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Research about Great Lakes

Source-linked reports with geographic coverage including Great Lakes.

At least 127 records · Page 7Linked to original sources

Factors related to northern goshawk landscape use in the western Great Lakes region

Northern Goshawks ( Accipiter gentilis ) are a species of special conservation concern in the western Great Lakes bioregion and elsewhere in North America, and exhibit landscape-scale spatial use patterns. However, little information exists about Northern Goshawk habitat relations at broad spatial extents, as most existing published information comes from a few locations of relatively small spatial extent and, in some cases, short durations. We used an information-theoretic approach to evaluate competing hypotheses regarding factors (forest canopy cover, successional stage, and heights of the canopy top and base) related to odds of Northern Goshawk landscape use throughout the western Great Lakes bioregion based on an occupancy survey completed in 2008 ( Bruggeman et al. 2011 ). We also combined these data with historical data of Northern Goshawk nest locations in the bioregion from 1979–2006 to evaluate the same competing hypotheses to elucidate long-term trends in use. The odds of Northern Goshawk use in 2008, and from 1979–2008, were positively correlated with average percent canopy cover. In the best-approximating models developed using 1979–2008 data, the odds of landscape use were positively correlated with the percentages of the landscape having canopy heights between 10 m and 25 m, and 25 m and 50 m, and the amount of variability in canopy base height. Also, the odds of landscape use were negatively correlated with the average height at the canopy base. Our results suggest multiple habitat factors were related to Northern Goshawk landscape-scale habitat use, similar to habitat use described at smaller spatial scales in the western Great Lakes bioregion and in western North America and Europe.

Michigan, Minnesota, Wisconsin↗

Assessment of suitable habitat for Phragmites australis (common reed) in the Great Lakes coastal zone

In the Laurentian Great Lakes, the invasive form of Phragmites australis (common reed) poses a threat to highly productive coastal wetlands and shorelines by forming impenetrable stands that outcompete native plants. Large, dominant stands can derail efforts to restore wetland ecosystems degraded by other stressors. To be proactive, landscape-level management of Phragmites requires information on the current spatial distribution of the species and a characterization of areas suitable for future colonization. Using a recent basin-scale map of this invasive plant’s distribution in the U.S. coastal zone of the Great Lakes, environmental data (e.g., soils, nutrients, disturbance, climate, topography), and climate predictions, we performed analyses of current and predicted suitable coastal habitat using boosted regression trees, a type of species distribution modeling. We also investigated differential influences of environmental variables in the upper lakes (Lakes Superior, Michigan, and Huron) and lower lakes (Lakes St. Clair, Erie, and Ontario). Basin-wide results showed that the coastal areas most vulnerable to Phragmites expansion were in close proximity to developed lands and had minimal topographic relief, poorly drained soils, and dense road networks. Elevated nutrients and proximity to agriculture also influenced the distribution of Phragmites . Climate predictions indicated an increase in suitable habitat in coastal Lakes Huron and Michigan in particular. The results of this study, combined with a publicly available online decision support tool, will enable resource managers and restoration practitioners to target and prioritize Phragmites control efforts in the Great Lakes coastal zone.

Great Lakes↗

Geographic setting influences Great Lakes beach microbiological water quality

Understanding of factors that influence Escherichia coli (EC) and enterococci (ENT) concentrations, pathogen occurrence, and microbial sources at Great Lakes beaches comes largely from individual beach studies. Using 12 representative beaches, we tested enrichment cultures from 273 beach water and 22 tributary samples for EC, ENT, and genes indicating the bacterial pathogens Shiga-toxin producing E. coli (STEC), Shigella spp., Salmonella spp, Campylobacter jejuni/coli, and methicillin-resistant Staphylococcus aureus, and 108–145 samples for Bacteroides human, ruminant, and gull source-marker genes. EC/ENT temporal patterns, general Bacteroides concentration, and pathogen types and occurrence were regionally consistent (up to 40 km), but beach catchment variables (drains/creeks, impervious surface, urban land cover) influenced exceedances of EC/ENT standards and detections of Salmonella and STEC. Pathogen detections were more numerous when the EC/ENT Beach Action Value (but not when the Geometric Mean and Statistical Threshold Value) was exceeded. EC, ENT, and pathogens were not necessarily influenced by the same variables. Multiple Bacteroides sources, varying by date, occurred at every beach. Study of multiple beaches in different geographic settings provided new insights on the contrasting influences of regional and local variables, and a broader-scale perspective, on significance of EC/ENT exceedances, bacterial sources, and pathogen occurrence.

