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R. Randy Wilson

Publications and source records attributed to R. Randy Wilson.

17 recordsLinked to original sources
USGS2026

Vessel surveys for marine birds in the Gulf of America (Gulf of Mexico): Protocol manual (2017–2025)

This manual provides contextual background and detailed instructions for survey procedures that were used to collect, enter, archive, and analyze digitally-entered field data collected for marine birds during three large-vessel survey projects in the northern Gulf of America (also Gulf of Mexico, hereafter “Gulf”). Projects using this protocol included the Gulf of Mexico Marine Assessment Program for Protected Species (GoMMAPPS: 2017-2018), the Vessel Surveys for Abundance and Distribution of Marine Mammals and Seabirds (VSAD: 2023-2024), and certain repositioning legs of the Atlantic Marine Assessment Program for Protected Species (AMAPPS: 2021, 2023). Based substantially on Ballance and Force (2011), the protocol used in the Gulf is based on a standardized 300 m wide strip transect using software architecture available in program SeeBird (v.4.3.7), a data collection program developed by Robert Holland at the National Oceanic and Atmospheric Administration (NOAA) Southwest Fisheries Science Center, La Jolla, CA. Survey protocols relied on the digital entering of standardized metadata linked to sightings of seabirds (e.g., species, number, location, distance bin, association[s], behavior, flight direction, flight height, angle from ship, age, sex, plumage, and optional comments) as well as other protected species. SeeBird has additional modules that may be used to input data on mixed-species seabird flocks and flying fish (only the latter was used in the Gulf). SeeBird includes “user-driven” commands issued through the keyboard, touch screen, and/or mouse, many of which are aided by drop-down menus that prompt the user options that are available for input. In addition, the program features “event–driven” inputs updated at intervals selected by the user, and that also record exact time and location of such edits. We describe how SeeBird was adapted in the Gulf both to run directly via serial connections from the ship’s navigation system and to operate independently from the ship through a satellite-derived GPS feed. This user manual also gives information about: procedures used to document flying fish abundance on Gulf surveys; how to use the Beaufort scale for standardizing wind force and sea state environmental conditions; how to make and use a hand-held rangefinder for delimiting strip transect bin distances; additional functionalities in SeeBird for which user guides are not available; file-naming, file-archiving, and file location conventions used on Gulf marine bird surveys; protocols used in the Gulf for describing relative abundance of the macro-alga Sargassum as an environmental covariate; the standardized four-letter taxa codes (summarized in a SpeciesCode.txt file) used in the Gulf for denoting species, other taxonomic, and event categories as inputs to SeeBird; an Excel example spreadsheet that has successfully imported SeeBird’s ASCII text file; procedural steps to import, edit, and archive into Excel ASCII text files from SeeBird; initialization instructions for starting up program SeeBird on a Panasonic ToughBook™ that relies on integrated, satellite-derived GPS, and for writing “S” (seabirds + other taxa) and “E” (flying fish) ASCII files to a folder or subfolder other than the default root location on the laptop that contains program SeeBird; how to trouble-shoot miscellaneous start-up problems with program SeeBird; how to add new four-letter codes (into a SpeciesCode.txt file) when additional taxa or objects of interest are encountered. The primary objective of this manual is to archive a permanent, replicable history of survey methodology to facilitate continuity with comparable future projects that may be conducted in the Gulf. A secondary objective is to describe why and how program SeeBird was applied and adapted specifically for operating in Gulf survey conditions. As of late 2025, user guides for software program SeeBird were no longer readily accessible, so this manual serves also to address that gap.

Northern Gulf of America↗
USGS2023

Structured decision making to prioritize regional bird monitoring needs

Conservation planning for large ecosystems has multiple benefits but is often challenging to implement because of the multiple jurisdictions, species, and habitats involved. In addition, decision making at large spatial scales can be hampered because many approaches do not explicitly incorporate potentially competing values and concerns of stakeholders. After the Deepwater Horizon oil spill, establishing baselines was challenging because of (1) variation in study designs, (2) inconsistent use of explicit objectives and hypotheses, (3) inconsistent use of standardized monitoring protocols, and (4) variation in spatial and temporal scope associated with avian monitoring projects before the spill. Herein, we show how the Gulf of Mexico Avian Monitoring Network members used structured decision making to identify bird monitoring priorities. We used multiple tools and techniques to clearly define the problem and stakeholder objectives and to identify bird monitoring priorities at the scale of the entire northern Gulf of Mexico region. Although our example is specific to the northern Gulf of Mexico, this approach provides an example of how stakeholder values can be incorporated into the coordination process of broad-scale monitoring programs to address management, restoration, and scientific questions in other ecosystems and for other taxa.

