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At least 217 records · Page 12Linked to original sources

Crustal stress and structure at Kīlauea Volcano inferred from seismic anisotropy

Seismic anisotropy, measured through shear wave splitting (SWS) analysis, can be indicative of the state of stress in Earth's crust. Changes in SWS at Kīlauea Volcano, Hawai‘i, associated with the onset of summit eruptive activity in 2008 hint at the potential of the technique for tracking volcanic activity. To use SWS observations as a monitoring tool, however, it is important to understand the cause of seismic anisotropy at the volcano throughout the eruptive cycle. To address this need, we analyzed SWS results from across Kīlauea in combination with macroscopic surface structures (mapped fractures, faults, and fissures) and stress orientations inferred from fault plane solutions. Seismic anisotropy seems to be due to pervasive aligned structures in most regions of the volcano. The upper East and Southwest Rift Zones, however, show a bimodality in stress and SWS, suggesting a stress discontinuity with depth, perhaps related to magma conduits that trend obliquely to the dominant structure. Other areas in and around Kīlauea Caldera display principal stresses of similar magnitudes, indicating that small stress perturbations can rotate the maximum horizontal compressive stress direction by up to 90 ° . In these locations, static structures generally control SWS, but dynamic conditions due to magmatic activity can override the structural control. Monitoring of SWS may therefore provide important signs of impending volcanism.

Hawaii

Genetic analysis of harvest samples reveals population structure in a highly mobile generalist carnivore

Delineating wildlife population boundaries is important for effective population monitoring and management. The bobcat ( Lynx rufus ) is a highly mobile generalist carnivore that is ecologically and economically important. We sampled 1225 bobcats harvested in South Dakota, USA (2014–2019), of which 878 were retained to assess genetic diversity and infer population genetic structure using 17 microsatellite loci. We assigned individuals to genetic clusters ( K ) using spatial and nonspatial Bayesian clustering algorithms and quantified differentiation ( F ST and GST″ ) among clusters. We found support for population genetic structure at K = 2 and K = 4, with pairwise F ST and GST″ values indicating weak to moderate differentiation, respectively, among clusters. For K = 2, eastern and western clusters aligned closely with historical bobcat management units and were consistent with a longitudinal suture zone for bobcats previously identified in the Great Plains. We did not observe patterns of population genetic structure aligning with major rivers or highways. Genetic divergence observed at K = 4 aligned roughly with ecoregion breaks and may be associated with environmental gradients, but additional sampling with more precise locational data may be necessary to validate these patterns. Our findings reveal that cryptic population structure may occur in highly mobile and broadly distributed generalist carnivores, highlighting the importance of considering population structure when establishing population monitoring programs or harvest regulations. Our study further demonstrates that for elusive furbearers, harvest can provide an efficient, broad-scale sampling approach for genetic population assessments.

South Dakota

Evaluating the importance of characterizing soil structure and horizons in parameterizing a hydrologic process model

Incorporating the influence of soil structure and horizons into parameterizations of distributed surface water/groundwater models remains a challenge. Often, only a single soil unit is employed, and soil-hydraulic properties are assigned based on textural classification, without evaluating the potential impact of these simplifications. This study uses a distributed physics-based model to assess the influence of soil horizons and structure on effective parameterization. This paper tests the viability of two established and widely used hydrogeologic methods for simulating runoff and variably saturated flow through layered soils: (1) accounting for vertical heterogeneity by combining hydrostratigraphic units with contrasting hydraulic properties into homogeneous, anisotropic units and (2) use of established pedotransfer functions based on soil texture alone to estimate water retention and conductivity, without accounting for the influence of pedon structures and hysteresis. The viability of this latter method for capturing the seasonal transition from runoff-dominated to evapotranspiration-dominated regimes is also tested here. For cases tested here, event-based simulations using simplified vertical heterogeneity did not capture the state-dependent anisotropy and complex combinations of runoff generation mechanisms resulting from permeability contrasts in layered hillslopes with complex topography. Continuous simulations using pedotransfer functions that do not account for the influence of soil structure and hysteresis generally over-predicted runoff, leading to propagation of substantial water balance errors. Analysis suggests that identifying a dominant hydropedological unit provides the most acceptable simplification of subsurface layering and that modified pedotransfer functions with steeper soil-water retention curves might adequately capture the influence of soil structure and hysteresis on hydrologic response in headwater catchments.