Great Lakes↗

Rivermouth alteration of agricultural impacts on consumer tissue δ 15 N

Terrestrial agricultural activities strongly influence riverine nitrogen (N) dynamics, which is reflected in the δ 15 N of riverine consumer tissues. However, processes within aquatic ecosystems also influence consumer tissue δ 15 N. As aquatic processes become more important terrestrial inputs may become a weaker predictor of consumer tissue δ 15 N. In a previous study, this terrestrial-consumer tissue δ 15 N connection was very strong at river sites, but was disrupted by processes occurring in rivermouths (the ‘rivermouth effect’). This suggested that watershed indicators of N loading might be accurate in riverine settings, but could be inaccurate when considering N loading to the nearshore of large lakes and oceans. In this study, the rivermouth effect was examined on twenty-five sites spread across the Laurentian Great Lakes. Relationships between agriculture and consumer tissue δ 15 N occurred in both upstream rivers and at the outlets where rivermouths connect to the nearshore zone, but agriculture explained less variation and had a weaker effect at the outlet. These results suggest that rivermouths may sometimes be significant sources or sinks of N, which would cause N loading estimates to the nearshore zone that are typically made at discharge gages further upstream to be inaccurate. Identifying definitively the controls over the rivermouth effect on N loading (and other nutrients) will require integration of biogeochemical and hydrologic models.

Great Lakes↗

Genetic population structure of muskellunge in the Great Lakes

We quantified genetic relationships among Muskellunge Esox masquinongy from 15 locations in the Great Lakes to determine the extent and distribution of measurable population structure and to identify appropriate spatial scales for fishery management and genetic conservation. We hypothesized that Muskellunge from each area represented genetically distinct populations, which would be evident from analyses of genotype data. A total of 691 Muskellunge were sampled ( n = 10–127/site) and genetic data were collected at 13 microsatellite loci. Results from a suite of analyses (including pairwise genetic differentiation, Bayesian admixture prediction, analysis of molecular variance, and tests of isolation by distance) indicated the presence of nine distinct genetic groups, including two that were approximately 50 km apart. Geographic proximity and low habitat complexity seemed to facilitate genetic similarity among areas, whereas Muskellunge from areas of greater habitat heterogeneity exhibited high differentiation. Muskellunge from most areas contained private alleles, and mean within-area genetic variation was similar to that reported for other freshwater fishes. Management programs aimed at conserving the broader diversity and long-term sustainability of Muskellunge could benefit by considering the genetically distinct groups as independent fisheries, and individual spawning and nursery habitats could subsequently be protected to conserve the evolutionary potential of Muskellunge.

Great Lakes↗

Association of toxin-producing Clostridium botulinum with the macroalga Cladophora in the Great Lakes

Avian botulism, a paralytic disease of birds, often occurs on a yearly cycle and is increasingly becoming more common in the Great Lakes. Outbreaks are caused by bird ingestion of neurotoxins produced by Clostridium botulinum, a spore-forming, gram-positive, anaerobe. The nuisance, macrophytic, green alga Cladophora (Chlorophyta; mostly Cladophora glomerata L.) is a potential habitat for the growth of C. botulinum. A high incidence of botulism in shoreline birds at Sleeping Bear Dunes National Lakeshore (SLBE) in Lake Michigan coincides with increasingly massive accumulations of Cladophora in nearshore waters. In this study, free-floating algal mats were collected from SLBE and other shorelines of the Great Lakes between June and October 2011. The abundance of C. botulinum in algal mats was quantified and the type of botulism neurotoxin (bont) genes associated with this organism were determined by using most-probable-number PCR (MPN-PCR) and five distinct bont gene-specific primers (A, B, C, E, and F). The MPN-PCR results showed that 16 of 22 (73%) algal mats from the SLBE and 23 of 31(74%) algal mats from other shorelines of the Great Lakes contained the bont type E (bont/E) gene. C. botulinum was present up to 15 000 MPN per gram dried algae based on gene copies of bont/E. In addition, genes for bont/A and bont/B, which are commonly associated with human diseases, were detected in a few algal samples. Moreover, C. botulinum was present as vegetative cells rather than as dormant spores in Cladophora mats. Mouse toxin assays done using supernatants from enrichment of Cladophora containing high densities of C. botulinum (>1000 MPN/g dried algae) showed that Cladophora-borne C. botulinum were toxin-producing species (BoNT/E). Our results indicate that Cladophora provides a habitat for C. botulinum, warranting additional studies to better understand the relationship between this bacterium and the alga, and how this interaction potentially contributes to botulism outbreaks in birds.