INFORMS Journal on Applied Analytics↗
USGS2022

Errors in aerial survey count data: Identifying pitfalls and solutions

Accurate estimates of animal abundance are essential for guiding effective management, and poor survey data can produce misleading inferences. Aerial surveys are an efficient survey platform, capable of collecting wildlife data across large spatial extents in short timeframes. However, these surveys can yield unreliable data if not carefully executed. Despite a long history of aerial survey use in ecological research, problems common to aerial surveys have not yet been adequately resolved. Through an extensive review of the aerial survey literature over the last 50 years, we evaluated how common problems encountered in the data (including nondetection, counting error, and species misidentification) can manifest, the potential difficulties conferred, and the history of how these challenges have been addressed. Additionally, we used a double-observer case study focused on waterbird data collected via aerial surveys and an online group (flock) counting quiz to explore the potential extent of each challenge and possible resolutions. We found that nearly three quarters of the aerial survey methodology literature focused on accounting for nondetection errors, while issues of counting error and misidentification were less commonly addressed. Through our case study, we demonstrated how these challenges can prove problematic by detailing the extent and magnitude of potential errors. Using our online quiz, we showed that aerial observers typically undercount group size and that the magnitude of counting errors increases with group size. Our results illustrate how each issue can act to bias inferences, highlighting the importance of considering individual methods for mitigating potential problems separately during survey design and analysis. We synthesized the information gained from our analyses to evaluate strategies for overcoming the challenges of using aerial survey data to estimate wildlife abundance, such as digital data collection methods, pooling species records by family, and ordinal modeling using binned data. Recognizing conditions that can lead to data collection errors and having reasonable solutions for addressing errors can allow researchers to allocate resources effectively to mitigate the most significant challenges for obtaining reliable aerial survey data.

Alabama, Florida, Louisiana, Mississippi, Texas↗
USGS2021

Structured decision making and optimal bird monitoring in the northern Gulf of Mexico

The avian conservation community struggles to design and implement large scale, long-term coordinated bird monitoring programs within the northern Gulf of Mexico due to the complexity of the conservation enterprise in the region; this complexity arises from the diverse stakeholders, multiple jurisdictions, complex ecological processes, myriad habitats, and over 500 species of birds using the region for at least some part of their annual cycle. In addition, long-term monitoring over large spatial scales is difficult because of the need for monitoring data to both (1) evaluate management and restoration outcomes, and (2) provide reliable information about the status and trends of bird populations over time. To address these challenges, the Gulf of Mexico Avian Monitoring Network developed a problem statement: “How can a cost-effective monitoring strategy for the Gulf Coast bird community and ecosystem be developed that evaluates ongoing conservation activities and chronic and acute threats; maximizes learning; and is flexible and holistic enough to detect novel ecological threats and evaluate new and emerging conservation activities?” A structured decision-making framework was then used to articulate and quantify stakeholder values related to the problem statement. One use of the stakeholder values was to develop a regional, strategic plan for bird monitoring, which is presented elsewhere. A formal and complete decision support tool for conservation investments in monitoring and research guided by the stakeholder values is presented in this report. The technical aspects of the stakeholder value model and a portfolio analysis that could be used to guide decision making when allocating resources for monitoring activities is described. Whereas the decision analysis presented here could be useful to any decision maker faced with difficult choices about resource allocation, it is designed for decision makers who request monitoring study proposals and then determine which combination of proposals to fund. The portfolio decision support tool is designed to help funding agencies and organizations identify resource allocation strategies to maximize stated objectives. To begin the decision analysis, an objectives hierarchy and quantitative performance metrics from the values of the Gulf of Mexico bird conservation community were created by a panel of regional stakeholders. Each fundamental objective and sub-objective in the hierarchy is composed of several performance metrics. To test the decision support tool, the authors evaluated a combination of monitoring study proposals written for the region and simulated proposals. Each proposal was scored against the performance metrics and used multi-attribute utility theory to combine the multiple objectives into a measure of total monitoring benefit. The total monitoring benefit and costs of each proposal were then used in a constrained optimization routine to identify optimal monitoring portfolios, that is, a combination of activities that maximizes monitoring benefits while meeting cost and other constraints of interest to stakeholders. A graphical solution based on the concept of Pareto efficiency, which is useful in situations when cost constraints and exact budgets are not known, is also provided. Finally, an evaluation of the sensitivity of the decision-making framework to the weights assigned to objectives by stakeholders is included. This decision support tool allows decision makers to identify an optimal suite of monitoring proposals with a transparent portfolio analysis that includes user-defined constraints (such as costs).

