on the west by the 2500 meter isobath. This area encompasses the whole of the southern California Borderland, the continental shelves of central and northern California, and the upper part of the continental slope from Mexico on the south to Oregon on the north. The area includes both leased and unleased tracts and lies adjacent to highly productive coastal basins of central and northern California . Because the geology of the south and north parts of this large area differ, they are considered separately in this report.

California↗

USGS2014

The geochemistry of deep-sea coral skeletons: a review of vital effects and applications for palaeoceanography

Deep-sea corals were discovered over a century ago, but it is only over recent years that focused efforts have been made to explore the history of the oceans using the geochemistry of their skeletal remains. They offer a promising archive of past oceanic environments given their global distribution, layered growth patterns, longevity and preservation as well as our ability to date them using radiometric techniques. This paper provides an overview of the current state-of-the-art in terms of geochemical approaches to using deep-sea coral skeletons to explore the history of the ocean. Deep-sea coral skeletons have a wide array of morphologies (e.g. solitary cup corals, branching colonial corals) and materials (calcite, aragonite and proteins). As such their biomineralization strategies are diverse, leading to complex geochemistry within coral skeletons. Notwithstanding these complications, progress has been made on developing methods for reconstructing the oceanographic environment in the past using trace elements and isotopic methods. Promising approaches within certain coral groups include clumped isotopes and Mg/Li for temperature reconstructions, boron isotopes and radiocarbon for carbon cycling, εNd, and radiocarbon for circulation studies and δ 15 N, P/Ca and Ba/Ca for nutrient tracer studies. Likewise there is now a range of techniques for dating deep-sea corals skeletons (e.g. U-series, radiocarbon), and determining their growth rates (e.g. radiocarbon and 210 Pb). Dating studies on historic coral populations in the Atlantic, Southern Ocean and Pacific point to climate and environmental changes being dominant controls on coral populations over millennial and orbital timescales. This paper provides a review of a range of successes and promising approaches. It also highlights areas in which further research would likely provide new insights into biomineralization, palaeoceanography and distribution of past coral populations.

Deep-Sea Research Part II: Topical Studies in Ocea↗

USGS2025

A critical perspective on the Society of Environmental Toxicology and Chemistry’s adherence to founding principles—Opportunities for the future

The Society of Environmental Toxicology and Chemistry (SETAC) is a global organization whose mission is the advancement of environmental science and the promotion of science-informed decision making. On SETAC’s 45th anniversary, the following question was raised: Are the 1979 founding principles of SETAC, multidisciplinary approaches to solving environmental problems, multisector engagement and scientific objectivity, still relevant to the fulfillment of its mission? In a special session held at the 45th Annual Meeting in Fort Worth, Texas, U.S., a critical evaluation of the founding principles was initiated by reviewing SETAC’s history and ongoing activities, and recommendations were made for the future. With few exceptions, participants appreciated SETAC’s purposeful efforts to approach challenging environmental issues through multisector balance, an approach that is unique amongst scientific societies. We recognized that scientists have biases and views of what they find important, regardless of employing organization, and that objectivity is best served by being aware of these biases and views. SETAC’s founding principles have stood the test of time and continue to provide a strong foundation for the Society’s mission, and with a few suggested improvements, will continue to be instrumental in guiding environmental science, stewardship and policy into the future. The significance of SETAC’s contribution of robust science grounded in reliable evidence and data was recognized as being especially crucial at this time of triple planetary crisis (climate change, pollution and biodiversity loss), compounded by rapid technological developments and geopolitical issues.

Environmental Toxicology and Chemistry↗

USGS2022

Use of riparian spiders as sentinels of persistent and bioavailable chemical contaminants in aquatic ecosystems: A review