Hydrological Processes

Influence of population density and length structure on angler catch rate in kokanee fisheries

Management agencies are often charged with providing fisheries that lead to angler participation. Catch rate is one of the primary drivers of angler participation but can be influenced by a suite of factors, including population structure (e.g., density and size structure). The complexity of understanding how population structure influences angler catch rate is typified in kokanee Oncorhynchus nerka fisheries. Previous research suggests that angler catch rates of kokanee are positively influenced by fish density and total length. However, that research was based on data collected using size-selective midwater trawls. Due to the potential limitation of previous research, we sought to (1) understand the relative bias of midwater trawls and gill nets for describing the size structure of kokanee available to anglers and (2) re-evaluate the influence of fish density and fish length on angler catch rates in kokanee fisheries. Midwater trawl, gill-net, and creel data were collected on five prominent kokanee fisheries throughout Idaho in 2016 and 2017. Catch composition and percent overlap of midwater trawls, gill nets, and angler-caught fish were compared to understand the efficacy of midwater trawls and gill nets for representing the size structure of kokanee available to anglers. In addition, the influence of kokanee density and length on angler catch rates was evaluated. Midwater trawls primarily sampled small kokanee (<330 mm) and exhibited little overlap with angler-caught fish, whereas gill nets sampled more large fish (>330 mm) and exhibited higher overlap with angler-caught fish when compared to midwater trawls. Fish length was not positively associated with angler catch rates. However, fish density exhibited a positive relationship with angler catch rates. Our results highlight the importance of gear choice for understanding how kokanee populations function and elucidate the tradeoffs associated with population density, fish length, and resulting kokanee fisheries.

Idaho

Fish assemblage structure and habitat associations in a large western river system

Longitudinal gradients of fish assemblage and habitat structure were investigated in the Kootenai River of northern Idaho. A total of 43&thinsp;500-m river reaches was sampled repeatedly with several techniques (boat-mounted electrofishing, hoop nets and benthic trawls) in the summers of 2012 and 2013. Differences in habitat and fish assemblage structure were apparent along the longitudinal gradient of the Kootenai River. Habitat characteristics (e.g. depth, substrate composition and water velocity) were related to fish assemblage structure in three different geomorphic river sections. Upper river sections were characterized by native salmonids (e.g. mountain whitefish Prosopium williamsoni ), whereas native cyprinids (peamouth Mylocheilus caurinus , northern pikeminnow Ptychocheilus oregonensis ) and non-native fishes (pumpkinseed Lepomis gibbosus , yellow perch Perca flavescens ) were common in the downstream section. Overall, a general pattern of species addition from upstream to downstream sections was discovered and is likely related to increased habitat complexity and additions of non-native species in downstream sections. Assemblage structure of the upper sections were similar, but were both dissimilar to the lower section of the Kootenai River. Species-specific hurdle regressions indicated the relationships among habitat characteristics and the predicted probability of occurrence and relative abundance varied by species. Understanding fish assemblage structure in relation to habitat could improve conservation efforts of rare fishes and improve management of coldwater river systems.

Idaho

RAD-seq refines previous estimates of genetic structure in Lake Erie walleye

Delineating population structure helps fishery managers to maintain a diverse “portfolio” of local spawning populations (stocks), as well as facilitate stock-specific management. In Lake Erie, commercial and recreational fisheries for Walleye Sander vitreus exploit numerous local spawning populations, which cannot be easily differentiated using traditional genetic data (e.g., microsatellites). Here, we used genomic information (12,264 polymorphic loci) generated using restriction site-associated DNA sequencing to investigate stock structure in Lake Erie Walleye. We found low genetic divergence (genetic differentiation index F ST = 0.0006–0.0019) among the four Lake Erie western basin stocks examined, which resulted in low classification accuracies for individual samples (40–60%). However, more structure existed between western and eastern Lake Erie basin stocks ( F ST = 0.0042–0.0064), resulting in greater than 95% classification accuracy of samples to a lake basin. Thus, our success in using genomics to identify stock structure varied with spatial scale. Based on our results, we offer suggestions to improve the efficacy of this new genetic tool for refining stock structure and eventually determining relative stock contributions in Lake Erie Walleye and other Great Lakes populations.

