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At least 199 records · Page 11Linked to original sources

Morphological traits related to potential invasiveness of two subspecies of the crayfish Faxonius neglectus

Biological invasions have major environmental and economic impacts, and pose a serious threat to global biodiversity. Invasive crayfish species are one of the greatest threats to native crayfish biodiversity. Additionally, almost 50% of US and Canadian species are considered at risk, making crayfish one of the most imperiled taxonomic groups in the world. Small-scale (extralimital) invasions are often overlooked and may be more common than large-scale (extraregional) invasions. One example of an extraregional and extralimital invader is the Ringed Crayfish ( Faxonius neglectus ), which has been independently introduced multiple times to drainages throughout the United States, including those adjacent to its native range. Traits related to invasiveness, such as chelae size, are suggested to differ between invasive crayfish from extralimital and extraregional source populations with larger size related to increased invasiveness. We examined morphological traits related to invasion potential for both subspecies of F . neglectus , F . neglectus neglectus and F . neglectus chaenodactylus . We sampled 28 stream sites within the known native range of F . neglectus , including the Neosho and Upper White River drainages in Oklahoma, Arkansas and Missouri. Total carapace length, chelae length and chelae width of 30 adult male F . neglectus were measured from each site. We found significant differences in crayfish morphological characteristics among stream sites. We found significantly greater chelae length:carapace length (ChL:CarL) and chelae width:chelae length (ChW:ChL) in F . neglectus chaenodactylus than F . neglectus neglectus . Even among F . neglectus neglectus populations, there were significant differences in ChL:CarL and ChW:ChL. Morphological characteristics suggest that F . neglectus chaenodactylus and some populations of F . neglectus neglectus may be pre-adapted to the role of invader.

Arkansas, Kansas, Missouri, Oklahoma

Evaluating hydrologic region assignment techniques for ungaged basins in Alaska, USA

Building continental-scale hydrologic models in data-sparse regions requires an understanding of spatial variation in hydrologic processes. Extending these models to ungaged locations requires techniques to group ungaged locations with gaged ones to make process importance and model parameter transfer decisions to ungaged locations. This analysis (1) tested the utility of fundamental streamflow statistics (FDSS) in defining hydrologic regions across Alaska, USA; (2) evaluated if the hydrologic regions represented different hydrologic processes; and (3) tested the ability of random forest and direct assignment techniques, informed by statistically estimated FDSS (FDSSest) and basin characteristics (BCs), to correctly assign ungaged locations to hydrologic regions. Six hydrologic regions were identified across the domain using FDSS. Differences in mean flow, phase shift of the seasonal cycle, and skewness were the primary characteristics defining each region. Two regions represented arctic and continental climates, generally in the northern portion of the domain; four regions represented the southern, maritime portion of the domain. Random forest modeling with BCs (67% success rate) outperformed FDSSest (58% success rate) suggesting that no statistically estimated streamflow was needed to assign ungaged locations to a region. For regions with many sites, most region assignment techniques performed similarly. Random forest modeling performance declined when BCs and FDSSest were both used to predict region membership, suggesting FDSSest had little information in addition to BCs. This analysis demonstrated that FDSS-based hydrologic regions discern process differences across a data-sparse and hydrologically diverse landscape. Process importance rankings from random forest-derived BCs provided model-independent information for making modeling decisions.

Alaska

Identifying nutrient sources and sinks to the South Platte River and Cherry Creek, Denver, CO, during low-flow conditions in 2019–2020

Elevated concentrations and loads of nutrients in the South Platte River and Cherry Creek in Denver, Colorado, may have adverse effects on those streams and downstream water bodies, including increased production of algae, eutrophication, and decreased recreational opportunities. This article describes streamflow and concentrations and loads of nutrients for the South Platte River and Cherry Creek in Denver based on data collected during two longitudinal Lagrangian sampling campaigns in low-flow conditions in fall of 2019 and 2020. The results are used to assess sources and sinks of nutrients in the study area and help to establish baseline conditions against which future changes in nutrient concentrations and loads can be assessed. Discharges from Chatfield and Cherry Creek Reservoirs, storm drains, and most tributaries to the South Platte River, and Cherry Creek were generally small sources of streamflow and nutrient loads in both years. The Marcy Gulch, South Platte Water Renewal, and Robert W. Hite wastewater treatment plants were larger sources of streamflow and nutrient loads. The Burlington Ditch was a sink for streamflow and nutrient loads, diverting more than 95% of the South Platte River during the two sampling campaigns. Most other sinks were associated with decreases in streamflow between sampling sites. Golf courses were a potential source of nutrients for Cherry Creek but not for the South Platte River.