Aquatic ecosystems around the world are contaminated with a wide range of anthropogenic chemicals, including metals and organic pollutants, that originate from point and nonpoint sources. Many of these chemical contaminants have complex environmental cycles, are persistent and bioavailable, can be incorporated into aquatic food webs, and pose a threat to the health of wildlife and humans. Identifying appropriate sentinels that reflect bioavailability is critical to assessing and managing aquatic ecosystems impacted by contaminants. The objective of the present study is to review research on riparian spiders as sentinels of persistent and bioavailable chemical contaminants in aquatic ecosystems. Our review of the literature on riparian spiders as sentinels suggests that significant progress has been made during the last two decades of research. We identified 55 published studies conducted around the world in which riparian spiders (primarily of the families Tetragnathidae, Araneidae, Lycosidae, and Pisauridae) were used as sentinels of chemical contamination of lotic, lentic, and estuarine systems. For several contaminants, such as polychlorinated biphenyls (PCBs), Hg, and Se, it is now clear that riparian spiders are appropriate sentinels. However, many contaminants and factors that could impact chemical concentrations in riparian spiders have not been well characterized. Further study of riparian spiders and their potential role as sentinels is critical because it would allow for development of national-scale programs that utilize riparian spiders as sentinels to monitor chemical contaminants in aquatic ecosystems. A riparian spider sentinel program in the United States would be complementary to existing national sentinel programs, including those for fish and immature dragonflies.

Environmental Toxicology and Chemistry↗

USGS1995

Oil, biological communities and contingency planning

The Oil Pollution Act of 1990 mandates the inclusion of a fish and wildlife response plan in the National Contingency Plan (NCP) and the creation of Area Committees that must develop an Area Contingency Plan (ACP). Area Contingency Plans must include a detailed annex containing a Fish and Wildlife and Sensitive Environments Plan. Tank vessels, offshore facilities, and certain onshore facilities must have response plans consistent with the requirements of the NCP and the ACP. New regulations to supersede the Type A and B procedures of the Natural Resources Damage Assessment Regulations are being developed for oil spills. Currently, four assessment methods have been proposed: (1) Type A, (2) comprehensive (Type B), (3) intermediate (between types A and B), and (4) compensation tables. The Oil Spill Liability Trust Fund is approaching its ceiling of $1 billion, but only $50 million has been appropriated. Effective biological contingency planning requires extensive knowledge of (1) the environmental fate of petroleum, (2) the effects of petroleum on organisms, (3) the existing biological resources, and (4) the establishment of a system of biological priorities. The characteristics and fate of petroleum and the biological effects of petroleum are reviewed. Assessment of biological resources includes plant and animal distributions, important habitat, endangered or threatened species, and economic considerations. The establishment by Area Committees of priorities for environmental protection, injury assessment, and restoration will promote efficient spill response. Three special issues are discussed: (1) improving our ability to restore natural resources, (2) the potential role of biological diversity in spill response planning, and (3) planning for animal rehabilitation.

Book chapter↗

USGS2020

Bellwether sites for evaluating changes in landslide frequency and magnitude in cryospheric mountainous terrain: A call for systematic, long-term observations to decipher the impact of climate change

Permafrost and glaciers are being degraded by the warming effects of climate change. The impact that this degradation has on slope stability in mountainous terrain is the subject of ongoing research efforts. The relatively new availability of high-resolution (≤ 10 m) imagery with worldwide coverage and short (≤ 30 days) repeat acquisition times, as well as the emerging field of environmental seismology, presents opportunities for making remote, systematic observations of landslides in cryospheric mountainous terrain. I reviewed the literature and evaluated landslide activity in existing imagery to select five ~ 5000-km 2 sites where long-term, systematic observations could take place. The five proposed sites are the northern and eastern flanks of the Northern Patagonia Ice Field, the Western European Alps, the eastern Karakoram Range in the Himalayan Mountains, the Southern Alps of New Zealand, and the Fairweather Range in Southeast Alaska. Systematic observations of landslide occurrence, triggers, size, and travel distance at these sites, especially if coupled with observations from in situ instrumental monitoring, could lead to a better understanding of changes in slope stability induced by climate change. The suggested sites are not meant to be absolute and unalterable. Rather, they are intended as a starting point and discussion starter for new work in this expanding landslide research frontier.

Landslides↗

USGS1991

The interaction between biology and the management of aquatic macrophytes

‘Management’ refers to controlling nuisance aquatic species and to restoring or restructing aquatic plant communities. Producing stable, diverse, aquatic plant communities containing a high percentage of desirable species is a primary management goal. There are a variety of techniques including harvesting, herbicides, water-level fluctuation, sediment alteration, nutrient limitation, light alteration, and biological controls which can be used for managing macrophytes. These techniques are briefly reviewed along with discussions of biological considerations important to the efficacy of the technique and the environmental impacts of the technique. There is a growing interest in restoring and restructing aquatic plant communities. Techniques for community restoration are discussed as are emerging management technologies using growth regulators and bioengineering. New management technologies will probably be limited by costs and environmental impacts. In the near future, better macrophyte management will come through better planning and more effective use of present technology. The challenge is to make current planning and management techniques more effective through increased biological inputs. The potential for biological input ranges from subcellular biology to species biology, to community and ecosystem biology. Some information needs are identified.