Great Lakes↗

Ecotoxicology of organochlorine chemicals in birds of the Great Lakes

Silent Spring was fulfilled in the United States with passage of environmental legislation such as the Clean Water Act, the Federal Insecticide, Fungicide, and Rodenticide Act, and the Toxic Substance Control Act in the 1970s. Carson's writings, television interviews, and testimony before Congress alerted a nation and the world to the unintended effects of persistent, bioaccumulative chemicals on populations of fish, wildlife, and possibly humans. Her writings in the popular press brought attention to scientific findings that declines in populations of a variety of birds were directly linked to the widespread use of dichlorodiphenyltrichloroethane (DDT) in agriculture, public health, and horticulture. By the 1970s, DDT and other persistent organic pollutants (POPs) were being banned or phased out, and the intent of these regulatory acts became apparent in a number of locations across the United States, including the Great Lakes. Concentrations of DDT and its major product of transformation, dichlorodiphenylchloroethane (DDE), were decreasing in top predators, such as bald eagles (Haliaeetus leucocephalus), osprey (Pandion haliaetus), colonial waterbirds, and other fish-eating wildlife. Eggshell thinning and the associated mortality of bird embryos caused by DDE had decreased in the Great Lakes and elsewhere by the early 1980s.

Environmental Toxicology and Chemistry↗

Comparative susceptibility among three stocks of yellow perch, Perca flavescens (Mitchill), to viral haemorrhagic septicaemia virus strain IVb from the Great Lakes

The Great Lakes strain of viral haemorrhagic septicaemia virus IVb (VHSV-IVb) is capable of infecting a wide number of naive species and has been associated with large fish kills in the Midwestern United States since its discovery in 2005. The yellow perch, Perca flavescens (Mitchill), a freshwater species commonly found throughout inland waters of the United States and prized for its high value in sport and commercial fisheries, is a species documented in several fish kills affiliated with VHS. In the present study, differences in survival after infection with VHSV IVb were observed among juvenile fish from three yellow perch broodstocks that were originally derived from distinct wild populations, suggesting innate differences in susceptibility due to genetic variance. While all three stocks were susceptible upon waterborne exposure to VHS virus infection, fish derived from the Midwest (Lake Winnebago, WI) showed significantly lower cumulative % survival compared with two perch stocks derived from the East Coast (Perquimans River, NC and Choptank River, MD) of the United States. However, despite differences in apparent susceptibility, clinical signs did not vary between stocks and included moderate-to-severe haemorrhages at the pelvic and pectoral fin bases and exophthalmia. After the 28-day challenge was complete, VHS virus was analysed in subsets of whole fish that had either survived or succumbed to the infection using both plaque assay and quantitative PCR methodologies. A direct correlation was identified between the two methods, suggesting the potential for both methods to be used to detect virus in a research setting.

Great Lakes↗

Distribution and abundance of freshwater polychaetes, Manayunkia speciosa (Polychaeta), in the Great Lakes with a 70-year case history for western Lake Erie

Manayunkia speciosa has been a taxonomic curiosity for 150 years with little interest until 1977 when it was identified as an intermediate host of a fish parasite (Ceratomyxa shasta) responsible for fish mortalities (e.g., chinook salmon). Manayunkia was first reported in the Great Lakes in 1929. Since its discovery, the taxon has been reported in 50% (20 of 40 studies) of benthos studies published between 1960 and 2007. When found, Manayunkia comprised < 1% of benthos in 70% of examined studies. In one extensive study, Manayunkia occurred in only 26% of 378 sampled events (1991–2009). The taxon was found at higher densities in one area of Lake Erie (mean = 3658/m 2 ) and Georgian Bay (1790/m 2 ) than in five other areas (mean = 60 to 553/m 2 ) of the lakes. A 70-year history of Manayunkia in western Lake Erie indicates it was not found in 1930, was most abundant in 1961 (mean = 8039, maximum = 67,748/m 2 ), and decreased in successive periods of 1982 (3529, 49,639/m 2 ), 1993 (1876, 25,332/m 2 ), and 2003 (79, 2583/m 2 ). It occurred at 48% of stations in 1961, 58% in 1982, 52% in 1993, and 6% of stations in 2003. In all years, Manayunkia was distributed primarily near the mouth of the Detroit River. Causes for declines in distribution and abundance are unknown, but may be related to pollution-abatement programs that began in the 1970s, and invasion of dreissenid mussels in the late-1980s which contributed to de-eutrophication of western Lake Erie. At present, importance of the long-term decline of Manayunkia in Lake Erie is unknown.