Alabama, Florida, Louisiana, Mississippi, Texas↗
USGS2020

Mississippi Alluvial Valley Forest-breeding landbird population & quantitative habitat objectives

The Mississippi Alluvial Valley (MAV) is a 9 million ha (22-million-acre) floodplain that supports a diverse and ecologically rich bottomland hardwood forest ecosystem – one of the most productive in North America. It extends from roughly Cape Girardeau, Missouri, to the Gulf of Mexico and features a mosaic of ridges, swales, meander belts, and backswamps. Small changes in elevation (<1 foot) in the MAV are associated with large shifts in hydrology, which in turn, strongly affect plant and animal community composition and structure. The resultant diversity contributes to a fertile and productive floodplain. General forest types in the MAV include: Oak-gum-cypress (41%), elm-ash-cottonwood (29%), oakhickory (17%), and the remainder is other forest types (Oswalt 2013). Within the oak-gum-cypress and elm-ash-cottonwood categories, sugarberry-hackberry-elm-green ash and sweetgum-Nuttall oak-willow oak forest types account for close to one-half of MAV bottomland forest acreage, while baldcypress-tupelo forests are about 16 percent (Oswalt 2013). Although we emphasize bottomland hardwood habitat and associated bird species, this planning effort includes analyses based upon all forest types within the MAV. Hence, the term ‘forest’ refers to all forest types in the MAV.

Arkansas, Kentucky, Louisiana, Mississippi, Missou↗
USGS2017

Breeding birds in managed forests on public conservation lands in the Mississippi Alluvial Valley

Managers of public conservation lands in the Mississippi Alluvial Valley have implemented forest management strategies to improve bottomland hardwood habitat for target wildlife species. Through implementation of various silvicultural practices, forest managers have sought to attain forest structural conditions (e.g., canopy cover, basal area, etc.) within values postulated to benefit wildlife. We evaluated data from point count surveys of breeding birds on 180 silviculturally treated stands (1049 counts) that ranged from 1 to 20 years post-treatment and 134 control stands (676 counts) that had not been harvested for >20 years. Birds detected during 10-min counts were recorded within four distance classes and three time intervals. Avian diversity was greater on treated stands than on unharvested stands. Of 42 commonly detected species, six species including Prothonotary Warbler ( Prothonotaria citrea ) and Acadian Flycatcher ( Empidonax virescens ) were indicative of control stands. Similarly, six species including Indigo Bunting ( Passerina cyanea ) and Yellow-breasted Chat ( Icteria virens ) were indicative of treated stands. Using a removal model to assess probability of detection, we evaluated occupancy of bottomland forests at two spatial scales (stands and points within occupied stands). Wildlife-forestry treatment improved predictive models of species occupancy for 18 species. We found years post treatment (range = 1–20), total basal area, and overstory canopy were important species-specific predictors of occupancy, whereas variability in basal area was not. In addition, we used a removal model to estimate species-specific probability of availability for detection, and a distance model to estimate effective detection radius. We used these two estimated parameters to derive species densities and 95% confidence intervals for treated and unharvested stands. Avian densities differed between treated and control stands for 16 species, but only Common Yellowthroat ( Geothlypis trichas ) and Yellow-breasted Chat had greater densities on treated stands.