Colorado

The hydroclimate niche: A tool for predicting and managing riparian plant community responses to streamflow seasonality

Habitat suitability is a consequence of interacting environmental factors. In riparian ecosystems, suitable plant habitat is influenced by interactions between stream hydrology and climate, hereafter referred to as “hydroclimate”. We tested the hypothesis that hydroclimate variables would improve the fit of ecological niche models for a suite of riparian species using occurrence data from the western United States. We focus on the climate conditions (temperature, precipitation and vapor pressure deficit) during the months of lowest and highest streamflow as integrative hydroclimate metrics of resource and stress levels. We found that the inclusion of hydroclimate variables improved model fit for all species in the western USA dataset. We then tested the utility of the improved habitat suitability models by projecting them onto a regulated segment of the Colorado River to assess potential impacts of streamflow seasonality on vegetation metrics of management concern. Species frequency derived from independent survey data in the Colorado River segment was significantly higher for species with predicted suitable habitat than for species without predicted suitable habitat. Under different simulated hydrographs for the Colorado River, overall species richness was predicted to be greatest with peak streamflows during summer, and native-to-non-native species ratios were predicted to be greatest with lowest streamflows in winter. Summer high flows were particularly associated with higher predicted habitat suitability for species that have increased in cover over recent decades (e.g., Pluchea sericea, Baccharis species). We conclude that hydroclimate covariates can be useful tools for predicting how riparian vegetation communities respond to changes in the seasonal timing of low and high streamflows.

River Research and Applications

High resolution spatiotemporal patterns of flow at the landscape scale in montane non-perennial streams

Intermittent and ephemeral streams in dryland environments support diverse assemblages of aquatic and terrestrial life. Understanding when and where water flows provide insights into the availability of water, its response to external controlling factors, and potential sensitivity to climate change and a host of human activities. Knowledge regarding the timing of drying/wetting cycles can also be useful to map critical habitats for species and ecosystems that rely on these temporary water sources. However, identifying the locations and monitoring the timing of streamflow and channel sediment moisture remains a challenging endeavor. In this paper, we analyzed daily conductivity from 37 sensors distributed along 10 streams across an arid mountain front in Arizona (United States) to assess spatiotemporal patterns in flow permanence, defined as the timing and extent of water in streams. Conductivity sensors provide information on surface flow and sediment moisture, supporting a stream classification based on seasonal flow dynamics. Our results provide insight into flow responses to seasonal rainfall, highlighting stream reaches very reactive to rainfall versus those demonstrating more stable streamflow. The strength of stream responses to precipitation are explored in the context of surficial geology. In summary, conductivity data can be used to map potential stream habitat for water-dependent species in both space and time, while also providing the basis upon which sensitivity to ongoing climate change can be evaluated.

Arizona

Climate change impacts on native cutthroat trout habitat in Colorado streams

Headwater streams support vital aquatic habitat yet are vulnerable to changing climate due to their high elevation and small size. Coldwater fish are especially sensitive to the altered streamflow and water temperature regimes during summer low flow periods. Though previous studies have provided insights on how changes in climate and alterations in stream discharge may affect habitat availability for various native cutthroat trout species, suitable physical habitats have not been evaluated under future climate projections for the threatened Greenback Cutthroat Trout (GBCT) native to headwater regions of Colorado, USA. Thus, this study used field data collected from selected headwater streams across the current distribution of GBCT to construct one-dimensional hydraulic models to evaluate streamflow and physical habitat under four future climate projections. Results illustrate reductions in both predicted streamflow and physical habitat for all future climate projections across study sites. The projected mean summer streamflow shows greater decline (−52% on average) compared to the projected decline in mean August flow (−21% on average). Moreover, sites located at a relative higher elevation with larger substrate and steeper slope were projected to experience more reductions in physical habitat due to streamflow reductions. Specifically, streams with step-pool morphologies may experience grater changes in available habitat compared to pool-riffle streams. Future climate change studies related to coldwater fish that examine spatial variation in flow alteration could provide novel data to complement the existing literature on the thermal characteristics. Tailoring reintroduction and management efforts for GBCT to the individual headwater stream with adequate on-site monitoring could provide a more holistic conservation approach.

