Geology topics

Research about Spain

Source-linked reports with geographic coverage including Spain.

At least 19 recordsLinked to original sources

USGS2026

In vitro assessment of human alveolar epithelial responses to acute combined exposure to volcanic ash and sulphur dioxide gas

Volcanic activity releases sulphur dioxide (SO₂) gas and fine particulate matter that may degrade air quality in populated areas, yet the effects of combined exposure remain insufficiently characterised. We examined the biological effects of acute combined exposure to volcanic ash and SO₂ in human alveolar epithelial cells (A549) maintained at the air-liquid interface. Cells were exposed to respirable ash from the 2021 Tajogaite eruption (La Palma; 100 µg/cm²) and SO₂ gas (nominal concentration 50 ppm; 1, 2, or 3 h), separately or sequentially. Cellular outcomes evaluated after exposure included cytotoxicity, pro-inflammatory and redox responses. Tajogaite ash alone had limited effects on cells, while exposure to SO 2 altered inflammatory responses (increasing gene transcription while reducing extracellular cytokine levels). Co-exposure responses were largely comparable to SO₂ alone, with no statistically significant evidence of synergistic effects under the conditions tested. These findings highlight the importance of considering gas-phase exposures in volcanic environments.

Canary Islands, La Palma, Tajogaite volcano↗

USGS2026

Characterization and modeling approach for planning restoration strategies in a complex basin affected by acid mine drainage

The management of acid mine drainage (AMD) impacted catchments, such as the Odiel River basin, in southwestern Spain, prioritizes reclamation to meet water resources needs. Assessing water composition across its watercourses is needed to identify major AMD contributors and potentially guide remediation efforts. An equilibrium-based mixing model was developed to simulate AMD pollutant load reductions and estimate the impact of selective restorations on water quality in the Odiel River, particularly at the planned Alcolea Reservoir near its outlet. Sampling under varying flow conditions (average vs. high flow) showed a reduction in acidity transport (from 54.0 to 42.5 ton/day), attributed to greater neutralization effects during high flows. Over 90 % of metal-fluxes originate alone from the Riotinto (73.6 %), Tharsis (14.5 %), and San Telmo (5.00 %) mining districts, among many other mines. While geochemical model estimates fit well with observed data (R 2 = 0.99), some deviations in non-conservative constituents (i.e. pH, Fe and Al) were observed (R 2 = 0.73–0.99), likely due to uncertainties in solubility constants and redox/Fe speciation. After evaluating model reliability, two reduction scenarios (50 % and 100 %) were applied to the three primary sources. Full removal of contamination could subtantially improve impounded reservoir water quality, with pH values of 4.93 and 7.64, and a net acidity between 8.75 and 4.63 mg/L eq. CaCO 3 in both average and high flows, respectively. Such differences may be related to flow regime effects on water quality. However, a 50 % reduction is insufficient to meet drinking or irrigation standards, highlighting the need for full and appropriate reclamation. The model offers a management tool for decision-making in the restoration of the Odiel River basin and could be transferable to similar AMD-affected basins worldwide.

Odiel River basin, southwestern Spain↗

USGS2025

Post-traumatic stress disorder in adult population of La Palma (Spain) after the 2021 Tajogaite eruption: Environmental and sociodemographic predictors

The adverse mental health effects of exposure to disasters have long been recognized, with earthquakes, floods, tsunamis, and volcanoes (among others) contributing to anxiety, depression, and post-traumatic stress disorder (PTSD). Concerning PTSD and volcanic eruptions specifically, most studies have highlighted risk factors related to demographic and social factors (e.g., gender, age, and education), mental health, and those associated with the experience of the volcanic eruption (e.g., having evacuated). Establishing the demographic scope of those more likely to suffer PTSD following volcanic eruptions as well as understanding the aggravating factors is key to best tailor precautionary measurements and psychological interventions. In this study, we supplement these analyses by investigating the role of environmental contaminants (specifically, exposure to tephra falls and degraded air quality) together with demographic, health, and hazard experiences in contributing to PTSD. We focus on data collected after the 2021 eruption of the Tajogaite volcano (Cumbre Vieja, La Palma, Spain). Our results show that in addition to sex, education, distance from the volcano, evacuation status, health issues related to the eruption, and mental health antecedents, exposure to volcanic gas but not tephra is associated with higher PTSD scores. Our findings establish a picture of the prevalence of PTSD symptoms in various demographics groups and aggravating factors following the 2021 eruption of the Tajogaite volcano. We further call for an extended investigation (and potentially, ultimately, a mechanistic explanation) of the role of air quality (as a potential synergic hazard) in shaping mental health following exposure to a primary hazard, such as a volcanic eruption.