Mississippi Alluvial Valley↗
USGS2017

How do en route events around the Gulf of Mexico influence landbird populations

Habitats around the Gulf of Mexico (GOM) provide critical resources for Nearctic–Neotropical migratory landbirds, the majority of which travel across or around the GOM every spring and fall as they migrate between temperate breeding grounds in North America and tropical wintering grounds in the Caribbean and Central and South America. At the same time, ecosystems in the GOM are changing rapidly, with unknown consequences for migratory landbird populations, many of which are experiencing population declines. In general, the extent to which events encountered en route limit migratory bird populations is not well understood. At the same time, information from weather surveillance radar, stable isotopes, tracking, eBird, and genetic datasets is increasingly available to address many of the unanswered questions about bird populations that migrate through stopover and airspace habitats in the GOM. We review the state of the science and identify key research needs to understand the impacts of en route events around the GOM region on populations of intercontinental landbird migrants that breed in North America, including: (1) distribution, timing, and habitat associations; (2) habitat characteristics and quality; (3) migratory connectivity; and (4) threats to and current conservation status of airspace and stopover habitats. Finally, we also call for the development of unified and comprehensive long-term monitoring guidelines and international partnerships to advance our understanding of the role of habitats around the GOM in supporting migratory landbird populations moving between temperate breeding grounds and wintering grounds in Mexico, Central and South America, and the Caribbean.

The Condor↗
USGS2016

Object-based forest classification to facilitate landscape-scale conservation in the Mississippi Alluvial Valley

The Mississippi Alluvial Valley is a floodplain along the southern extent of the Mississippi River extending from southern Missouri to the Gulf of Mexico. This area once encompassed nearly 10 million ha of floodplain forests, most of which has been converted to agriculture over the past two centuries. Conservation programs in this region revolve around protection of existing forest and reforestation of converted lands. Therefore, an accurate and up to date classification of forest cover is essential for conservation planning, including efforts that prioritize areas for conservation activities. We used object-based image analysis with Random Forest classification to quickly and accurately classify forest cover. We used Landsat band, band ratio, and band index statistics to identify and define similar objects as our training sets instead of selecting individual training points. This provided a single rule-set that was used to classify each of the 11 Landsat 5 Thematic Mapper scenes that encompassed the Mississippi Alluvial Valley. We classified 3,307,910&plusmn;85,344 ha (32% of this region) as forest. Our overall classification accuracy was 96.9% with Kappa statistic of 0.96. Because this method of forest classification is rapid and accurate, assessment of forest cover can be regularly updated and progress toward forest habitat goals identified in conservation plans can be periodically evaluated.

Arkansas, Louisiana, Mississippi, Missouri↗
USGS2016

Forest structure of oak plantations after silvicultural treatment to enhance habitat for wildlife

During the past 30 years, thousands of hectares of oak-dominated bottomland hardwood plantations have been planted on agricultural fields in the Mississippi Alluvial Valley. Many of these plantations now have closed canopies and sparse understories. Silvicultural treatments could create a more heterogeneous forest structure, with canopy gaps and increased understory vegetation for wildlife. Lack of volume sufficient for commercial harvest in hardwood plantations has impeded treatments, but demand for woody biomass for energy production may provide a viable means to introduce disturbance beneficial for wildlife. We assessed forest structure in response to prescribed pre-commercial perturbations in hardwood plantations resulting from silvicultural treatments: 1) row thinning by felling every fourth planted row; 2) multiple patch cuts with canopy gaps of <1 0.25 – 2 ha; and 3) tree removal on intersecting corridors diagonal to planted rows. These 3 treatments, and an untreated control, were applied to oak plantations (20 - 30 years post-planting) on three National Wildlife Refuges (Cache River, AR; Grand Cote, LA; and Yazoo, MS) during summer 2010. We sampled habitat using fixed-radius plots in 2009 (pre-treatment) and in 2012 (post-treatment) at random locations. Retained basal area was least in diagonal corridor treatments but had greater variance in patch-cut treatments. All treatments increased canopy openness and the volume of coarse woody debris. Occurrence of birds using early successional habitats was greater on sites treated with patch cuts and diagonal intersections. Canopy openings on row-thinned stands are being filled by lateral crown growth of retained trees whereas patch cut and diagonal intersection gaps appear likely to be filled by regenerating saplings.

