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Research about Washington

Source-linked reports with geographic coverage including Washington.

At least 289 records · Page 16Linked to original sources
USGS2020

Cryptic and extensive hybridization between ancient lineages of American crows

Most species and therefore most hybrid zones have historically been defined using phenotypic characters. However, both speciation and hybridization can occur with negligible morphological differentiation. Recently developed genomic tools provide the means to better understand cryptic speciation and hybridization. The Northwestern Crow ( Corvus caurinus ) and American Crow ( Corvus brachyrhynchos ) are continuously distributed sister taxa that lack reliable traditional characters for identification. In this first population genomic study of Northwestern and American crows, we use genomic SNPs (nuDNA) and mtDNA to investigate the degree of genetic differentiation between these crows and the extent to which they may hybridize. Our results indicate that American and Northwestern crows have distinct evolutionary histories, supported by two nuDNA ancestry clusters and two 1.1%‐divergent mtDNA clades dating to the late Pleistocene, when glacial advances may have isolated crow populations in separate refugia. We document extensive hybridization, with geographic overlap of mtDNA clades and admixture of nuDNA across >900 km of western Washington and western British Columbia. This broad hybrid zone consists of late‐generation hybrids and backcrosses, but not recent (e.g., F1) hybrids. Nuclear DNA and mtDNA clines had concordant widths and were both centred in southwestern British Columbia, farther north than previously postulated. Overall, our results suggest a history of reticulate evolution in American and Northwestern crows, perhaps due to recurring neutral expansion(s) from Pleistocene glacial refugia followed by lineage fusion(s). However, we do not rule out a contributing role for more recent potential drivers of hybridization, such as expansion into human‐modified habitats.

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USGS2020

Deglaciation of the Puget Lowland, Washington

Recently obtained radiocarbon ages from the southern Puget Lowland and reevaluation of limiting ages from the Olympic Peninsula in the light of new light detection and ranging (LiDAR) data suggest that the Juan de Fuca and Puget lobes of the Cordilleran ice sheet reached their maximum extents after 16,000 calibrated yr B.P. Source areas for both lobes fed through a common conduit, likely requiring that downstream responses to changes in either source area were similar. Dates for ice-sheet retreat are sparse and contradictory, but they suggest that retreat was rapid. Depositional and geomorphic evidence shows that retreat of the Juan de Fuca lobe predated retreat of the Puget lobe. No recessional end moraines have been identified in the Puget Lowland, in contrast to numerous recessional end moraines constructed by the Okanogan lobe east of the Cascade Range, and in contrast to later ice-sheet retreat in western Whatcom County north of the Puget Lowland. These observations lead to the hypothesis that collapse of the Juan de Fuca lobe, hastened by the instability of a marine-based ice sheet, steepened the ice-sheet surface over the eastern Strait of Juan de Fuca and diverted ice flow upstream of the Puget lobe to the west. Starved of ice, the Puget lobe retreated quickly.

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USGS2020

Between the supercontinents: Mesoproterozoic Deer Trail Group, an intermediate age unit between the Mesoproterozoic Belt–Purcell Supergroup and the Neoproterozoic Windermere Supergroup in northeastern Washington, USA

Mesoproterozoic and Neoproterozoic basins in western North America record the evolving position of the Laurentian craton within two supercontinents during their growth and dismemberment: Columbia (Nuna) and Rodinia. The western-most exposures of the Columbia rift-related Belt–Purcell Supergroup are preserved in northeastern Washington, structurally overlain by the Deer Trail Group and depositionally overlying the Neoproterozoic Windermere Supergroup. It has been disputed whether the Deer Trail Group is correlative with the Belt–Purcell Supergroup, or younger. To help resolve the uncertain correlation of these units and their bearing on supercontinent evolution, we characterized the detrital zircon age populations of units from the Deer Trail Group, the Windermere Supergroup, and the Belt–Purcell Supergroup in northeastern Washington. These data show that the western part of the Columbia supercontinent (now located in Australia and eastern Antarctica) remained attached to western Laurentia and continued to supply 1600–1500 Ma detrital zircon grains to the Belt–Purcell Supergroup until after ca. 1391 Ma. The Deer Trail Group is younger than the Belt–Purcell strata, with the basal unit younger than ca. 1362 Ma and a middle unit younger than ca. 1300 Ma. The Deer Trail Group has a pre-Grenville-age provenance from the southwestern USA and possibly east Antarctica. The Buffalo Hump Formation is younger than the Deer Trail Group, with Grenville-age (ca. 1112 Ma) detrital zircon grains and a detrital zircon signature like that of the overlying Neoproterozoic Windermere Supergroup. We interpret the Deer Trail Group to have been deposited during the rift-demise of supercontinent Columbia and before the Grenville-age assembly of the supercontinent Rodinia.