Canary Islands, Island of La Palma↗

USGS2025

Triple-oxygen isotopic evidence of prolonged direct bioleaching of pyrite with O2

Sulfate is often touted as containing atmospheric oxygen whose isotopic signature can constrain redox, environmental conditions, and biological activity. Yet, the amount and isotopic fractionation associated with air-O 2 incorporation during sulfate formation is still debated, making its verification difficult. In this study, we identify a distinct, microbially dominated environment with the potential to preserve maximum signals of air-O 2 in sulfate. We report triple-oxygen isotope data for sulfate produced from pyrite oxidation in microbial and abiotic experiments, and from natural dissolved sulfate from the Rio Tinto, Spain, an acid mine drainage site. The oxygen isotope systematics of sulfate in these environments define a unique kinetic isotope effect associated with initial stage pyrite oxidation by Acidithiobacillus ferrooxidans that preserves >80 % oxygen from air-O 2 in sulfate. Unlike experiments, which evolve toward water-oxygen dominated sulfate on short time scales, Rio Tinto, Spain hosts a microbe rich environment with distinct geochemistry that maintains high O 2 -oxygen in sulfate. Therefore, in addition to containing isotopic records from water and air, sulfates can also contain a biosignature that is promising for understanding conditions on Mars and early Earth.

Earth and Planetary Science Letters↗

USGS2024

Remote sensing evapotranspiration in ensemble-based framework to enhance cascade routing and re-infiltration concept in integrated hydrological model applied to support decision making

Integrated hydrological models (IHMs) help characterize the complexity of surface–groundwater interactions. The cascade routing and re-infiltration (CRR) concept, recently applied to a MODFLOW 6 IHM, improved conceptualization and simulation of overland flow processes. The CRR controls the transfer of rejected infiltration and groundwater exfiltration from upslope areas to adjacent downslope areas where that water can be evaporated, re-infiltrated back to subsurface, or discharged to streams as direct runoff. The partitioning between these three components is controlled by uncertain parameters that must be estimated. Thus, by quantifying and reducing those uncertainties, next to uncertainties of the other model parameters (e.g. hydraulic and storage parameters), the reliability of the CRR is improved and the IHM is better suited for decision support modelling, the two key objectives of this work. To this end, the remotely sensed MODIS-ET product was incorporated into the calibration process for complementing traditional hydraulic head and streamflow observations. A total of approximately 150,000 observations guided the calibration of a 13-year MODFLOW 6 IHM simulation of the Sardon catchment (Spain) with daily stress periods. The model input uncertainty was represented by grid-cell-scale parameterization, yielding approximately 500,000 unknown input parameters to be conditioned. The calibration was carried out through an iterative ensemble smoother. Incorporating the MODIS-ET data improved the CRR implementation, and reduced uncertainties associated with other model parameters. Additionally, it significantly reduced the uncertainty associated with net recharge, a critical flux for water management that cannot be directly measured and rather is commonly estimated by IHM simulations.

Sardon catchment↗

USGS2023

Effects of feeding and habitat on resting metabolic rates of the Pacific walrus

Arctic marine mammals live in a rapidly changing environment due to the amplified effects of global warming. Pacific walruses ( Odobenus rosmarus divergens ) have responded to declines in Arctic sea-ice extent by increasingly hauling out on land farther from their benthic foraging habitat. Energy models can be useful for better understanding the potential implications of changes in behavior on body condition and reproduction but require behavior-specific metabolic rates. Here we measured the resting metabolic rates (RMR) of three captive, adult female Pacific walruses through breath-by-breath respirometry when fed and fasted resting out of water (sitting and lying down) and while fed resting in water. RMR in and out of water were positively related with pretrial energy intake when not fasted and 25% higher than RMR when walruses were fasted and out of water. Overall, RMR was higher than previously estimated for this species. Fasting RMR out of water was only 25% lower than subsurface swimming metabolic rates suggestive of relatively efficient swimming in adult females. Our results identify the importance of considering feeding status and species-specific differences in affecting metabolic costs. Further research is needed to better understand potential energetic costs of thermoregulation at temperatures experienced by wild walruses.