Michigan, New York, Ohio, Ontario, Pennsylvania

A crystal-chemical classification of borate structures with emphasis on hydrated borates

The rules governing formation of hydrated borate polyanions that were proposed by C.L. Christ in 1960 are critically reviewed and new rules added on the basis of recent crystal structure determinations. Principles and classifications previously published by others are also critically reviewed briefly. The fundamental building blocks from which borate polyanions can be constructed are defined on the basis of the number n of boron atoms, and the fully hydrated polyanions are illustrated. Known structures are grouped accordingly, and a shorthand notation using n and symbols ?? = triangle, T = tetrahedron is introduced so that the polyanions can be easily characterized. For example, 3:??+2T describes [B3O3(OH)5]2-. Correct structural formulas are assigned borates with known structures whereas borates of unknown structure are grouped separately. ?? 1977 Springer-Verlag.

Physics and Chemistry of Minerals

Temporal genetic population structure and interannual variation in migration behavior of Pacific Lamprey Entosphenus tridentatus

Studies using neutral loci suggest that Pacific lamprey, Entosphenus tridentatus , lack strong spatial genetic population structure. However, it is unknown whether temporal genetic population structure exists. We tested whether adult Pacific lamprey: (1) show temporal genetic population structure; and (2) migrate different distances between years. We non-lethally sampled lamprey for DNA in 2009 and 2010 and used eight microsatellite loci to test for genetic population structure. We used telemetry to record the migration behaviors of these fish. Lamprey were assignable to three moderately differentiated genetic clusters ( F ST = 0.16–0.24 for all pairwise comparisons): one cluster was composed of individuals from 2009, and the other two contained individuals from 2010. The F ST value between years was 0.13 and between genetic clusters within 2010 was 0.20. A total of 372 (72.5%) fish were detected multiple times during their migrations. Most fish (69.9%) remained in the mainstem Willamette River; the remaining 30.1% migrated into tributaries. Eighty-two lamprey exhibited multiple back-and-forth movements among tributaries and the mainstem, which may indicate searching behaviors. All migration distances were significantly greater in 2010, when the amplitude of river discharge was greater. Our data suggest genetic structuring between and within years that may reflect different cohorts.

Oregon

Predicting bird guilds using vegetation composition and structure on a wild and scenic river in Arizona

Riparian areas are among the most ecologically diverse terrestrial ecosystems but make up <2% of landscape area in southwestern USA. Many species of resident and neotropical migratory birds utilize riparian habitats for breeding, foraging, and nesting. We quantified vegetation composition and structure to predict bird guilds on Wild and Scenic portions of the Verde River, Arizona. We grouped plant species into guilds based on similar functional traits to describe composition. We surveyed birds during the breeding and migrating season to determine abundance and categorized species into guilds using preferences of breeding habitat, foraging substrate, and nest placement. Riparian obligate and facultative breeding guilds were most common. Both vegetation composition and structure were useful predictors of birds. Vegetation structure was most complex in gallery riparian forest. Abundance of riparian-obligate birds in the breeding guild were positively associated with vegetation structure of dense, multi-canopy canopy and tall trees. Abundance of most bird guilds were positively associated with composition of tall trees ( Populus fremontii , Salix gooddingii ) and drought tolerant shrubs ( Prosopis velutina, Celtis reticulata ). Our findings show complex riparian habitat important to wildlife is created by both composition and structure of near-stream vegetation that is tied to hydrology and sensitive to flow change.