Great Lakes↗

Great Lakes rivermouth ecosystems: scientific synthesis and management implications

At the interface of the Great Lakes and their tributary rivers lies the rivermouths, a class of aquatic ecosystem where lake and lotic processes mix and distinct features emerge. Many rivermouths are the focal point of both human interaction with the Great Lakes and human impacts to the lakes; many cities, ports, and beaches are located in rivermouth ecosystems, and these human pressures often degrade key ecological functions that rivermouths provide. Despite their ecological uniqueness and apparent economic importance, there has been relatively little research on these ecosystems as a class relative to studies on upstream rivers or the open-lake waters. Here we present a synthesis of current knowledge about ecosystem structure and function in Great Lakes rivermouths based on studies in both Laurentian rivermouths, coastal wetlands, and marine estuarine systems. A conceptual model is presented that establishes a common semantic framework for discussing the characteristic spatial features of rivermouths. This model then is used to conceptually link ecosystem structure and function to ecological services provided by rivermouths. This synthesis helps identify the critical gaps in understanding rivermouth ecology. Specifically, additional information is needed on how rivermouths collectively influence the Great Lakes ecosystem, how human alterations influence rivermouth functions, and how ecosystem services provided by rivermouths can be managed to benefit the surrounding socioeconomic networks.

Great Lakes↗

Sensitivity of fish density estimates to standard analytical procedures applied to Great Lakes hydroacoustic data

Standardized methods of data collection and analysis ensure quality and facilitate comparisons among systems. We evaluated the importance of three recommendations from the Standard Operating Procedure for hydroacoustics in the Laurentian Great Lakes (GLSOP) on density estimates of target species: noise subtraction; setting volume backscattering strength (S v ) thresholds from user-defined minimum target strength (TS) of interest (TS-based S v threshold); and calculations of an index for multiple targets (N v index) to identify and remove biased TS values. Eliminating noise had the predictable effect of decreasing density estimates in most lakes. Using the TS-based Sv threshold decreased fish densities in the middle and lower layers in the deepest lakes with abundant invertebrates (e.g., Mysis diluviana). Correcting for biased in situ TS increased measured density up to 86% in the shallower lakes, which had the highest fish densities. The current recommendations by the GLSOP significantly influence acoustic density estimates, but the degree of importance is lake dependent. Applying GLSOP recommendations, whether in the Laurentian Great Lakes or elsewhere, will improve our ability to compare results among lakes. We recommend further development of standards, including minimum TS and analytical cell size, for reducing the effect of biased in situ TS on density estimates.

Great Lakes↗

A spatial age-structured model for describing sea lamprey ( Petromyzon marinus ) population dynamics

The control of invasive sea lampreys ( Petromyzon marinus ) presents large scale management challenges in the Laurentian Great Lakes. No modeling approach has been developed that describes spatial dynamics of lamprey populations. We developed and validated a spatial and age-structured model and applied it to a sea lamprey population in a large river in the Great Lakes basin. We considered 75 discrete spatial areas, included a stock-recruitment function, spatial recruitment patterns, natural mortality, chemical treatment mortality, and larval metamorphosis. Recruitment was variable, and an upstream shift in recruitment location was observed over time. From 1993–2011 recruitment, larval abundance, and the abundance of metamorphosing individuals decreased by 80, 84, and 86%, respectively. The model successfully identified areas of high larval abundance and showed that areas of low larval density contribute significantly to the population. Estimated treatment mortality was less than expected but had a large population-level impact. The results and general approach of this work have applications for sea lamprey control throughout the Great Lakes and for the restoration and conservation of native lamprey species globally.