Colorado

Ecogeomorphic interactions of Russian olives (Elaeagnus angustifolia) and point-bar morphology along Powder River, Montana, USA

Point bars are crucial alluvial features that help sustain meandering river ecosystems. They are influenced by interactions among physical processes and biological components, such as riparian vegetation. Understanding the mechanisms underlying these interactions can inform management decisions. The meandering Powder River in southeastern Montana, USA, provides a valuable study system for fluvial interactions because it has no major human alterations. Riparian stands of non-native Russian olive trees ( Elaeagnus angustifolia ) became noticeable after ~50-year flood in 1978. Fortuitously, 20 channel cross-sections had been established in 1975 and 1977 and have been surveyed periodically ever since. This temporally extensive dataset was coupled with a spatially extensive dataset of 85 point-bar cross-sections derived from high-resolution LiDAR that enabled evaluation of interactions between Russian olives and point-bar morphology through time and space. Based on published literature and field observations, we hypothesised that an increase in the width of Russian olive stands would result in (1) a decrease in the overall point-bar slope, (2) an increase in point-bar heights and (3) in point-bar toe-slopes. The results indicated that point-bar slopes did decrease with an increase in stand width, supporting the first hypothesis ( R 2 = 0.87). However, point-bar heights decreased with an increase in stand width ( R 2 = 0.63), contradicting the second hypothesis and there was no relation between point-bar toe-slopes and stand width ( R 2 < 0.1). The reverse hypothesis that point-bar morphology controls Russian olive stand widths makes more biophysical sense. Thus, Russian olive stand width increases as the point-bar slope and height decrease ( R 2 = 0.79 and 0.50), which provide a greater area of inundation for colonisation. This study demonstrates that geomorphology is controlling non-native vegetation colonisation along Powder River, but that the subsequent stabilisation associated with Russian oliveexpansion may in turn upset the critical geomorphic balance between bank erosion and point-bar aggradation necessary for a stable meandering system.

Montana

The Toolbox for River Velocimetry using Images from Aircraft (TRiVIA)

Accurate knowledge of the speed at which water moves along a river is essential for understanding ecohydraulic processes and managing natural resources. Measuring flow velocity via remote sensing can be more efficient than conventional field methods, and powerful computational techniques for inferring velocity fields from videos or image time series have been developed. The development of dedicated software tools for particle image velocimetry (PIV) could facilitate greater use of these methods by the river community. This paper introduces a standalone app designed for this exact purpose: the Toolbox for River Velocimetry using Images from Aircraft, or TRiVIA. The program provides a complete workflow for producing spatially distributed velocity vectors from a video or sequence of images, all within an accessible graphical user interface. TRiVIA includes modules for extracting and resampling frames, stabilization and geo-referencing images, defining a region of interest, enhancing images, performing PIV with an efficient ensemble correlation algorithm, visualizing results, assessing accuracy assessment, and exporting PIV output. We illustrate the software's capabilities using an example data set from a large river in Alaska. The initial release of the toolbox is now freely available. Augmenting TRiVIA to incorporate bathymetric information could enable discharge calculation functionality.

River Research and Applications

No evidence for cottonwood forest decline along a flow-augmented western U.S. river

In contrast to many other arid region rivers, streamflow in the South Platte River is heavily augmented by trans-basin water imports and irrigation return flows. Hydrological changes began in the 1880s, resulting in channel narrowing and the development of a continuous Populus-Salix forest by the mid-twentieth century. We assessed the composition, structure and regeneration status of the riparian forest and identified environmental variables affecting annual Populus deltoides tree growth. We sampled forest structure at four sites in 2015, and conducted dendroecological analysis at seven additional sites in 2019. The riparian forest was dominated by P. deltoides , which occurred at all sites, comprising 79% of total tree basal area and 62% of total tree density. Age structure data indicated ongoing though episodic recruitment of P. deltoides , at least over the past ~130 years. We tested 14 linear mixed effects models to describe the effect of climate and streamflow on individual tree growth (modeled as the log of BAI, n = 237 trees). The most parsimonious model selected with AICc explained 28.6% of BAI variability, and included hydrology and climate factors during the growing season (i.e., June–August streamflow, June–July PDSI), some aspects of off-season (i.e., previous November and March) streamflow, along with tree age and study site effects. The riparian forest developed in response to, and has been maintained by, current climate conditions and water management regimes. It may be negatively affected by future climate change and increased urban water demand in the basin.