Arizona

The crystal structure of cesium biuranyl trisulphate, Cs2(UO2)2(SO4)3

The crystal structure of the new compound Cs(UO 2 ) 2 (SO 4 ) 3 has been determined by X-ray diffraction methods. The compound is tetragonal, space group P 4 2 1 m "> P42 1 m ( D 2 d 3 ), with a = 9·62 ± 0·02, c = 8&#xB7;13 &#xB1; 0&#xB7;01 A &#x30A; "> c = 8·13 ± 0·01Å , and Z = 2; s.g. (calc.) = 4·80 ± 0·03, s.g. (obs.) = 4·74 ± 0·05. The compound forms plates parallel to (001) bounded by the form (110). Intensity data were obtained from Buerger precession photographs of the ( hk 0) and (0 kl ) reciprocal lattice nets. No corrections for absorption were made. The co-ordinates of the U and Cs atoms were obtained by interpretation of the Patterson projections normal to (001) and (100) and a plausible structure was derived from electron density projections. The final parameters of the structure were determined from subtraction electron density maps, least squares analysis of the structure factors, and spatial considerations. The compound has a layer structure consisting of (UO 2 ) 2 (SO 4 ) 3 ] n 2 n − sheets paralle to (001), tied together by cesium ions. The UO 2 2+ group is co-ordinated by five sulphate oxygens which form a nearly plane pentagon approximately normal to the uranyl axis. The Cs 2 atom is co-ordinated by twelve oxygen atoms and the Cs 1 atom by eight oxygen atoms. X-ray and optical data are also given for the compound Rb 2 UO 2 (SO 4 ) 2 ·2H 2 O.

Journal of Inorganic and Nuclear Chemistry

Faunal impact on vegetation structure and ecosystem function in mangrove forests: A review

The last 20 years witnessed a real paradigm shift concerning the impact of biotic factors on ecosystem functions as well as on vegetation structure of mangrove forests. Before this small scientific revolution took place, structural aspects of mangrove forests were viewed to be the result of abiotic processes acting from the bottom-up, while, at ecosystem level, the outwelling hypothesis stated that mangroves primary production was removed via tidal action and carried to adjacent nearshore ecosystems where it fuelled detrital based food-webs. The sesarmid crabs were the first macrofaunal taxon to be considered a main actor in mangrove structuring processes, thanks to a number of studies carried out in the Indo-Pacific forests in the late 1970s and early 1980s. Following these classical papers, a number of studies on Sesarmidae feeding and burrowing ecology were carried out, which leave no doubts about the great importance of these herbivorous crabs in structuring and functioning Old world ecosystems. Although Sesarmidae are still considered very important in shaping mangrove structure and functioning, recent literature emphasizes the significance of other invertebrates. The Ocypodidae have now been shown to have the same role as Sesarmidae in terms of retention of forest products and organic matter processing in New world mangroves. In both New and Old world mangroves, crabs process large amounts of algal primary production, contribute consistently to retention of mangrove production and as ecosystem engineers, change particle size distribution and enhance soil aeration. Our understanding of the strong impact of gastropods, by means of high intake rates of mangrove products and differential consumption of propagules, has changed only recently. The role of insects must also be stressed. It is now clear that older techniques used to assess herbivory rates by insects strongly underestimate their impact, both in case of leaf eating and wood boring species and that herbivorous insects can potentially play a strong role in many aspects of mangrove ecology. Moreover, researchers only recently realized that ant&ndash;plant interactions may form an important contribution to our understanding of insect&ndash;plant dynamics in these habitats. Ants seem to be able to relieve mangroves from important herbivores such as many insects and sesarmid crabs. It thus seems likely that ants have positive effects on mangrove performance.