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USGS2020

Distribution and transport of Olympia oyster, Ostrea lurida, larvae in northern Puget Sound, Washington, USA

As efforts for restoring Olympia oyster (Ostrea lurida) populations have expanded, there is an increased need to understand local factors that could influence the long-term success of these projects. To address concerns over potential limitations to recruitment at a restoration site in northern Puget Sound, Washington, USA, a study was developed to characterize physical processes governing larval transport in conjunction with larval abundance and environmental factors. Larval presence was not associated with tide cycle, season, or a combination of tide cycle and season. In terms of location, larvae were more likely to be present at offshore and intertidal sites versus the estuarine lagoon where the adult population resides. Larval density was higher during late summer ebbs versus early summer floods. Across sampling dates and locations, larval sizes ranged from 184 to 263 µm, indicating that larvae were released into the water column throughout the reproductive season and retained in the embayment for at least ~16 days. Throughout different tidal cycles in Skagit Bay, acoustic Doppler current profilers were used to measure current direction and velocities, concurrent with plankton sampling. Surface currents in the study area alternated between a clockwise and counterclockwise gyre during initial ebb and flood tides, respectively. Larvae exported from the source population during initial to mid-ebbs are swept into a northward gyre, and potentially retained at intertidal sites alongshore. These results will provide resource managers attempting to restore native bivalves with the ability to expand populations by identifying optimal areas for habitat enhancement through natural recruitment.

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USGS2020

Variable prey consumption leads to distinct regional differences in Chinook salmon growth during the early marine critical period

Growth during the early marine critical period is positively associated with survival and recruitment for Pacific salmon Oncorhynchus spp., so it is important to understand how certain foraging strategies may bolster growth in estuarine and marine environments. To elucidate how spatiotemporal and demographic differences in diet contribute to growth rate variability, we analyzed stomach contents in tandem with morphometric and hormonal indices of growth for subyearling Chinook salmon O. tshawytscha captured in Puget Sound, Washington, USA. Regional dietary patterns indicated that fish caught in northern Puget Sound ate insects in the estuarine and nearshore habitats, followed by decapod larvae, euphausiids, or forage fish in the offshore zone. In southern Puget Sound, fish ate insects in the estuary but were more likely to eat mysids and other crustaceans in the nearshore zone. In the marine habitats adjacent to the San Juan Islands, subyearlings ate forage fish, and their stomachs were as much as 1.4 to 3 times fuller than salmon captured in other regions. Scale-derived growth rates and insulin-like growth factor-1 levels showed distinct growth advantages for San Juan Islands fish which were strongly associated with the early adoption of piscivory. However, consumption of larger crustaceans such as mysids and euphausiids was also associated with greater relative growth regardless of where individuals were captured. These findings highlight how spatiotemporal differences in prey quantity, prey profitability, and individual foraging strategies result in variable growth rates among salmon populations. Specifically, they emphasize the role of piscivory in promoting early marine growth for out-migrating Chinook salmon.

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USGS2020

Pre-late Wisconsin valley-glacier erratics between Leavenworth and Peshastin, Wenatchee valley, Washington

The late Wisconsin Icicle Creek alpine glacier transported tonalite boulders from the Mount Stuart batholith to arcuate end moraines in Icicle valley and Wenatchee valley near Leavenworth. Some previous workers considered sparsely weathered Mount Stuart boulders lying outside these moraines and draped by silt as having been ice rafted in a late Wisconsin lake. But the boulders are more likely within drift or end moraines of pre-late Wisconsin glaciations correlative to lateral moraines on Boundary Butte. The silt—a younger deposit that covers the boulders—derives not from a physically ponded lake but apparently from brief Missoula flood(s) backflooding up the valley.