Great Lakes↗

A synthesized mating pheromone component increases adult sea lamprey ( Petromyzon marinus ) trap capture in management scenarios

Application of chemical cues to manipulate adult sea lamprey (Petromyzon marinus) behavior is among the options considered for new sea lamprey control techniques in the Laurentian Great Lakes. A male mating pheromone component, 7a,12a,24-trihydroxy-3-one-5a-cholan-24-sulfate (3kPZS), lures ovulated female sea lamprey upstream into baited traps in experimental contexts with no odorant competition. A critical knowledge gap is whether this single pheromone component influences adult sea lamprey behavior in management contexts containing free-ranging sea lampreys. A solution of 3kPZS to reach a final in-stream concentration of 10 -12 mol·L -1 was applied to eight Michigan streams at existing sea lamprey traps over 3 years, and catch rates were compared between paired 3kPZS-baited and unbaited traps. 3kPZS-baited traps captured significantly more sexually immature and mature sea lampreys, and overall yearly trapping efficiency within a stream averaged 10% higher during years when 3kPZS was applied. Video analysis of a trap funnel showed that the likelihood of sea lamprey trap entry after trap encounter was higher when the trap was 3kPZS baited. Our approach serves as a model for the development of similar control tools for sea lamprey and other aquatic invaders.

Great Lakes↗

Hydraulic and water-quality data collection for the investigation of Great Lakes tributaries for Asian carp spawning and egg-transport suitability

If the invasive Asian carps (bighead carp Hypophthalmichthys nobilis and silver carp Hypophthalmichthys molitrix ) migrate to the Great Lakes, in spite of the efforts to stop their advancement, these species will require the fast-flowing water of the Great Lakes tributaries for spawning and recruitment in order to establish a growing population. Two Lake Michigan tributaries (the Milwaukee and St. Joseph Rivers) and two Lake Erie tributaries (the Maumee and Sandusky Rivers) were investigated to determine if these tributaries possess the hydraulic and water-quality characteristics to allow successful spawning of Asian carps. To examine this issue, standard U.S. Geological Survey sampling protocols and instrumentation for discharge and water-quality measurements were used, together with differential global positioning system data for georeferencing. Non-standard data-processing techniques, combined with detailed laboratory analysis of Asian carp egg characteristics, allowed an assessment of the transport capabilities of each of these four tributaries. This assessment is based solely on analysis of observed data and did not utilize the collected data for detailed transport modeling. All four tributaries exhibited potential settling zones for Asian carp eggs both within the estuaries and river mouths and within the lower 100 kilometers (km) of the river. Dams played a leading role in defining these settling zones, with the exception of dams on the Sandusky River. The impoundments created by many of the larger dams on these rivers acted to sufficiently decelerate the flows and allowed the shear velocity to drop below the settling velocity for Asian carp eggs, which would allow the eggs to fall out of suspension and settle on the bottom where it is thought the eggs would perish. While three rivers exhibited these settling zones upstream of the larger dams, not all settling zones are likely to have such effects on egg transport. The Milwaukee River exhibited only a short settling zone upstream of the Grafton Dam, whereas the St. Joseph and Maumee Rivers both had extensive settling zones (>5 km) behind major dams. These longer settling zones are likely to capture more eggs than shorter settling reaches. All four rivers exhibited settling zones at their river mouths, with the Lake Erie tributaries having much larger settling zones extending more than 10 km up the tributaries. While hydraulic data from all four rivers indicated settling of eggs is possible in some locations, all four rivers also exhibited sufficient temperatures, water-quality characteristics, turbulence, and transport times outside of settling zones for successful suspension and development of Asian carp eggs to the hatching stage before the threat of settlement. These observed data indicate that these four Great Lakes tributaries have sufficient hydraulic and water-quality characteristics to support successful spawning and recruitment of Asian carps. The data indicate that with the right temperature and flow conditions, river reaches as short as 25 km may allow Asian carp eggs sufficient time to develop to hatching. Additionally, examining the relation between critical shear velocity and mean velocity, egg settling appears to take place at mean velocities in the range of 15&ndash;25 centimeters per second, a much lower value than is generally cited in the literature. A first-order estimate of the minimum transport velocity for Asian carp eggs in a river can be obtained by using mean flow depth and river substrate data, and curves were constructed to show this relation. These findings would expand the number of possible tributaries suitable for Asian carp spawning and contribute to the understanding of how hydraulic and water-quality information can be used to screen additional rivers in the future.