Aquatic Botany

Sensitivity of system stability to model structure

A community is stable, and resilient, if the levels of all community variables can return to the original steady state following a perturbation. The stability properties of a community depend on its structure, which is the network of direct effects (interactions) among the variables within the community. These direct effects form feedback cycles (loops) that determine community stability. Although feedback cycles have an intuitive interpretation, identifying how they form the feedback properties of a particular community can be intractable. Furthermore, determining the role that any specific direct effect plays in the stability of a system is even more daunting. Such information, however, would identify important direct effects for targeted experimental and management manipulation even in complex communities for which quantitative information is lacking. We therefore provide a method that determines the sensitivity of community stability to model structure, and identifies the relative role of particular direct effects, indirect effects, and feedback cycles in determining stability. Structural sensitivities summarize the degree to which each direct effect contributes to stabilizing feedback or destabilizing feedback or both. Structural sensitivities prove useful in identifying ecologically important feedback cycles within the community structure and for detecting direct effects that have strong, or weak, influences on community stability. The approach may guide the development of management intervention and research design. We demonstrate its value with two theoretical models and two empirical examples of different levels of complexity. ?? 2009 Elsevier B.V. All rights reserved.

Ecological Modelling

Space matters: Host spatial structure and the dynamics of plague transmission

The development of models to elucidate the transmission pathways and dynamics of wildlife diseases remains challenging. Sylvatic plague, caused by the bacterium Yersinia pestis ( Yp ), is an infectious zoonotic disease that primarily affects wild rodents, including prairie dogs ( Cynomys spp.) in North America. Proposed transmission pathways for Yp include flea bites, direct contacts between hosts, and environmental reservoirs (e.g. soil, carcasses). We developed a spatially explicit, agent-based model of Yp transmission to explore the effects of alternative transmission pathways, different disease initiation mechanisms (host or fleas), parameter uncertainty, and spatial structure of hosts. A particularly novel aspect of our model was the integration of ecological models with traditional disease models. Specifically, we used estimates from spatial capture-recapture models to generate data-driven spatial distributions, densities, and contact rates to capture the spatial structure of prairie dogs. We simulated ~9 million scenarios across a wide range of parameter values and conducted sensitivity analyses to determine the most influential parameters on the number of flea-days (sum of the mean number of fleas on hosts each day of the simulation), number of newly infected hosts per day, the time to depopulation (<20 prairie dogs remaining), and the proportion of the prairie dog population remaining at the end of the simulation (after 150 days). When including spatial structure, we found the probability of transmission via environmental sources of Yp (i.e. carcasses) had the greatest influence on model results when Yp infection was initiated in prairie dog hosts, rather than in fleas. Conversely, the mechanism of transmission by fleas to prairie dogs had the greatest influence on model results when Yp infection was initiated in fleas (i.e. via introduction by carnivores, a migrant prairie dog, or other mammalian host). Uncertainty in parameter estimates, particularly those related to the transmission pathways of Yp, continue to hamper efforts to realistically model plague dynamics in wild rodents. Our results elucidate the complexity of the flea-plague-prairie dog system and reiterate the importance of research on Yp transmission mechanisms to provide a full understanding of this disease. Our results also emphasize the importance of realistic estimates of spatial structure for exploring transmission dynamics of wildlife diseases and provide a framework for generating a data-driven description of spatial structure.

Ecological Modelling

Structural characterization of terrestrial microbial Mn oxides from Pinal Creek, AZ

The microbial catalysis of Mn(II) oxidation is believed to be a dominant source of abundant sorption- and redox-active Mn oxides in marine, freshwater, and subsurface aquatic environments. In spite of their importance, environmental oxides of known biogenic origin have generally not been characterized in detail from a structural perspective. Hyporheic zone Mn oxide grain coatings at Pinal Creek, Arizona, a metals-contaminated stream, have been identified as being dominantly microbial in origin and are well studied from bulk chemistry and contaminant hydrology perspectives. This site thus presents an excellent opportunity to study the structures of terrestrial microbial Mn oxides in detail. XRD and EXAFS measurements performed in this study indicate that the hydrated Pinal Creek Mn oxide grain coatings are layer-type Mn oxides with dominantly hexagonal or pseudo-hexagonal layer symmetry. XRD and TEM measurements suggest the oxides to be nanoparticulate plates with average dimensions on the order of 11 nm thick × 35 nm diameter, but with individual particles exhibiting thickness as small as a single layer and sheets as wide as 500 nm. The hydrated oxides exhibit a 10-Å basal-plane spacing and turbostratic disorder. EXAFS analyses suggest the oxides contain layer Mn(IV) site vacancy defects, and layer Mn(III) is inferred to be present, as deduced from Jahn–Teller distortion of the local structure. The physical geometry and structural details of the coatings suggest formation within microbial biofilms. The biogenic Mn oxides are stable with respect to transformation into thermodynamically more stable phases over a time scale of at least 5 months. The nanoparticulate layered structural motif, also observed in pure culture laboratory studies, appears to be characteristic of biogenic Mn oxides and may explain the common occurrence of this mineral habit in soils and sediments.

