Geology topics

Marybeth K. Brey

Publications and source records attributed to Marybeth K. Brey.

At least 19 recordsLinked to original sources

USGS2026

Applying a metapopulation model to compare native and invasive fish passage scenarios in the Upper Mississippi River

The U.S. Geological Survey assessed the ability of metapopulation models to evaluate population-level effects of fish passage projects in the Upper Mississippi River for six native species: Ictalurus furcatus (Valenciennes in Cuvier and Valenciennes, 1840; blue catfish), Ictiobus cyprinellus (Valenciennes in Cuvier and Valenciennes, 1844; bigmouth buffalo), Acipenser fulvescens (Rafinesque, 1817; lake sturgeon), Polyodon spathula (Walbaum, 1792; American paddlefish [paddlefish]), Ictiobus bubalus (Rafinesque, 1818; smallmouth buffalo), Scaphirhynchus platorynchus (Rafinesque, 1818; shovelnose sturgeon), and one invasive species: Hypophthalmichthys molitrix (Valenciennes in Cuvier and Valenciennes, 1844; silver carp). The study had four objectives: to (1) compile parameter values and identify gaps, (2) parameterize species-specific models, (3) apply the models to multiple scenarios, and (4) develop a point-and-click interface. The literature review revealed a paucity of data, for all species except silver carp and paddlefish; however, length–weight relations, female length–maturity curves, growth curves, spawning and recruitment estimates, and movement data among pools were identified. These data allowed the parameterization of native species metapopulation models, in addition to the use of an existing silver carp model. The models were applied to scenarios representing increased passage, decreased passage, combined passage and deterrents effects, and increased native passage with decreased invasive passage. Increased passage scenarios (depicting potential passage projects) produced greater movement and larger upriver populations. Decreased passage scenarios (depicting potential deterrent projects) yielded reduced movement. Similar population dynamic patterns were observed across species for each scenario because of a lack of species-specific empirical movement data. Simulations for longer lived, slow growing species required longer response times, including time to reach equilibria. A point-and-click model was created to allow others to use the model. Data availability for movement limited the ability of the model to be used to understand fish-movement biology and thereby inform management strategies. More fish-movement data could improve model projections of these scenarios.

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

USGS2026

Evaluating the effects of a multi-modal deterrent on upstream passage by invasive carp and native species

Invasive carp, including Silver Carp ( Hypophthalmichthys molitrix ) and Grass Carp ( Ctenopharyngodon idella ), are increasing their range within the Mississippi River Basin, threatening native fishes and ecosystem function. We conducted a 3-year field test of a multi-modal deterrent that uses sound, bubbles, and light to test its ability to impede upstream passage by invasive carp. The test was performed at a lock and dam on the Cumberland River, Kentucky, USA, using acoustic telemetry to track the movements of Silver Carp, Grass Carp, and several native fish species. When the deterrent was operating, the estimated probability of upstream lock passage was 53% lower (95% CI: 41%–63%) for Silver Carp but 74% higher (95% CI: 3%–190%) for Grass Carp compared to times when the deterrent was not operating. Paddlefish ( Polyodon spathula ) passage was reduced by 43% (95% CI: 71% reduction to 9% increase), although this was not statistically significant. We detected no significant effects on Freshwater Drum ( Aplodinotus grunniens ) or Smallmouth Buffalo ( Ictiobus bubalus ). However, sample sizes were small for these native species, limiting our ability to identify effects. Water temperature, tailwater elevation, and vessel lockages affected passage rates, although this varied among species.

Kentucky↗

USGS2026

Silver Carp Hypophthalmichthys molitrix broad movement patterns and dispersal dynamics in the upper Mississippi River

Rivers are important movement corridors for fishes that allow for population connectivity. Within the upper Mississippi River Basin, invasive Silver Carp ( Hypophthalmichthys molitrix ) use tributaries including the Des Moines, Iowa, and Cedar rivers; however, the extent Silver Carp move and intermix among these systems and pools of the upper Mississippi River (UMR) is largely unknown. We used acoustic telemetry and multistate models to (1) identify individual and spatial variability in movement strategies, (2) use survival and transition probability estimates to evaluate population dynamics and connectivity, and (3) evaluate the influence of temporal and environmental variables on survival and transition probabilities. Weekly transition probabilities generally increased with discharge and ranged between 0.01 from UMR Pool 20 to outside the study area at 575 m 3 /s to 0.75 from UMR Pool 18 to outside the study area at 7200 m 3 /s. Survival decreased as temperature increased from 0 to 30 °C and was higher in tributaries than mainstem pools correlated with lower discharge. Overall, Silver Carp displayed diverse movement strategies with high connectivity that can inform invasive species management strategies.