Great Lakes↗

Developing and implementing the use of predictive models for estimating water quality at Great Lakes beaches

Predictive models have been used at beaches to improve the timeliness and accuracy of recreational water-quality assessments over the most common current approach to water-quality monitoring, which relies on culturing fecal-indicator bacteria such as Escherichia coli (E. coli.) . Beach-specific predictive models use environmental and water-quality variables that are easily and quickly measured as surrogates to estimate concentrations of fecal-indicator bacteria or to provide the probability that a State recreational water-quality standard will be exceeded. When predictive models are used for beach closure or advisory decisions, they are referred to as “nowcasts.” During the recreational seasons of 2010-12, the U.S. Geological Survey (USGS), in cooperation with 23 local and State agencies, worked to improve existing nowcasts at 4 beaches, validate predictive models at another 38 beaches, and collect data for predictive-model development at 7 beaches throughout the Great Lakes. This report summarizes efforts to collect data and develop predictive models by multiple agencies and to compile existing information on the beaches and beach-monitoring programs into one comprehensive report. Local agencies measured E. coli concentrations and variables expected to affect E. coli concentrations such as wave height, turbidity, water temperature, and numbers of birds at the time of sampling. In addition to these field measurements, equipment was installed by the USGS or local agencies at or near several beaches to collect water-quality and metrological measurements in near real time, including nearshore buoys, weather stations, and tributary staff gages and monitors. The USGS worked with local agencies to retrieve data from existing sources either manually or by use of tools designed specifically to compile and process data for predictive-model development. Predictive models were developed by use of linear regression and (or) partial least squares techniques for 42 beaches that had at least 2 years of data (2010-11 and sometimes earlier) and for 1 beach that had 1 year of data. For most models, software designed for model development by the U.S. Environmental Protection Agency (Virtual Beach) was used. The selected model for each beach was based on a combination of explanatory variables including, most commonly, turbidity, day of the year, change in lake level over 24 hours, wave height, wind direction and speed, and antecedent rainfall for various time periods. Forty-two predictive models were validated against data collected during an independent year (2012) and compared to the current method for assessing recreational water quality-using the previous day’s E. coli concentration (persistence model). Goals for good predictive-model performance were responses that were at least 5 percent greater than the persistence model and overall correct responses greater than or equal to 80 percent, sensitivities (percentage of exceedances of the bathing-water standard that were correctly predicted by the model) greater than or equal to 50 percent, and specificities (percentage of nonexceedances correctly predicted by the model) greater than or equal to 85 percent. Out of 42 predictive models, 24 models yielded over-all correct responses that were at least 5 percent greater than the use of the persistence model. Predictive-model responses met the performance goals more often than the persistence-model responses in terms of overall correctness (28 versus 17 models, respectively), sensitivity (17 versus 4 models), and specificity (34 versus 25 models). Gaining knowledge of each beach and the factors that affect E. coli concentrations is important for developing good predictive models. Collection of additional years of data with a wide range of environmental conditions may also help to improve future model performance. The USGS will continue to work with local agencies in 2013 and beyond to develop and validate predictive models at beaches and improve existing nowcasts, restructuring monitoring activities to accommodate future uncertainties in funding and resources.