Conference Paper↗
USGS2014

Avian response to conservation buffers in agricultural landscapes during winter

Native herbaceous vegetation cover along row‐crop field edges (i.e., field buffers) increases breeding densities of many bird species. However, the effect of field buffers on bird species during the non‐breeding season is less understood. We compared density, avian richness, and avian conservation value on row‐crop fields containing buffers strategically designed for wildlife versus fields without buffers in 3 southeastern U.S. states during winter 2007 and 2008. Fields with buffers were enrolled in U.S. Department of Agriculture, Conservation Reserve Program practice Habitat Buffers for Upland Birds (CP33), which targets restoration of northern bobwhite ( Colinus virginianus ) and other upland bird species. Overall species richness did not differ on fields with buffers versus fields without buffers in 2007, but was 29% greater on fields with buffers in 2008. Swamp sparrows ( Melospiza georgiana ), song sparrows ( M. melodia ), field sparrows ( Spizella pusilla ), and red‐bellied woodpeckers ( Melanerpes carolinus ) had greater densities on fields with buffers compared with fields without buffers. Increasing field‐buffer width did not result in greater bird densities. Our results suggest a small change in primary land use (≈7%) produced a disproportionate population response by some grassland‐dependent and woodland bird species during winter. Because field buffers provide a direct source of winter food and cover resources, they may be a pragmatic means to provide critical non‐breeding habitat with little alteration of existing agricultural systems.

Arkansas, Kentucky, Mississippi↗
USGS2011

Distribution and habitat associations of breeding secretive marsh birds in Louisiana's Mississippi Alluvial Valley

Populations of many North American secretive marsh birds (SMBs) have declined in recent decades, partially as a function of wetland loss. Protecting and restoring appropriate habitat for these species is contingent upon understanding the habitat features they utilize. We investigated breeding distributions of SMBs in northeast Louisiana at 118 wetlands in 2007 and 2008 and modeled species occupancy (psi) as a function of habitat variables measured at local (<= 100 m) and landscape (<= 1 km) scales. Common Moorhens (Gallinula chloropus), Least Bitterns (Ixobrychus exilis), and Purple Gallinules (Porphyrula martinica) were the most commonly detected species, whereas breeding King Rails (Rallus elegans) and American Coots (Fulica americana) were rare. Local habitat features consistently played a greater role in predicting psi than landscape features for the three most common species. The proportion of local wetland area dominated by robust emergent vegetation (i.e., Typha spp. and Zizaniopsis miliacea) positively influenced psi for all species, while other wetland vegetation types tended to have a minimal or negative effect. Our results suggest the habitat characteristics preferred by breeding SMBs differ from those used by migrating shorebirds and wintering waterfowl and management and restoration objectives for those species may be inadequate for enhancing SMB habitat.

Louisana↗
USGS2010

Spatial models of Northern Bobwhite populations for conservation planning

Since 1980, northern bobwhite (Colinus virginianus) range-wide populations declined 3.9% annually. Within the West Gulf Coastal Plain Bird Conservation Region in the south-central United States, populations of this quail species have declined 6.8% annually. These declines sparked calls for land use change and prompted implementation of various conservation practices. However, to effectively reverse these declines and restore northern bobwhite to their former population levels, habitat conservation and management efforts must target establishment and maintenance of sustainable populations. To provide guidance for conservation and restoration of habitat capable of supporting sustainable northern bobwhite populations in the West Gulf Coastal Plain, we modeled their spatial distribution using landscape characteristics derived from 1992 National Land Cover Data and bird detections, from 1990 to 1994, along 10-stop Breeding Bird Survey route segments. Four landscape metrics influenced detections of northern bobwhite: detections were greater in areas with more grassland and increased aggregation of agricultural lands, but detections were reduced in areas with increased density of land cover edge and grassland edge. Using these landscape metrics, we projected the abundance and spatial distribution of northern bobwhite populations across the entire West Gulf Coastal Plain. Predicted populations closely approximated abundance estimates from a different cadre of concurrently collected data but model predictions did not accurately reflect bobwhite detections along species-specific call-count routes in Arkansas and Louisiana. Using similar methods, we also projected northern bobwhite population distribution circa 1980 based on Land Use Land Cover data and bird survey data from 1976 to 1984. We compared our 1980 spatial projections with our spatial estimate of 1992 populations to identify areas of population change. Additionally, we used our projection of the spatial distribution and abundance of bobwhite to predict areas of population sustainability. Our projections of population change and sustainability provide guidance for targeting habitat conservation and rehabilitation efforts for restoration of northern bobwhite populations in the West Gulf Coastal Plain.