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USGS2020

Canine distemper virus in the sea otter population (Enhydra lutris) in Washington State, USA

Before 2001, all serosurveys for morbilliviruses in sea otters ( Enhydra lutris ) in California, Washington, and Alaska, USA, documented a 0% seroprevalence. The first published serologic detections of morbillivirus in sea otters occurred in 2001–02 in live-captured Washington sea otters, with a documented 80% seroprevalence. We conducted a retrospective study of sea otter cases from 1989 to 2010 compiled at the US Geological Survey, National Wildlife Health Center to identify cases of morbilliviral disease in Washington sea otters and to characterize the disease using immunohistochemistry, reverse transcription (RT)-PCR, genetic sequencing, virus isolation, and serology. We identified six cases of morbilliviral disease and 12 cases of morbilliviral infection in this population of sea otters during 2000–10. Significant histologic findings included inflammation in the white and gray matter of the brain characterized by lymphoplasmacytic perivascular cuffing, neuronal necrosis, and satellitosis in gray matter and by spongiosis, myelin degeneration, spheroids, and gemistocytes in white matter. Intranuclear and intracytoplasmic viral inclusion bodies were found in neurons, Purkinje cells, and glia. Immunohistochemistry for canine distemper virus (CDV) showed positive staining in neurons, glial cells, and cell processes. A pan-morbillivirus RT-PCR with subsequent restriction endonuclease digestion or sequencing identified CDV. Virus isolation was not successful. Two sea otters with morbilliviral encephalitis showed greater antibody titers to CDV than phocine distemper virus. Histologic changes were confined to the central nervous system and resembled neurologic canine distemper in domestic dogs. Cases of sea otters with morbilliviral infection without histologic changes could represent early infections or incompletely cleared sublethal infections. We found that morbillivirus was present in the Washington sea otter population as early as 2000, and we provide a description of the pathology of canine distemper in sea otters.

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USGS2020

Wind River subbasin restoration: Annual report of US..Geological Survey activities, January 2018 through December 2018