Geochimica et Cosmochimica Acta

A field investigation of the basaltic ring structures of the Channeled Scabland and the relevance to Mars

The basaltic ring structure (BRS) is a class of peculiar features only reported in the Channeled Scabland of eastern Washington State. They have been suggested to be good analogs, however, for some circular features on Mars. BRSs are found where Pleistocene floods scoured the Columbia River Basin, stripping off the uppermost part of the Miocene Columbia River Basalt Group and exposing structures that were previously embedded in the lava. The &ldquo;Odessa Craters,&rdquo; near Odessa, WA, are 50&ndash;500-m-wide BRSs that are comprised of discontinuous, concentric outcrops of subvertically-jointed basalt and autointrusive dikes. Detailed field investigation of the Odessa Craters in planform and a cross-sectional exposure of a similar structure above Banks Lake, WA, lead us to propose that BRSs formed by concurrent phreatovolcanism and lava flow inflation. In this model, phreatovolcanic (a.k.a., &ldquo;rootless&rdquo;) cones formed on a relatively thin, active lava flow; the lava flow inflated around the cones, locally inverting topography; tensile stresses caused concentric fracturing of the lava crust; lava from within the molten interior of the flow exploited the fractures and buried the phreatovolcanic cones; and subsequent erosive floods excavated the structures. Another population of BRSs near Tokio Station, WA, consists of single-ringed, raised-rimmed structures that are smaller and more randomly distributed than the Odessa Craters. We find evidence for a phreatovolcanic component to the origin as well, and hypothesize that they are either flood-eroded phreatovolcanic cones or Odessa Crater-like BRSs. This work indicates that BRSs are not good analogs to the features on Mars because the martian features are found on the uneroded surfaces. Despite this, the now superseded concepts for BRS formation are useful for understanding the formation of the martian features.

Washington

A review of sea lamprey dispersal and population structure in the Great Lakes and the implications for control

Understanding the population structure of invasive sea lamprey ( Petromyzon marinus ) in the Great Lakes basin is essential for an effective control program. We review knowledge of lake connectivity, dispersal during the parasitic stage, and results from phenotypic, demographic, and genetic studies to evaluate how sea lamprey populations are structured. There is no evidence for contemporary movement between Lake Ontario and the Atlantic population, although it appears possible. Dispersal between Lake Ontario and the Finger Lakes is more likely, as is contemporary movement between Lakes Ontario and Erie via the Welland Canal, although neither has been directly observed. Downstream movement from Lake Erie to Lake Ontario via the Niagara River has been reported. Bidirectional movement between Lakes Erie and Huron has been observed, and movement of sea lamprey among the upper Great Lakes (especially between Lakes Huron and Michigan) is relatively common, although complete mixing likely does not occur. The maximum straight-line dispersal distance reported for a tagged sea lamprey was 628 km between the St. Marys River and western Lake Erie. Genetic population studies using a variety of molecular markers generally found weak but significant broad-scale population structure (e.g., between freshwater and anadromous populations, and among Lake Ontario, Lake Erie, and the upper Great Lakes), but finer-scale structure was rarely detected. Nevertheless, some within-basin structure is suggested by regional differences in phenotypic and demographic traits (e.g., sex ratio, body size). Further study will be important because management is most efficiently targeted when the geography of demographically independent populations is well-characterized.