upper Mississippi River↗

USGS2025

Silver Carp passage at three locks and dams on the Tennessee and Cumberland rivers from 2016–2019

Bigheaded carps (i.e., Silver Carp Hypophthalmichthys molitrix and Bighead Carp H. nobilis ) are non-native species that continue to expand their ranges throughout North American river systems, including the Tennessee and Cumberland river systems in the southeastern United States. These species are known to have deleterious effects on native fishes. Management efforts have focused on reducing upstream passage at dams coupled with intensive removal of individuals occupying upstream reaches. Understanding mechanisms affecting the success of upstream fish passage through lock chambers is critical to determining immigration rates into upstream habitats, the likelihood of success for upstream removal efforts, and potential effects of upstream passage reduction using deterrents. We used acoustic telemetry to examine the timing of Silver Carp upstream and downstream dam passages, patterns of fish movement throughout the river systems, and the relationship between fish size and upstream passage through the lock chamber. During 2016–2019, 465 Silver Carp were surgically implanted with transmitters within the two river systems. We documented 37 upstream passages and 57 downstream passages of Barkley, Kentucky, and Pickwick dams. During the two years in which dam passages were observed, most upstream dam passages (89%) occurred during April–August when water temperature ranged from 12–31°C. Most downstream passages (89%) occurred from late-February through July. The relatively small portion of Silver Carp tagged in Pickwick Lake (11% of total tagged), the uppermost reach of the study area, accounted for nearly two-thirds of all dam passages. Our findings may help managers model Silver Carp populations and inform decisions regarding fish deterrent placement and operation.

Alabama, Arkansas, Illinois, Kentucky, Mississippi↗

USGS2025

Tailwater residency patterns of Silver Carp at Kentucky Lock and Dam

Objective The management of invasive Silver Carp Hypophthalmichthys molitrix in the Tennessee River basin focuses on removal, and there is interest in extending removal efforts to the tailwater environments of high-head locks and dams along the Tennessee River, such as Kentucky Dam. We used acoustic telemetry data from Silver Carp to understand important ecological associations underlying their residence in the Kentucky Dam tailwater, measured by daily fish counts and mean residence time. Methods We used time-series-informed regression models, variance partitioning, and cross-correlation function analysis to associate six predictors, including lock and dam operations (total, spill gate, and turbine discharge and number of lockages), hydrology (tailwater elevation), and water temperature, with two measures of Silver Carp residency (daily counts and mean residence time). Results We found that spill-induced hydrology (total discharge + spill discharge + tailwater elevation) was negatively associated with daily counts but not with residence time, whereas temperature was positively associated with counts and negatively associated with residence times. Variance partitioning indicated that nearly all the variance in counts and residence times was jointly explained by temporal effects, lock and dam operations (discharge, tailwater elevation, and lockages), and temperature. The cross-correlations indicated that the counts were lagged by all predictors, sometimes up to 5 d, whereas residence times were lagged by both total and spill discharge and number of lockages. Conclusions We found that discharge and water temperature were principally associated with residency of Silver Carp in the Kentucky Dam tailwater. However, these associations were entirely temporally constrained, which can affect how strongly and how quickly Silver Carp respond to changing environmental conditions across different time scales. Managers can leverage these associations to plan removal periods where daily tailwater conditions/dam operations are favorable to invasive carp residence (e.g., >10°C and <2,500 m 3 /s) and adjust fishing effort to optimize removal rates in response to changing conditions.

Kentucky Lock and Dam↗

USGS2025

Considerations for using tag-returns to monitor targeted removal of invasive fishes

Objective Targeted removals are used for management of some invasive fish populations. Tag–return studies are one approach that can be used to assess the efficacy of targeted removals. However, there are many decisions to make when designing a tag–return study. We used simulation modeling to outline general guidelines for consideration when designing efficient tag–return studies to measure annual removal rates of invasive fish, particularly invasive carps. Methods We simulated data sets using scenarios with varying numbers of fish tagged per year, removal rates, tag reporting rates, tag retention rates, and study durations. We generated the data sets under a set of “known” parameters with added stochasticity; we then fitted the simulated data sets to a Bayesian tag–return model and measured the precision and accuracy of the model-estimated removal rates. Results We found that the model was able to predict removal rates without bias for most of the scenarios. However, we did find patterns in the precision of the predictions that could help to inform tag–return studies. When the proportion of the population removed through harvest was constant, the proportion of the population removed per year and the probability that harvested tags were reported had the largest effect on precision. The number of tags released per year and the study duration also had moderate effects. For scenarios testing the ability of the model to predict removal rates in stochastic populations, the precision of the model was primarily influenced by the number of fish tagged, the underlying nature of the stochasticity, and whether fish were tagged during the year of the prediction. Conclusions Based on our simulations, we outline how study objectives, the underlying population variability, and the tolerance range for error can guide decisions regarding the number of fish to tag, how to monitor tag return rates, and how long to conduct a study.