Journal of Wildlife Management↗
USGS2006

The role of the Wetland Reserve Program in conservation efforts in the Mississippi River Alluvial Valley

The Mississippi River Alluvial Valley includes the floodplain of the Mississippi River from Cairo, Illinois, USA, to the Gulf of Mexico. Originally this region supported about 10 million ha of bottomland hardwood forests, but only about 2.8 million ha remain today. Furthermore, most of the remaining bottomland forest is highly fragmented with altered hydrologic processes. During the 1990s landscape-scale conservation planning efforts were initiated for migratory birds and the threatened Louisiana black bear ( Ursus americanus luteolus ). These plans call for large-scale reforestation and restoration efforts in the region, particularly on private lands. In 1990 the Food, Agriculture, Conservation and Trade Act authorized the Wetlands Reserve Program (WRP). The WRP is a voluntary program administered by the United States Department of Agriculture that provides eligible landowners with financial incentives to restore wetlands and retire marginal farmlands from agricultural production. As of 30 September 2005, over 275,700 ha have been enrolled in the program in the Mississippi River Alluvial Valley, with the greatest concentration in Louisiana, Arkansas, and Mississippi, USA. Hydrologic restoration is common on most sites, with open-water wetlands, such as moist-soil units and sloughs, constituting up to 30% of a given tract. Over 33,200 ha of open-water wetlands have been created, potentially providing over 115,000,000 duck-use days. Twenty-three of 87 forest-bird conservation areas have met or exceed core habitat goals for migratory songbirds and another 24 have met minimum area requirements. The WRP played an integral role in the fulfillment of these goals. Although some landscape goals have been attained, the young age of the program and forest stands, and the lack of monitoring, has limited evaluations of the program's impact on wildlife populations.

Mississippi River Alluvial Valley↗
USGS2003

Spring bird migration in Mississippi Alluvial Valley forests

We surveyed forest songbirds during migration in bottomland hardwood forest stands and managed cottonwood ( Populus deltoides ) plantations in northeast Louisiana and west-central Mississippi between 24 March and 24 May 1996 and 1997. We detected more bird species in bottomland hardwood stands than in cottonwood stands. Within hardwood stands, we detected more individuals in stands subjected to uneven-aged timber harvest than in unmanaged stands. Early in migration, avian species composition was similar in both forest types, being comprised mainly of short-distance migrants. Bird species composition in these forest types became increasingly disparate as long-distance neotropical-nearctic migrants arrived. Ten bird species were characteristic of bottomland hardwood forests, whereas eight different species were characteristic of managed cottonwood plantations. Because these two forest types supported different bird communities, both forest types provide important inland stopover habitat during migration. Silvicultural management of bottomland hardwood forests that increases their understory vegetation will provide forested habitat for a more species rich and abundant population of songbirds during migration.

American Midland Naturalist↗
USGS2002

Avian response to bottomland hardwood reforestation: The first 10 years

Bottomland hardwood forests were planted on agricultural fields in Mississippi and Louisiana predominantly using either Quercus species (oaks) or Populus deltoides (eastern cottonwood). We assessed avian colonization of these reforested sites between 2 and 10 years after planting. Rapid vertical growth of cottonwoods (circa 2–3 m/year) resulted in sites with forest structure that supported greater species richness of breeding birds, increased Shannon diversity indices, and supported greater territory densities than on sites planted with slower-growing oak species. Grassland birds ( Spiza americana [Dickcissel] and Sturnella magna [Eastern Meadowlark]) were indicative of species breeding on oak-dominated reforestation no more than 10 years old. Agelaius phoeniceus (Red-winged Blackbird) and Colinus virginianus (Northern Bobwhite) characterized cottonwood reforestation no more than 4 years old, whereas 14 species of shrub-scrub birds (e.g., Passerina cyanea [Indigo Bunting]) and early-successional forest birds (e.g., Vireo gilvus [Warbling vireo]) typified cottonwood reforestation 5 to 9 years after planting. Rates of daily nest survival did not differ between reforestation strategies. Nest parasitism increased markedly in older cottonwood stands but was overwhelmed by predation as a cause of nest failure. Based on Partners in Flight prioritization scores and territory densities, the value of cottonwood reforestation for avian conservation was significantly greater than that of oak reforestation during their first 10 years. Because of benefits conferred on breeding birds, we recommend reforestation of bottomland hardwoods should include a high proportion of fast-growing early successional species such as cottonwood.

Restoration Ecology↗