Lake Ontario

Assessing fire effects on forest spatial structure using a fusion of Landsat and airborne LiDAR data in Yosemite National Park

Mosaics of tree clumps and openings are characteristic of forests dominated by frequent, low- and moderate-severity fires. When restoring these fire-suppressed forests, managers often try to reproduce these structures to increase ecosystem resilience. We examined unburned and burned forest structures for 1937 0.81 ha sample areas in Yosemite National Park, USA. We estimated severity for fires from 1984 to 2010 using the Landsat-derived Relativized differenced Normalized Burn Ratio (RdNBR) and measured openings and canopy clumps in five height strata using airborne LiDAR data. Because our study area lacked concurrent field data, we identified methods to allow structural analysis using LiDAR data alone. We found three spatial structures, canopy-gap, clump-open, and open, that differed in spatial arrangement and proportion of canopy and openings. As fire severity increased, the total area in canopy decreased while the number of clumps increased, creating a patchwork of openings and multistory tree clumps. The presence of openings > 0.3 ha, an approximate minimum gap size needed to favor shade-intolerant pine regeneration, increased rapidly with loss of canopy area. The range and variation of structures for a given fire severity were specific to each forest type. Low- to moderate-severity fires best replicated the historic clump-opening patterns that were common in forests with frequent fire regimes. Our results suggest that managers consider the following goals for their forest restoration: 1) reduce total canopy cover by breaking up large contiguous areas into variable-sized tree clumps and scattered large individual trees; 2) create a range of opening sizes and shapes, including ~ 50% of the open area in gaps > 0.3 ha; 3) create multistory clumps in addition to single story clumps; 4) retain historic densities of large trees; and 5) vary treatments to include canopy-gap, clump-open, and open mosaics across project areas to mimic the range of patterns found for each forest type in our study.

California

Remote sensing of dryland ecosystem structure and function: Progress, challenges, and opportunities

Drylands make up roughly 40% of the Earth's land surface, and billions of people depend on services provided by these critically important ecosystems. Despite their relatively sparse vegetation, dryland ecosystems are structurally and functionally diverse, and emerging evidence suggests that these ecosystems play a dominant role in the trend and variability of the terrestrial carbon sink. More, drylands are highly sensitive to climate and are likely to have large, non-linear responses to hydroclimatic change. Monitoring the spatiotemporal dynamics of dryland ecosystem structure (e.g., leaf area index) and function (e.g., primary production and evapotranspiration) is therefore a high research priority. Yet, dryland remote sensing is defined by unique challenges not typically encountered in mesic or humid regions. Major challenges include low vegetation signal-to-noise ratios, high soil background reflectance, presence of photosynthetic soils (i.e., biological soil crusts), high spatial heterogeneity from plot to regional scales, and irregular growing seasons due to unpredictable seasonal rainfall and frequent periods of drought. Additionally, there is a relative paucity of continuous, long-term measurements in drylands, which impedes robust calibration and evaluation of remotely-sensed dryland data products. Due to these issues, remote sensing techniques developed in other ecosystems or for global application often result in inaccurate, poorly constrained estimates of dryland ecosystem structural and functional dynamics. Here, we review past achievements and current progress in remote sensing of dryland ecosystems, including a detailed discussion of the major challenges associated with remote sensing of key dryland structural and functional dynamics. We then identify strategies aimed at leveraging new and emerging opportunities in remote sensing to overcome previous challenges and more accurately contextualize drylands within the broader Earth system. Specifically, we recommend: 1) Exploring novel combinations of sensors and techniques (e.g., solar-induced fluorescence, thermal, microwave, hyperspectral, and LiDAR) across a range of spatiotemporal scales to gain new insights into dryland structural and functional dynamics; 2) utilizing near-continuous observations from new-and-improved geostationary satellites to capture the rapid responses of dryland ecosystems to diurnal variation in water stress; 3) expanding ground observational networks to better represent the heterogeneity of dryland systems and enable robust calibration and evaluation; 4) developing algorithms that are specifically tuned to dryland ecosystems by utilizing expanded ground observational network data; and 5) coupling remote sensing observations with process-based models using data assimilation to improve mechanistic understanding of dryland ecosystem dynamics and to better constrain ecological forecasts and long-term projections.

Remote Sensing of Environment