Great Lakes↗

Great Lakes rivermouths: a primer for managers

Between the North American Great Lakes and their tributaries are the places where the confluence of river and lake waters creates a distinct ecosystem: the rivermouth ecosystem. Human development has often centered around these rivermouths, in part, because they provide a rich array of ecosystem services. Not surprisingly, centuries of intense human activity have led to substantial pressures on, and alterations to, these ecosystems, often diminishing or degrading their ecological functions and associated ecological services. Many Great Lakes rivermouths are the focus of intense restoration efforts. For example, 36 of the active Great Lakes Areas of Concern (AOCs) are rivermouths or areas that include one or more rivermouths. Historically, research of rivermouth ecosystems has been piecemeal, focused on the Great Lakes proper or on the upper reaches of tributaries, with little direct study of the rivermouth itself. Researchers have been divided among disciplines, agencies and institutions; and they often work independently and use disparate venues to communicate their work. Management has also been fragmented with a focus on smaller, localized, sub-habitat units and socio-political or economic elements, rather than system-level consideration. This Primer presents the case for a more holistic approach to rivermouth science and management that can enable restoration of ecosystem services with multiple benefits to humans and the Great Lakes ecosystem. A conceptual model is presented with supporting text that describes the structures and processes common to all rivermouths, substantiating the case for treating these ecosystems as an identifiable class. 1 Ecological services provided by rivermouths and changes in how humans value those services over time are illustrated through case studies of two Great Lakes rivermouths—the St. Louis River and the Maumee River. Specific ecosystem services are identified in italics throughout this Primer and follow definitions described by the Millennium Ecosystem Assessment (Table1). Collectively, this primer synthesizes existing information in a new way that aims to support management of rivermouths as distinct and important ecosystems. The development and management decisions made around rivermouths today will shape the future of these ecosystems, and the human communities within them, well into the future. 1 The information presented in this paper was derived from discussions and draft documents of the Great Lakes Rivermouth Collaboratory. The Great Lakes Rivermouth Collaboratory was established by the U.S. Geological Survey's Great Lakes Science Center (USGS-GLSC) in collaboration with the Great Lakes Commission to engage the Great Lakes scientific community in sharing and documenting knowledge about freshwater rivermouth ecosystems. For more information, see http://www.glc.org/habitat/Rivermouth-Collaboratory.html.

Great Lakes↗

Limitations of quantitative analysis of deep crustal seismic reflection data: Examples from GLIMPCE

Amplitude preservation in seismic reflection data can be obtained by a relative true amplitude (RTA) processing technique in which the relative strength of reflection amplitudes is preserved vertically as well as horizontally, after compensating for amplitude distortion by near-surface effects and propagation effects. Quantitative analysis of relative true amplitudes of the Great Lakes International Multidisciplinary Program on Crustal Evolution seismic data is hampered by large uncertainties in estimates of the water bottom reflection coefficient and the vertical amplitude correction and by inadequate noise suppression. Processing techniques such as deconvolution, F-K filtering, and migration significantly change the overall shape of amplitude curves and hence calculation of reflection coefficients and average reflectance. Thus lithological interpretation of deep crustal seismic data based on the absolute value of estimated reflection strength alone is meaningless. The relative strength of individual events, however, is preserved on curves generated at different stages in the processing. We suggest that qualitative comparisons of relative strength, if used carefully, provide a meaningful measure of variations in reflectivity. Simple theoretical models indicate that peg-leg multiples rather than water bottom multiples are the most severe source of noise contamination. These multiples are extremely difficult to remove when the water bottom reflection coefficient is large (>0.6), a condition that exists beneath parts of Lake Superior and most of Lake Huron.

Great Lakes↗

Modeling species invasions in Ecopath with Ecosim: an evaluation using Laurentian Great Lakes models

Invasive species affect the structure and processes of ecosystems they invade. Invasive species have been particularly relevant to the Laurentian Great Lakes, where they have played a part in both historical and recent changes to Great Lakes food webs and the fisheries supported therein. There is increased interest in understanding the effects of ecosystem changes on fisheries within the Great Lakes, and ecosystem models provide an essential tool from which this understanding can take place. A commonly used model for exploring fisheries management questions within an ecosystem context is the Ecopath with Ecosim (EwE) modeling software. Incorporating invasive species into EwE models is a challenging process, and descriptions and comparisons of methods for modeling species invasions are lacking. We compared four methods for incorporating invasive species into EwE models for both Lake Huron and Lake Michigan based on the ability of each to reproduce patterns in observed data time series. The methods differed in whether invasive species biomass was forced in the model, the initial level of invasive species biomass at the beginning of time dynamic simulations, and the approach to cause invasive species biomass to increase at the time of invasion. The overall process of species invasion could be reproduced by all methods, but fits to observed time series varied among the methods and models considered. We recommend forcing invasive species biomass when model objectives are to understand ecosystem impacts in the past and when time series of invasive species biomass are available. Among methods where invasive species time series were not forced, mediating the strength of predator–prey interactions performed best for the Lake Huron model, but worse for the Lake Michigan model. Starting invasive species biomass at high values and then artificially removing biomass until the time of invasion performed well for both models, but was more complex than starting invasive species biomass at low values. In general, for understanding the effect of invasive species on future fisheries management actions, we recommend initiating invasive species biomass at low levels based on the greater simplicity and realism of the method compared to others.

Ecological Modelling↗