Aquatic Botany↗

USGS2025

Late Quaternary environmental change in eastern Beringia

Eastern Beringia (Alaska and western Yukon) is an extensive, high-latitude region of North America that remained largely unglaciated throughout the Quaternary. Consequently, its sedimentary deposits preserve long-term environmental records that have intrigued scientists for nearly a century. Recent advances in palaeoecological proxies and dating methods have proved critical in addressing long-standing questions about regional late Quaternary environmental change. At the same time, they have led to new and sometimes controversial hypotheses. This review covers recent discoveries and unresolved questions focused on the period 57,000–10,000 calendar years before C.E. 1950 (cal yr BP). The middle Wisconsin interstadial (57,000–30,000 cal yr BP) was a period of relative warmth in eastern Beringia, compared with the late Wisconsin (30,000–14,000 cal yr BP). Early in the interstadial occasional Picea woodland was present amongst widespread shrub tundra. Palaeoecological, sedimentary and isotopic data indicate that climate was cooler and drier than the Holocene, with high rates of aeolian activity. Megafauna typically associated with the ‘mammoth steppe’ ecosystem (woolly mammoth [ Mammuthus primigenius ], horse [ Equus ] and steppe-bison [ Bison priscus ]) were present in some abundance. The transition towards late Wisconsin cold-stage conditions (35,000–30,000 cal yr BP) coincided with the establishment of the Bering Land Bridge and featured expansion of spatially varied, herbaceous vegetation, sometimes associated with deep active layers. Sedimentary DNA ( seda DNA) and macrofossil evidence show vegetation was not a prairie-like grassland, and the term “steppe-tundra” is a better descriptor. Permafrost pore-ice isotopic (δ 18 O) records suggest a step change in one or more climate drivers ca. 30,000 cal yr BP, by which time steppe-tundra was established across eastern Beringia. It remains uncertain whether Picea survived cold-stage conditions within isolated refugia, or whether it recolonized from south of the Laurentide-Cordilleran ice sheets. Genetic data suggest that Picea probably survived in situ ; however, there is no definitive fossil evidence to support this. The end-Pleistocene transition from steppe-tundra to shrub tundra began ca. 15,000 cal yr BP and took place within decades at local scales. The expansion of woody taxa coincided with rising sea levels, reduced sea-ice extent and an abrupt shift in atmospheric circulation that enhanced precipitation. During this time, Earth's orbital configuration caused high early-summer temperatures and strong seasonality, creating growing conditions very different from today. The vegetation consisted of Salix and Betula shrub tundra with open areas of herbs and graminoids. During the deglacial warming trend, the Younger Dryas oscillation (12,800–11,700 cal yr BP) was variably expressed. It is generally evident in records affected by adjacent oceans but can be absent at sites in continental areas. These past conditions and paleoenvironmental changes have implications for contemporary issues: hypotheses about Pleistocene mammalian extinction; sensitivity of eastern Beringia to major oceanic reorganizations and high-frequency climate variability; the nature of woody plant expansion with climate warming; grazing, hydroclimate and fire as controls over ecosystems; the efficacy of “Pleistocene rewilding” for carbon capture.

Beringia↗

USGS2020

Storm impacts on phytoplankton community dynamics in lakes

In many regions across the globe, extreme weather events such as storms have increased in frequency, intensity, and duration due to climate change. Ecological theory predicts that such extreme events should have large impacts on ecosystem structure and function. High winds and precipitation associated with storms can affect lakes via short‐term runoff events from watersheds and physical mixing of the water column. In addition, lakes connected to rivers and streams will also experience flushing due to high flow rates. Although we have a well‐developed understanding of how wind and precipitation events can alter lake physical processes and some aspects of biogeochemical cycling, our mechanistic understanding of the emergent responses of phytoplankton communities is poor. Here we provide a comprehensive synthesis that identifies how storms interact with lake and watershed attributes and their antecedent conditions to generate changes in lake physical and chemical environments. Such changes can restructure phytoplankton communities and their dynamics, as well as result in altered ecological function (e.g., carbon, nutrient and energy cycling) in the short‐ and long‐term. We summarize the current understanding of storm‐induced phytoplankton dynamics, identify knowledge gaps with a systematic review of the literature, and suggest future research directions across a gradient of lake types and environmental conditions.