Valencia↗

USGS2021

Sentinel-2 and WorldView-3 atmospheric correction and signal normalization based on ground-truth spectroradiometric measurements

Remote sensing satellite Earth Observing Systems (EOS) provide a variety of products for monitoring Earth surface processes at varying spatial and spectral resolutions. Combining information from high and medium spatial resolution images is valuable for monitoring ground cover and vegetation status in cropland, grassland, forests, and other natural settings. However, coupling information from different EOS requires compensating for atmospheric and view angle effects before integrating comparable surface reflectance (SR) values. The objectives of this study were i) to assess how different atmospheric constituents affect the atmospheric correction results in Sentinel-2 and WorldView-3 imagery, ii) to establish a relationship with field spectra measurements, and iii) to develop an empirical approach to ensure that SR values extracted from different EOS can be normalized for use in monitoring vegetation and land cover status. We compared surface reflectance values derived from Sentinel-2 images corrected with Sen2Cor, MODTRAN or FLAASH atmospheric correction approaches for the visible-to-near infrared regions. Additionally, this information was compared to SR values extracted from WorldView-3 imagery acquired from the same dates and location (Central Spain) and corrected with MODTRAN and FLAASH approaches. Assessment of the atmospheric correction was conducted by comparing satellite image SR with ground-truth spectra acquired with a FieldSpec hand-held spectroradiometer. The results emphasized the importance of using common atmospheric parameters collected from ancillary data sources (i.e. MODIS Atmosphere & Land products) to ensure a reliable SR comparison. When compared to field-collected spectral data, SR from corrected Sentinel-2 push-broom imagery showed a reliable match (<4% difference in the visible bands and <0.52% difference in the near infrared bands). However, SR imagery from the pointable WorldView-3 instrument showed significant deviation, likely resulting from the effects of steep off-nadir acquisition angles (24.6° to 39.1°) combined with surface anisotropy. The magnitude and sign of the deviation in SR differed depending on the vegetation type, wavelength and sun-surface-sensor geometry. Therefore, it was necessary to account for angular effects to ensure reliable comparisons of imagery from the different EOS. In this study, an empirical angular correction approach was developed based on calibrating each WorldView-3 band against the ground-truth spectra. This correction allowed for the accurate signal normalization of WorldView-3 and Sentinel-2 imagery SR in the visible-to-near infrared regions.

ISPRS Journal of Photogrammetry and Remote Sensing↗

USGS2020

Quantitative paleoflood hydrology

This chapter reviews the paleohydrologic techniques and approaches used to reconstruct the magnitude and frequency of past floods using geological evidence. Quantitative paleoflood hydrology typically leads to two phases of analysis: (1) documentation and assessment of flood physical evidence (paleostage indicators), and (2) relating identified flood evidence to flood discharge, based on hydraulic calculations. Most paleoflood studies rely on stratigraphic sequences of fine-grained flood deposits found in slack-water and eddy environments in bedrock rivers to enable the estimates of paleodischarges for floods of past few centuries or millennia. Geochronology, commonly based on techniques such as optically stimulated luminescence (OSL) and radiocarbon, enable paleoflood age estimates. Such paleoflood discharge and age information can vastly improve flood frequency estimates, particularly for large and rare floods for which quantile estimates are typically poorly constrained by short historical records. The inclusion of such physical evidence of flooding into flood frequency assessments has been aided by new techniques of frequency analysis that can efficiently employ such data. Consequently, paleoflood analysis is supporting probability risk management of critical infrastructure such as nuclear facilities, dams, or bridges. Paleoflood studies also support understanding of the recurrence of geomorphically effective flows and assessment of non-stationarity in the frequency of large floods due to climate, land-use, or other environmental changes.

Llobregat River↗

USGS2019

Methane emissions from artificial waterbodies dominate the carbon footprint of irrigation: A study of transitions in the food-energy-water-climate nexus (Spain, 1900-2014)

Irrigation in the Mediterranean region has been used for millennia and has greatly expanded with industrialization. Irrigation is critical for climate change adaptation, but it is also an important source of greenhouse gas emissions. This study analyzes the carbon (C) footprint of irrigation in Spain, covering the complete historical process of mechanization. A 21-fold total, 6-fold area-based, and 4-fold product-based increase in the carbon footprint was observed during the 20th century, despite an increase in water use efficiency. CH 4 emissions from waterbodies, which had not previously been considered in the C footprint of irrigation systems, dominated the emission budget during most of the analyzed period. Technologies to save water and tap new water resources greatly increased energy and infrastructure demand, while improvements in power generation efficiency had a limited influence on irrigation emissions. Electricity production from irrigation dams may contribute to climate change mitigation, but the amount produced in relation to that consumed in irrigation has greatly declined. High uncertainty in CH 4 emission estimates from waterbodies stresses a need for more spatially resolved data and an improved empirical knowledge of the links between water quality, water level fluctuations, and emissions at the regional scale.