Colorado

Putting down roots: Afforestation and bank cohesion of Icelandic Rivers

Riparian vegetation is widely recognized as a critical component of functioning fluvial systems. Human pressures on woody vegetation including riparian areas have had lasting effects, especially at high latitude. In Iceland, prior to human settlement, native downy birch woodlands covered approximately 15%–40% of the land area compared to 1%–2% today. Afforestation efforts include planting seedlings, protecting native forest remnants, and acquiring land areas as national forests. The planted and protected nature of vegetation along rivers within forests provides a unique opportunity to evaluate the various taxa within riparian zones and the channel stabilizing characteristics of the vegetation used in afforestation. We investigated bank properties, sediment textures, and root characteristics within riparian zones along four rivers in forests in Iceland. Bank sediment textures are dominantly sandy loam overlying coarser textures. Undercut banks are common because of erosion of the less cohesive subsurface layer. Quantitative root data indicate that the woody taxa have greater root densities, rooting depths, and more complex root structures than forbs or graminoids. The native downy birch has the highest root densities, with <1 mm roots most abundant. Modeling of added bank cohesion indicates that willow provides up to six times and birch up to four times more added cohesion to the coarse sediment textures comprising stream banks compared to no vegetation. We conclude that planting and protecting the native birch and willow helps to reduce bank erosion, especially where long-term grazing exclusion can be maintained.

River Research and Applications

Turbulence near a sandbar island in the lower Missouri River

River turbulence is spatially variable due to interactions between morphology of rivers and physical mechanics of flowing water. Understanding the variation of turbulence in rivers is important for characterizing transport processes of soluble and particulate materials in these systems. We present an exploratory effort to understand ecologically relevant flow patterns using measurements of mean flow and turbulence in a highly engineered river channel around an island in the lower Missouri River. Specifically, the profiles of mean river velocities were investigated to examine the logarithmic relation and associated parameters, including shear velocity and bed roughness. Turbulence intensity and Reynolds shear stress were compared with classic open-channel profiles and previously reported river data in the hydraulics literature. With the capability of pulse-to-pulse coherent Doppler velocity profiling in high spatial resolution, we estimated the profiles of turbulence dissipation rate using resolved one-dimensional velocity spectra. These measurement data allow us to examine the validity of turbulence production-dissipation balance and the classic open-channel profiles of turbulence statistics, including turbulence intensity, Reynolds shear stress, dissipation rate, and eddy viscosity. The field data show a strong variation of turbulence profiles in close vicinity of the river island. In shallow water depths close to the island, turbulence is substantially enhanced in comparison with classic open-channel profiles. Such turbulence enhancement is likely attributed to non-uniformity of the flow structures.

Missouri

Expansive, positive changes to fish habitat diversity following the formation of a valley plug in a degraded desert river

Widespread hydrologic alterations have simplified in-stream habitats in rivers globally, driving population declines and extirpations of many native fishes. Here, we examine how rapid geomorphic change in a historically degraded desert river has influenced habitat diversification and ecosystem persistence. In 2010, a large reach of the degraded and simplified lower San Rafael River (SRR), Utah, was impacted by the formation of a valley plug and began to shift from a homogenous, single-thread channel to a complex, multi-threaded riverscape. We combined field measurements and drone-collected imagery to document changes in fish habitat due to the valley plug. Our results demonstrate that in 2021, the affected reach was more diverse than any other stream reach along the SRR, containing 641% more diverse habitat (e.g., pools, riffles, and backwaters) than what was measured in 2015. The plug reach also retained water for periods beyond what was expected during seasonal drying, with the total extent of inundation within the riverscape increasing by over 2800%. Since the formation of the valley plug, riparian habitat has increased by 230% and channel networks have expanded to more than 50 distinct channels throughout the zone of influence. Our results provide evidence of successful self-restoration in a formerly highly degraded reach of desert river, and encourage new methods of desert river restoration. We aim to inform the use of large-scale, disruptive restoration actions like intentional channel occlusions, with the goal of mitigating the impacts of simplification and increasing habitat persistence in the face of exacerbated aridity in the desert Southwest.