We sampled juvenile wild Steelhead Oncorhynchus mykiss in headwater streams of the Wind River, WA, to characterize populations and investigate life-history metrics, particularly migratory patterns. We used Passive Integrated Transponder (PIT)-tagging and a series of instream PIT-tag interrogation systems (PTISs) to track juveniles. The Wind River subbasin is considered a wild Steelhead refuge by Washington Department of Fish and Wildlife (WDFW). No hatchery Steelhead have been planted in the Wind River subbasin since 1997, and hatchery adults are estimated to be less than one percent of spawners in most years (pers comm. Thomas Buehrens, Washington Department of Fish and Wildlife). Our repeated headwater sampling of consistent sites in the Wind River subbasin has also allowed us to track relative abundance of Brook Trout, a non-native species to the Wind River. Our work is contributing to understanding of Steelhead population response to numerous restoration actions in the subbasin, including removal of Hemlock Dam from Trout Creek in 2009, where our PTISs are helping to quantify adult response. Data from our study, and companion work by Washington Department of Fish and Wildlife, are contributing to Bonneville Power Administration’s (BPA) Research Monitoring and Evaluation (RM&E) Program Strategy of Fish Population Status Monitoring (www.cbfish.org/ProgramStrategy.mvc/ViewProgramStrategySummary/1). Specifically this work addresses the sub-strategies of: 1) Assessing the Status and Trends of Diversity of Natural Origin Fish Populations and to Uncertainties Research regarding differing life histories of a wild Steelhead population, 2) Assessing the Status and Trend of Adult Natural Origin Fish Populations, and 3) Monitoring and Evaluating the Effectiveness of Tributary Habitat Actions Relative to Environmental, Physical, or Biological Performance Objectives. Our headwaters parr PIT tagging, WDFW parr, smolt, and adult tagging and our instream PTISs are providing data on movements and life histories of parr, smolt, and adult Steelhead. During summer 2018, we PIT-tagged 1,592 age-0 and age-1 Steelhead parr in headwater areas of the Wind River subbasin to characterize population traits and investigate life-history diversity, including growth and pre-smolt downstream movement. Repeat headwater sampling and smolt trap operations provide opportunities for recapture, and instream PTISs and Columbia River infrastructure provide opportunity for detection of PIT-tagged fish. Throughout the year, we maintained a series of six instream PTISs to monitor movement of tagged Steelhead parr, smolts, and adults. Detections at the instream PTISs have demonstrated trends of age-0 and age-1 parr emigration from natal areas during summer and fall, in addition to the expected movement of parr and smolts in spring. Substantial numbers of parr make downstream movements as age-1 fish. We have estimated that from 15 to 33 percent of parr tagged as age-0 fish make downstream migrations at age-1 for additional rearing. We have estimated that from 1 to 27 percent of parr tagged as age-1 fish make downstream migrations during fall. These findings raise many questions about parr rearing strategies, habitat use, and success of these migrants and suggest a need for broader monitoring of juvenile Steelhead in some river systems to fully document juvenile production. Long-term monitoring of PIT-tagged fish is providing information on contribution of various life-history strategies to smolt production and adult returns. Movements of PIT-tagged adult Steelhead were recorded at instream PTISs. These data have allowed assessment of adult returns to tributary watersheds within the Wind River subbasin. Detection efficiency of adult PIT-tagged Steelhead at our primary adult-monitoring PTIS in Trout Creek has been greater than 92 percent during 6 of the past 7 years. This is providing excellent data to estimate adult returns to this watershed. Determination of adult use of tributary watersheds is providing data to help evaluate the efficacy of the removal of Hemlock Dam on Trout Creek. Hemlock Dam, located at rkm 2.0 of Trout Creek, was removed in summer 2009. The dam contributed to hydrologic impairment of Trout Creek and had potential negative effects on Steelhead. The improvements made to the upper Wind River PTIS (site code WRU at rkm 28.3; better site characteristics and grid power) during 2016 and 2017, and a planned new site in the Mine Reach of the upper Wind River, will allow estimates of subbasin adult escapement like those in Trout Creek. During 2018, we also completed planning and permitting with U.S. Forest Service for a new PTIS site at rkm 36 of the Wind River (the Mine Reach, mentioned above). This site will replace two sites (one in Paradise Creek and one at rkm 41 of the Wind River), which had operational challenges due to lack of adequate solar power and winter difficulties. The new Mine Reach PTIS site at rkm 36, will have better solar exposure, fewer winter operations difficulties, and provide opportunity to detect fish from juvenile sampling sites that were downstream of the previous two PTISs. The more consistent operation of the new Mine Reach PTIS site will increase our ability to estimate migrant abundance as all the juveniles tagged upstream of it will be subject to the same potential detection history, instead of three different potential detection histories as before. Additionally, with the new Mine Reach PTIS site lower in the watershed, it will subject more PIT-tagged adult Steelhead to detection and provide ability to generate a nonbiased adult-detection efficiency estimate for the WRU PTIS at rkm 28.3 of the Wind River. This will provide the opportunity to estimate yearly adult Steelhead abundance to the upper Wind watershed area. Permitting is complete and some supplies have been purchased to build and install this new site in 2019. Repeat sampling at consistent locations in the subbasin has allowed investigation into juvenile Steelhead growth patterns. Growth rates (relative change in weight) of age-0 PIT-tagged parr during summer are similar across the subbasin, but lower for age-1 parr in the Trout Creek watershed than the upper Wind River watershed. Yearly growth for parr tagged at age-0 is similar across the subbasin. Yearly growth for parr tagged at age-1 is lowest in Martha Creek, but similar elsewhere. Non-native Brook Trout are present in portions of the subbasin, chiefly the Trout Creek watershed, and repeat sampling has allowed us to index their prevalence. Percentage of catch that is Brook Trout at each of four sample sites in Trout Creek have declined from the period 1998 – 2003 to the period 2011 – 2018. There was a pattern of decline in percent of catch and number of Brook Trout at the Trout Creek sites from 2011 through 2016, though a slight upward trend during 2017 and 2018 has been evident. Evaluating and planning restoration efforts are of interest to many managers and agencies to ensure efficient use of resources. The evaluation of various life-histories of Steelhead within the Wind River subbasin will provide information to better track populations, and to direct habitat restoration and water allocation planning. Movement of Steelhead parr raises many questions regarding estimating juvenile abundance, origin, and habitat use within watersheds. Improved PTISs and focused PIT tagging of age-0 and age-1 Steelhead parr are increasingly allowing us to investigate such questions. Increasingly detailed Viable Salmonid Population information, such as that provided by PIT-tagging and instream PTISs networks like those in the Wind River subbasin, provide data to inform policy and management, as life-history strategies and production bottlenecks are identified and understood.