Environmental Science & Technology↗

USGS2019

The antiquity of the Sahara Desert: New evidence from the mineralogy and geochemistry of Pliocene paleosols on the Canary Islands, Spain

The Sahara is the largest warm desert in the world, but its age has been controversial, with estimates ranging from Miocene to Holocene. Mineralogical and geochemical data show that paleosols of Pliocene to mid-Pleistocene age on Fuerteventura and Gran Canaria in the Canary Islands have developed in part from inputs of dust from Africa. These paleosols contain quartz and mica, minerals that are abundant in African dust but are rare in the basaltic rocks that dominate the Canary Islands. Trace elements with minimal mobility, Sc, Cr, Hf, Th, and Ta as well as the rare earth elements, show that paleosols have compositions that are intermediate between those of local rocks and African-derived dust. Thus, results reported here and in a recently published study by others indicate that 9 paleosols record delivery of African dust to the Canary Islands between ~4.8–2.8 Ma, ~3.0–2.9 Ma, ~2.3–1.46 Ma, and ~0.4 Ma. A long-term paleosol record of African dust input agrees with deep-sea records off the coast of western Africa that imply increased dust fluxes to the eastern Atlantic Ocean at ~4.6 Ma. It is concluded that the Sahara Desert has been in existence as an arid-region dust source, at least intermittently, for much of the Pliocene and continuing into the Pleistocene.

Canary Islands↗

USGS2018

Analysis of the impact of African dust storms on the presence of enteric viruses in the atmosphere in Tenerife, Spain

Airborne viruses and their relation to dust storms, as a possible route for dispersion, have not been widely investigated. There are, however, studies that have described the airborne dispersal of pathogenic viruses and their potential impact on public and agronomical health. Atmospheric samples were collected in an urban area of Tenerife during 2009, 2010, 2012 and 2013 and screened for the presence of enteric viruses using PCR and sequencing. The potential relationship of viral data with African dust storms and other climatic variables (viz., the seasonality, origin of the air mass and PM levels) was analyzed. Enteroviruses and Rotaviruses were detected in 15.4% (20/130) and 36.9% (48/130) of the samples, respectively. No significant statistical relationships were observed with African dust storms or the origin of the air masses, although higher percentages of positives were obtained for dust storm days. Enterovirus detection was significantly linked to warmer seasons, and PM 2.5 levels showed an inverse correlation with a rotaviral presence. This is the first multi-year report to describe the presence of Enterovirus and Rotavirus genetic sequences in air samples collected in an outdoor urban environment. The data illustrates the need for source region sampling to determine links and the influence of the weather and climatic and regional wind patterns on long-range atmospheric dispersion of viruses in future research efforts.

Tenerife↗

USGS2016

Ascent ability of brown trout, Salmo trutta , and two Iberian cyprinids − Iberian barbel, Luciobarbus bocagei , and northern straight-mouth nase, Pseudochondrostoma duriense − in a vertical slot fishway

Passage performance of brown trout ( Salmo trutta ), Iberian barbel ( Luciobarbus bocagei ), and northern straight-mouth nase ( Pseudochondrostoma duriense) was investigated in a vertical slot fishway in the Porma River (Duero River basin, Spain) using PIT telemetry. We analysed the effects of different fishway discharges on motivation and passage success. Both cyprinid species ascended the fishway easily, performing better than the trout despite their theoretically weaker swimming performance. Fishway discharge affected fish motivation although it did not clearly influence passage success. Observed results can guide design and operation criteria of vertical slot fishways for native Iberian fish.

Castilla y León region↗

USGS2015

Sprinting performance of two Iberian fish: Luciobarbus bocagei and Pseudochondrostoma duriense in an open channel flume

This paper presents sprinting data from Iberian barbel ( Luciobarbus bocagei ) and northern straight-mouth nase ( Pseudochondrostoma duriense ), volitionally swimming against high velocity flows (1.5, 2.5 and 3 m s −1 ) in an open channel flume. Swimming endurance and speed greatly exceeded previously published observations with both species attaining swim speeds >20 body lengths s −1 . Flow velocity was the primary variable limiting the distance both species were able to traverse. Barbel swam greater distances than nase at higher flow velocities, with longer individuals attaining greater distances than smaller ones. The results challenge established fish passage guidelines, suggesting that in some cases these species are capable of passing much higher velocities than was previously believed.