Utah

Expansion of smallmouth bass distribution and habitat overlap with juvenile Chinook salmon in the Willamette River, Oregon

Smallmouth bass populations have expanded far beyond their native range and these predatory fish present a pervasive threat to native aquatic species throughout North America. In the western United States, smallmouth bass are now present in river and reservoir habitats where Pacific salmon are found and are considered a potential threat to salmon recovery in many locations. We conducted a study to determine if smallmouth bass are expanding their range in the mainstem Willamette River, Oregon, and developed a model to assess habitat overlap between smallmouth bass and juvenile Chinook salmon. Sampling during 2011–2022 revealed that the distribution of smallmouth bass had expanded throughout that timeframe to encompass the entire mainstem Willamette River, including important rearing habitats for juvenile Chinook salmon. The model predicted that smallmouth bass and juvenile Chinook salmon habitat overlap was substantial, highlighting the need for additional research to evaluate for potential negative impacts to salmon recovery in the basin. The model was also used to evaluate the efficacy of using flow management to reduce interactions between these two species, but the scenarios we examined suggested that this was not a viable option. These results highlight the need for continued research to assess interactions between smallmouth bass and juvenile salmon, and other native species of concern, in the Willamette River Basin. The development of the model is useful for resource managers to understand interactions between these species to prioritize locations for sampling in the future.

Oregon

Upstream experience and experimental translocation of invasive bigheaded carps results in increased upstream passage success at a navigation lock in a large river

Fish movements in regulated rivers can be challenging to study because anthropogenic modifications, such as locks and dams, can influence animal behavior. Upper Mississippi River Lock and Dam 19 (LD 19), for example, is an invasive carp movement bottleneck due to an impassable dam. Upstream fish passage at LD19 is restricted to the lock chamber, making it an optimal location to test invasive fish deterrents that could limit further range expansion. Evaluating the effectiveness of experimental deterrents requires baseline knowledge of fish movements and suitable sample sizes of fish encountering the deterrents to ensure adequate statistical power. Some evidence indicates fish with prior upstream experience may return upstream or challenge potential deterrents at a higher rate than fish without such experience. To test how previous upstream experience could increase the rate at which fish moved upstream through a navigation lock chamber, we compared upstream passage through LD 19 using bigheaded carp captured below the dam (downstream-origin) and two groups of bigheaded carp captured upstream from the dam: those that swam downstream on their own volition (upstream-origin fish) and those that were captured upstream and translocated downstream of LD 19 (translocated upstream-origin). Translocated upstream-origin carp demonstrated the highest rate of upstream passage, with 59% of the fish detected downstream from LD 19 passing upstream during our study. In contrast, downstream-origin carp made no upstream passages over 2 years. Fish origin was shown to influence upstream passage success. This may be an important consideration for fish passage studies and deterrent evaluations.

Illinois, Iowa

Evaluating the context dependency of large river nursery habitats for fishes in the lower Red River catchment

Both fine- and coarse-scale physicochemical conditions affect the quantity and quality of nursery habitats within riverine ecosystems. Nursery habitats in large, braided, and sandbed streams such as the lower Red River of Oklahoma, Texas, and Arkansas are not well described and likely vary among species. Identification of nursery habitats is important for developing proper conservation and management actions. We used an occupancy model framework to determine how hierarchical habitat factors related to the occupancy of 38 juvenile fish species. Our findings indicate that large river nursery habitats can generally be defined by reaches with off-channel slackwater habitat, having deep pools but shallow thalweg depths, typically located further away from dams, and with low percentages of limestone lithology. Species within the same genera often exhibited variable relationships with river slope, amount of large woody debris, channel shape, discharge, and position of reaches within the stream network. Our results indicate important species-specific relationships that define nursery habitats, indicating an important context dependency of nursery habitats even within fishes that are taxonomically similar. If the goal is to improve recruitment by native fishes, then consideration of the important species-specific differences would be beneficial if improvements are made to nursery habitats. Moreover, careful consideration of the effects of dam operations will help maintain proper connectivity to off-channel habitats important in downriver portions of the river network.