Global Change Biology↗

USGS2010

New trends in species distribution modelling

Species distribution modelling has its origin in the late 1970s when computing capacity was limited. Early work in the field concentrated mostly on the development of methods to model effectively the shape of a species' response to environmental gradients (Austin 1987, Austin et al. 1990). The methodology and its framework were summarized in reviews 10–15 yr ago (Franklin 1995, Guisan and Zimmermann 2000), and these syntheses are still widely used as reference landmarks in the current distribution modelling literature. However, enormous advancements have occurred over the last decade, with hundreds – if not thousands – of publications on species distribution model (SDM) methodologies and their application to a broad set of conservation, ecological and evolutionary questions. With this special issue, originating from the third of a set of specialized SDM workshops (2008 Riederalp) entitled 'The Utility of Species Distribution Models as Tools for Conservation Ecology', we reflect on current trends and the progress achieved over the last decade.

Ecography↗

USGS2020

Considerations for incorporating quality control into water quality sampling strategies for the U.S. Geological Survey

This report describes considerations for incorporating routine quality-assessment and quality-control evaluations into U.S. Geological Survey discrete water-sampling programs and projects. U.S. Geological Survey water-data science in 2020 is characterized by robustness, external reproducibility, collaborative large-volume data analysis, and efficient delivery of water-quality data. Confidence in data, or robustness, can be increased by supplementing traditional field-based quality-control data with laboratory quality control (QC) data, such as third-party blind spikes and blind blanks, laboratory blanks, and laboratory-reagent spikes. Laboratory quality-control data can provide additional information about bias and variability, method performance, and false-positive and false-negative rates that are not available from field QC data alone. Reproducibility is supported by means of standardizing metadata and documentation. Collaborative analysis brings together disparate elements of various types of quality-control review and communicates persistent data quality issues for compounds to data users internal and external to the U.S. Geological Survey. Efficient delivery of water-quality data is achieved when quality-control review is accomplished in the same expedited (near real-time) time frame as distribution of environmental results to the public and might be improved with consideration given to data versioning or to a system of alerting data users to data interpretation that might differ from originally published data.

Open-File Report↗

USGS2013

Quagga and zebra mussels: biology, impacts, and control

Quagga and Zebra Mussels: Biology, Impacts, and Control , Second Edition provides a broad view of the zebra/quagga mussel issue, offering a historic perspective and up-to-date information on mussel research. Comprising 48 chapters, this second edition includes reviews of mussel morphology, physiology, and behavior. It details mussel distribution and spread in Europe and across North America, and examines policy and regulatory responses, management strategies, and mitigation efforts. In addition, this book provides extensive coverage of the impact of invasive mussel species on freshwater ecosystems, including effects on water clarity, phytoplankton, water quality, food web changes, and consequences to other aquatic fauna. It also reviews and offers new insights on how zebra and quagga mussels respond and adapt to varying environmental conditions. This new edition includes seven video clips that complement chapter text and, through visual documentation, provide a greater understanding of mussel behavior and distribution.

Book↗

USGS2005

Literature Review and Annotated Bibliography: Water Requirements of Desert Ungulates

Executive Summary Ungulates adapted to desert areas are able to survive extreme temperatures and limited water availability. This ability is largely due to behavioral, morphological, and physiological adaptations that allow these animals to avoid or tolerate extreme environmental conditions. The physiological adaptations possessed by ungulates for thermoregulation and maintenance of water balance have been the subject of numerous studies involving a wide range of species. In this report we review the behavioral, morphological, and physiological mechanisms used by ungulates and other desert mammals to maintain water and temperature balance in arid environments. We also review some of the more commonly used methods for studying the physiological mechanisms involved in water balance and thermoregulation, and the influence of dehydration on these mechanisms.

Open-File Report↗