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USGS2020

Quality Assurance Project Plan: Status and trends monitoring of small streams in the Puget Lowlands ecoregion for Stormwater Action Monitoring (SAM)

This Quality Assurance Project Plan (QAPP) details a long term status and trends monitoring study for small streams in the Puget Lowland as part of Stormwater Action Monitoring (SAM) program. SAM is the regional stormwater monitoring program funded by the Phase I Municipal Stormwater permit and the Western Washington Phase II Municipal Stormwater permit permittees. This study of small streams in the Puget Lowland Ecoregion (called Puget Small Streams, PSS study, or SAM_PSS study hereafter) is designed to answer the question, “Are regional conditions in receiving water quality and biota improving in concert with broad implementation of required stormwater management practices?” In 2015, the first round of monitoring evaluated the condition (status) of streams (DeGasperi et al., 2018). Beginning in 2020 and thereafter this study will monitor streams’ changes over time in urban, urbanizing and rural areas of the Puget Lowland. The PSS will follow the protocols developed for the on-going statewide stream health monitoring program-Status and Trends Monitoring for Watershed Health and Salmon Recovery (WHSR) for physical habitat, biological measurements, except for minor changes to water quality parameters to better capture the stormwater-related chemistry signals. In addition this effort will sample sieved sediments for stormwater-related chemistry signals. This QAPP ensures quality data collection, analysis, reporting and management of the SAM PSS monitoring study

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USGS2020

Development of a new open-source tool to map burned area and burn severity

Accurate and complete geospatial fire occurrence records are important in determining postfire effects, emissions, hazards, and fuel loading inventories. Currently, the Monitoring Trends in Burn Severity (MTBS) project maps the fire perimeter and burn severity of all large fires on public lands. Although the MTBS project maps a large proportion of the fire acreage, it maps a smaller proportion of the actual number of fires in the United States, thereby creating a data gap. To fill this data gap, fire scientists at the U.S. Geological Survey (USGS) Earth Resources Observation and Science Center (EROS; Sioux Falls, South Dakota) proposed creating an open-source Fire Mapping Tool (FMT; available at https://mtbs.gov/qgis-fire-mapping-tool) as part of a two-phase National Aeronautics and Space Administration (NASA) Applied Fire Science Program grant. Phase II developed the FMT to map burn perimeters and severity not included in the MTBS database. This paper will focus on Phase II and will explain the algorithms that enhance the FMT’s functionality, demonstrate fire mapping procedures, and provide an example comparison between MTBS analyst fire products and those mapped using the FMT. The overall goal in the production of the FMT was to provide a freely available tool that can be used to map fires anywhere in the world.

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USGS2020

Living with wildfire in the Squilchuck Drainage - Chelan County, Washington: 2020 data report

Research on the social dimensions of wildfire provides opportunities to understand how communities and the people who reside in those communities interact with the threat of wildfire. Overall, three findings from this project were particularly noteworthy. First, household survey results indicate that residents in the Squilchuck Drainage, Chelan County, Washington have high expectations of response services in the event of a wildfire. Second, the survey data indicated Chelan County Fire District 1 (CCFD1) was the most frequently reported source of wildfire risk information and was characterized as a source of useful information. Finally, the project in the Squilchuck Drainage was an opportunity to examine how heterogeneous communities inhabit a contiguous biophysical location. Reported findings highlight the fact that even in relatively small geographic spaces, the social variation could have important implications for how a wildfire mitigation program attends to the varying social conditions within the communities that they serve.

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USGS2020

Conservation genetics of imperiled striped whipsnake in Washington

Conservation of wide-ranging species is aided by population genetic information that provides insights into adaptive potential, population size, interpopulation connectivity, and even extinction risk in portions of a species range. The Striped Whipsnake (Masticophis taeniatus) occurs across 11 western U.S. states and into Mexico but has experienced population declines in parts of its range, particularly in the state of Washington. We analyzed nuclear and mitochondrial DNA extracted from 192 shed skins, 63 muscle tissue samples, and one mouth swab to assess local genetic diversity and differentiation within and between the last known whipsnake populations in Washington. We then placed that information in a regional context to better understand levels of differentiation and diversity among whipsnake populations in the northwestern portion of the range of the species. Microsatellite data analyses indicated that there was comparable genetic diversity between the two extant Washington populations, but gene flow may be somewhat limited. We found moderate to high levels of genetic differentiation among states across all markers, including five microsatellites, two nuclear genes, and two mitochondrial genes. Pairwise state-level comparisons and dendrograms suggested that Washington whipsnakes are most closely related to those in Oregon, and distinct from Idaho, Nevada, and Utah, approximately following an isolation by distance model. We conclude that Washington populations of whipsnakes have experienced recent isolating events, but they have yet to lose genetic diversity. The longevity and high vagility of the species may provide opportunity for conservation of whipsnakes in the state as long as shrubland habitat is available