Arkansas, Louisiana, Oklahoma, Texas

How low is too low? Partnering with stakeholders and managers to define ecologically based low-flow thresholds in a perennial temperate river

Managing aquatic ecosystems for people and nature can be improved by collaboration among scientists, managers, decision-makers, and other stakeholders. Many collaborative and interdisciplinary approaches have been developed to address the management of freshwater ecosystems; however, there are still barriers to overcome. We worked as part of a regional stakeholder group comprising municipal water utility operators, conservation organizations, academic partners, and other stakeholders to understand the effects of low-flow and drought on ecological functions of the upper Flint River, Georgia (USA), a free-flowing river important for municipal water supply, recreation, and native biota. We used published literature and locally targeted studies to identify quantitative flow targets that could be used to inform water management and drought planning. Drawing from principles of Translational Ecology, we relied on an iterative process to develop information needs for the group and maintained communication and engagement throughout data collection, analysis, and synthesis. We identified three quantitative flow benchmarks to evaluate the ecological impacts of drought in the river. The results were valuable to both the water utilities represented in the working group and State regional water planning, which is used to guide water management strategies and permitting for the basin. We identified principles that were important for the successful engagement in the working group and helped to overcome the challenge of working across sectors and without direct authority guiding the implementation of our work. Interdisciplinary work and creative solutions are crucial to plan for and adapt to greater pressure on our water resources.

River Research and Applications

Wood you believe it? Experimental addition of nonnative wood enhances instream habitat for native dryland fishes

Flow alteration and riparian vegetation encroachment are causing habitat simplification with severe consequences for native fishes. To assess the effectiveness of enhancing simplified habitat in a large dryland river, we experimentally added invasive wood at 19 paired treatment and reference (no wood added) subreaches (50–100 m) within the main channel of the San Juan River. Using a before-after-control-impact design, we sampled fishes and macroinvertebrates, and quantified habitat complexity. After wood addition, total native fish densities were 2.2× higher in treatments compared with references, whereas total nonnative fish densities exhibited no response. Macroinvertebrate densities were 6.8× higher, and habitat complexity increased in treatments. Counts of geomorphic features in treatments increased from 1 to a maximum of 11 following wood addition, while the number of features in references remained unchanged. Wood addition has potential to instigate natural riverine processes, ultimately enhancing native fish habitat by increasing macroinvertebrate densities and habitat complexity in dryland rivers. Water overallocation and increasing aridity will continue to challenge efforts to improve habitat conditions with environmental flows alone, and managers might consider integrating non-flow alternatives like addition of abundant, invasive wood to reduce habitat simplification.

New Mexico, Utah

Effects of 2D hydrodynamic model resolution on habitat estimates for rearing Coho Salmon in contrasting channel forms

Estimating the impacts of water allocation decisions on fish populations and habitat availability is an important part of environmental flow assessments, especially in locations where water resources are limited. Two-dimensional hydrodynamic models (2DHMs) are commonly coupled with biological models to estimate fish habitat quality, area, and capacity across a range of proposed streamflows. Increasingly, resource managers are relying on landscape-scale model domains with coarse model resolutions to maintain feasible computational loads, but this may affect habitat estimates if the mesh element size of the model exceeds the spatial scale relevant to the organism. We investigated how coarsening the resolution of a 2DHM influences the area and spatial distribution of estimated Coho Salmon ( Oncorhynchus kisutch ) fry habitats. We used an interpolation scheme that upscaled mesh elements from a high-resolution (0.25 m 2 ) 2DHM to quantify and visualize the effects of 2DHM resolution on estimates of Coho Salmon fry habitat for two contrasting channel morphologies and across a broad range of streamflows. Estimates of Coho Salmon fry habitat at increasingly coarser resolutions led to 20%–50% reductions in weighted usable habitat area (WUA) across several streamflow scenarios for a complex channel type, but did not impact estimates in a confined, flume-like channel. Additionally, flow-to-habitat area relationships were not congruent at a given streamflow when resolution coarsened. Along with almost 500% more high-quality habitat area estimated in the complex channel type over the confined, discrepancies in habitat area increased with higher flows in areas defined as optimal for rearing Coho Salmon fry. Considering that complex channel types contain critical habitat for Coho Salmon fry, this study suggests coarse 2DHM resolutions may exclude important wetted edge and off-channel habitats from environmental flow assessments.

California