Vadacondes↗

USGS2015

Uranium-series ages of fossil corals from Mallorca, Spain: The "Neotyrrhenian" high stand of the Mediterranean Sea revisited

The emergent marine deposits of the Mediterranean basin have been recognized as an important record of Quaternary sea level history for more than a century. Previous workers identified what have been interpreted to be two separate high stands of sea in the late Quaternary, namely the “Eutyrrhenian” (thought to be ~ 120 ka) and the “Neotyrrhenian” (thought to be either ~ 100 ka or ~ 80 ka). On Mallorca, Spain, both of these named deposits lie close to present sea level, implying paleo-sea levels slightly above present during both marine isotope stages (MIS) 5.5/5e and either 5.3/5c or 5.1/5a. If these interpretations are correct, they conflict, at least in part, with sea level records from far-field localities. We analyzed corals from the Neotyrrhenian beds on Mallorca, which gave U-series ages from ~ 126 ka to ~ 118 ka. These ages are consistent with previously published amino acid data that show that the Neotyrrhenian and Eutyrrhenian deposits are not significantly different in age. A fossil molluscan fauna from the Neotyrrhenian deposits on Mallorca has a warm-water paleozoogeographic aspect, with nine southward-ranging species and four extralimital southern species. When compared with sea surface temperatures obtained from planktonic foraminifera and alkenones from ODP core 977 in the nearby Alboran Sea, the only time period that shows comparable warmth is MIS 5.5/5e, consistent with the U-series ages of corals from the Neotyrrhenian deposits. We propose that the Neotyrrhenian deposits are a beachrock facies of the same age as the Eutyrrhenian deposits. This interpretation is consistent with the differences in physical sedimentology of the two deposits, explains the U-series and amino acid data indicating the same age, is consistent with the very slight elevation difference of the Neotyrrhenian and Eutyrrhenian beds, and explains the similar, though not identical paleozoogeographic aspects of their fossil faunas.

Mallorca↗

USGS2014

Surface-groundwater interactions in hard rocks in Sardon Catchment of western Spain: an integrated modeling approach

The structural and hydrological complexity of hard rock systems (HRSs) affects dynamics of surface–groundwater interactions. These complexities are not well described or understood by hydrogeologists because simplified analyses typically are used to study HRSs. A transient, integrated hydrologic model (IHM) GSFLOW (Groundwater and Surface water FLOW) was calibrated and post-audited using 18 years of daily groundwater head and stream discharge data to evaluate the surface–groundwater interactions in semi-arid, ∼80 km 2 granitic Sardon hilly catchment in Spain characterized by shallow water table conditions, relatively low storage, dense drainage networks and frequent, high intensity rainfall. The following hydrological observations for the Sardon Catchment, and more generally for HRSs were made: (i) significant bi-directional vertical flows occur between surface water and groundwater throughout the HRSs; (ii) relatively large groundwater recharge represents 16% of precipitation (P, 562 mm.y −1 ) and large groundwater exfiltration (∼11% of P) results in short groundwater flow paths due to a dense network of streams, low permeability and hilly topographic relief; deep, long groundwater flow paths constitute a smaller component of the water budget (∼1% of P); quite high groundwater evapotranspiration (∼5% of P and ∼7% of total evapotranspiration); low permeability and shallow soils are the main reasons for relatively large components of Hortonian flow and interflow (15% and 11% of P, respectively); (iii) the majority of drainage from the catchment leaves as surface water; (iv) declining 18 years trend (4.44 mm.y −1 ) of groundwater storage; and (v) large spatio-temporal variability of water fluxes. This IHM study of HRSs provides greater understanding of these relatively unknown hydrologic systems that are widespread throughout the world and are important for water resources in many regions.