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USGS2020

Reserve network design for prairie-dependent taxa in South Puget Sound

Conserving species requires managing threats, including habitat loss. One approach to managing habitat loss is to identify and protect habitat in networks of reserves. Reserve network design is a type of resource allocation problem: how can we choose the most effective reserve network design given available resources? We undertook development and implementation of a patch dynamics model to allow us to evaluate proposed reserve networks in terms of ability to sustain populations of several taxa that are dependent on native prairie in the South Puget Sound region of Washington, USA. We used expert input to build a patch dynamics model for each taxon and used the model to examine probability of persistence in 50 years under a variety of reserve network designs, including the existing reserve network. Results suggest that the existing reserve network offers varying levels of protection for the different taxa, from desirable (>90% certain that the probability of persistence is ≥75% in 50 years) to negligible. We identified a reserve network that was >90% certain to protect all 6 taxa of interest, which would require a combination of land protection and translocations of taxa to new or existing reserves. Post hoc, we also identified possible hybrid alternatives, involving addition of new reserves and growth of existing reserves, that protected all 6 taxa without translocations. The approach we demonstrate is technically tractable and allows for the evaluation of any proposed reserve network design, thereby allowing a decision maker to evaluate a set of reserve networks that meet resource constraints and determine which of those best meets conservation objectives.

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USGS2019

Preliminary assessment of shallow groundwater chemistry near Goodell Creek, North Cascades National Park, Washington

Goodell Creek is located within North Cascades National Park and is high-quality habitat for Chinook salmon, which are listed as threatened under the Endangered Species Act. The creation of a levee near the mouth of the creek where it enters the Skagit River has cut off the historical flood plain from the active channel. There is an effort to remove the levee along the left bank of the creek to restore this connection and preserve this high-quality habitat; however, construction debris and blasting waste from the past have been used as fill during the creation of the levee in the 1980s, and there is concern that contaminated groundwater could reach the creek if the levee is breached. As a result, the U.S. Geological Survey, in cooperation with the National Park Service, assessed near-channel shallow-groundwater chemistry to determine how levee remediation should proceed. Groundwater was sampled in late summer in 2017. Deep groundwater at that time of year limited water-quality sampling. Six samples were collected in September 2017 and analyzed for nutrients, dissolved metals, and a suite of semi-volatile organic compounds. Laboratory data were almost always reported as below the method detection limits with the exception of data from a single shallow-groundwater well. Elevated concentrations of metals detected in water samples from this well were possibly caused by the buried metal in the vicinity of the sampling location. One sample collected close to the active channel of Goodell Creek showed no signs of contamination.

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USGS2019

Seasonal use of a nonnatal marine basin by juvenile hatchery chinook salmon

Information on the movement patterns of fishes is essential for managers that are making critical resource decisions. We examined the frequency of a keystone species, Chinook Salmon Oncorhynchus tshawytscha that migrated from different marine basins to the Nisqually River estuary, which lies within the southernmost marine basin (hereafter, “South basin”) in Puget Sound (Washington, USA). Hatchery‐reared juvenile fish were sampled by using beach seine, lampara seine, and fyke nets to determine seasonal trends in frequency, habitat use, and the influence of different capture methods. The captured fish originated from three marine basins, nine Puget Sound rivers, and fourteen hatcheries. The data revealed a consistent pattern showing that most of the tagged fish (72%) were from the nearby Nisqually River (in the South basin), but fish from more northerly marine basins (hereafter, “Outbasin”) were also common. Although the majority of the tagged fish (99%) that were captured during April and May were originally released into rivers adjacent to the South basin, 90% of the fish that were captured in August and September had originated from rivers adjacent to Outbasin locations (up to 130 km distant). A comparison of sampling methods showed that the beach seine produced 27% Outbasin fish compared with 53% that were obtained with the lampara seine. The analysis of habitat use suggested that during June and July, more Outbasin fish (>40%) were captured in delta flats and nearshore habitats than in estuarine emergent marsh habitat (26%). Release location (river basin), but not distance, appeared to be an important factor that influenced the percentage of Outbasin fish that were captured in the South basin. However, it appeared that the fish that were released at light weights and early dates were more likely to be captured. Information on the movement of juvenile salmon to a nonnatal marine basin may help to increase our understanding of features of life history and survival, and it has application elsewhere, as many marine species are artificially propagated, released in large numbers, and have the potential to use nonnatal habitats.

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