Sardon Catchment↗

USGS2014

Uranium-series ages of corals, sea level history, and palaeozoogeography, Canary Islands, Spain: an exploratory study for two Quaternary interglacial periods

We present the first U-series ages of corals from emergent marine deposits on the Canary Islands. Deposits at + 20 m are 481 ± 39 ka, possibly correlative to marine isotope stage (or MIS) 11, while those at + 12 and + 8 m are 120.5 ± 0.8 ka and 130.2 ± 0.8 ka, respectively, correlative to MIS 5.5. The age, elevations, and uplift rates derived from MIS 5.5 deposits on the Canary Islands allow calculations of hypothetical palaeo-sea levels during the MIS 11 high sea stand. Estimates indicate that the MIS 11 high sea stand likely was at least + 9 m (relative to present sea level) and could have been as high as + 24 m. The most conservative estimates of palaeo-sea level during MIS 11 would require an ice mass loss equivalent to all of the modern Greenland and West Antarctic ice sheets; the more extreme estimates would require additional ice mass loss from the East Antarctic ice sheet. Extralimital southern species of mollusks, found in both MIS 11 and MIS 5.5 deposits on the Canary Islands, imply warmer-than-modern sea surface temperatures during at least a part of MIS 11 and much warmer sea surface temperatures during at least a part of MIS 5.5. Both MIS 11 and MIS 5.5 marine deposits on the Canary Islands contain extralimital northern species of mollusks as well, indicating cooler-than-present waters at times during these interglacial periods. We hypothesize that the co-occurrence of extralimital southern and northern species of marine invertebrates in the fossil record of the Canary Islands reflects its geographic location with respect to major synoptic-scale controls on climate and ocean currents. Previous interglacials may have been characterized by early, insolation-forced warming, along with northward migration of the intertropical convergence zone (ITCZ), accompanied by weakened trade winds and diminished upwelling. This allowed the arrival of extralimital southern taxa from the tropical Senegalese faunal province. During later parts of the MIS 11 and 5.5 interglacials, decreased insolation may have resulted in southward migration of the ITCZ, strengthened trade winds, and re-establishment of upwelling. Such conditions may have brought about not only local extinction of the Senegalese fauna, but allowed southward migration of the cooler-water Mediterranean fauna to the Canary Islands in the later parts of interglacials, a complex palaeoclimate record that is mirrored in the deep-sea core record.

Canary Islands↗

USGS2014

Demography of common toads after local extirpation of co-occurring midwife toads

Estimating demographic parameters like survival or recruitment provides insight into the state and trajectory of populations, but understanding the contexts influencing those parameters, including both biotic and abiotic factors, is particularly important for management and conservation. At a high elevation national park in Central Spain, common toads ( Bufo bufo ) are apparently taking advantage of the near-extirpation of the midwife toad ( Alytes obstetricans ), as colonization into new breeding ponds is evident. Within this scenario, we expected demographic parameters of common toad populations tobe affected favorably by the putative release from competition. However, we found the population growth rate was negative in 4 of 5 years at the long-standing population; survival probability at the long-standing population and newly-colonised breeding ponds was lower than reported for other toads living at high elevations and the probability of recruitment was inadequate to compensate for the survival rate in maintaining a positive trajectory for either of the breeding ponds. We assessed weather covariates and disease for their contribution to the context that may be limiting the common toad’s successful use of the niche vacated by the midwife toad.

Amphibia-Reptilia↗

USGS2013

Inferring the relative resilience of alternative states

Ecological systems may occur in alternative states that differ in ecological structures, functions and processes. Resilience is the measure of disturbance an ecological system can absorb before changing states. However, how the intrinsic structures and processes of systems that characterize their states affects their resilience remains unclear. We analyzed time series of phytoplankton communities at three sites in a floodplain in central Spain to assess the dominant frequencies or “temporal scales” in community dynamics and compared the patterns between a wet and a dry alternative state. The identified frequencies and cross-scale structures are expected to arise from positive feedbacks that are thought to reinforce processes in alternative states of ecological systems and regulate emergent phenomena such as resilience. Our analyses show a higher species richness and diversity but lower evenness in the dry state. Time series modeling revealed a decrease in the importance of short-term variability in the communities, suggesting that community dynamics slowed down in the dry relative to the wet state. The number of temporal scales at which community dynamics manifested, and the explanatory power of time series models, was lower in the dry state. The higher diversity, reduced number of temporal scales and the lower explanatory power of time series models suggest that species dynamics tended to be more stochastic in the dry state. From a resilience perspective our results highlight a paradox: increasing species richness may not necessarily enhance resilience. The loss of cross-scale structure (i.e. the lower number of temporal scales) in community dynamics across sites suggests that resilience erodes during drought. Phytoplankton communities in the dry state are therefore likely less resilient than in the wet state. Our case study demonstrates the potential of time series modeling to assess attributes that mediate resilience. The approach is useful for assessing resilience of alternative states across ecological and other complex systems.

Las Tablas de Daimiel floodplain wetland↗