North American Journal of Fisheries Management↗

USGS2025

Behavioral responses of Silver Carp to underwater acoustic deterrent sounds

Objective Invasive carps continue to spread across the Mississippi River basin, posing significant ecological risk. Identifying technologies to slow their dispersal is critical. The use of sound has been proposed as a method to modify the behavior of Silver Carp Hypophthalmichthys molitrix , offering a nonstructural deterrent strategy. Methods Silver Carp implanted with acoustic transmitters were released into earthen ponds equipped with telemetry arrays. The fish were exposed to a 30-min playback of three underwater sounds (chirp saw, chirp square, and 100-hp boat motor). Movement trajectories were analyzed using a two-state hidden Markov model to estimate the effects of environmental and experimental variables on fish behavior. Results The results of the hidden Markov model supported two behavioral states. State 1 was characterized by longer step lengths (distance between positions) and greater directional persistence in turning angle (change in direction between two intervals), indicative of heighted activity. State 2 was defined by shorter step lengths and less directional persistence, suggesting reduced activity. Silver Carp that were exposed to the chirp square sound had an increased likelihood of entering state 1, whereas the 100-hp boat motor sound promoted transitions to state 2. Conclusions Underwater sounds distinctly influenced the movement of Silver Carp in earthen ponds. The chirp square sound elicited heightened activity levels, demonstrating potential for use in acoustic deterrent applications. However, the response of Silver Carp to these sounds may be influenced by the size of the study environment or the absence of natural drivers of fish behavior, such as food or reproduction. This study contributes to the development of nonstructural, species-specific deterrent systems by identifying sounds that influence the behavior of invasive carps. The application of sound-based methods may play a critical role in integrated pest management strategies for invasive carps, potentially limiting their spread while minimizing effects on native species.

Transactions of the American Fisheries Society↗

USGS2025

Vertebrates in trade that pose high invasion risk to the United States

The United States imports thousands of live vertebrate species annually as part of legal trade. Escapes and releases from captivity are major pathways of invasion, however, the risk posed by the thousands of imported vertebrate species has not been systematically assessed. We conducted a horizon scan that used a data-driven climate match to filter a list of nearly 15,000 taxa drawn from across the globe of imported fish, amphibians, reptiles, birds, and mammals for rapid assessment by taxonomic experts. Experts evaluated 840 species and identified 32 (22 reptiles and 10 fishes) as having the highest risk for establishment, spread, and negative impacts. Of those high-risk species, the majority have the capacity to disrupt ecosystem processes via their role as top predators or the unique ecological niches that they occupy, while several of the snake species pose a threat to human health. High-risk species were often scored with high confidence while in contrast, low scores were attributed to a combination of ecological redundancy, low propagule pressure, or low climate match while low confidence arose from a lack of information in the literature (i.e., data deficiency). Our study therefore highlights legally imported species likely to cause the greatest harm with the recognition that many other species could also become invasive in the United States. The ranked list of vertebrate threats can be used to prioritize watchlists and inform the development of targeted regulations for importation can be applied to regions to provide a rapid, preliminary screening for large pools of potential invaders.

Biological Conservation↗

USGS2025

USGS invasive carp database management and integration support

Invasive carp tracking, monitoring, and contracted removal will continue throughout the Illinois River and upper Mississippi River as part of an adaptive management effort to mitigate, control, and contain invasive carp. To facilitate these actions, a need to compile and analyze invasive carp-related data from all agencies exists. Invasive carp-related data include all data sources that could inform the MRWG objectives or projects. This data, often in disparate formats, must be integrated into a common format that allows all agencies the opportunity to assess invasive carp monitoring, control, and removal efforts. Ensuring the interoperability of these datasets allows for their use in various analyses and modeling efforts. Implementing an interoperable data management framework provides mechanisms for endusers to find and use integrated data. Integrating data for use in modeling and analysis furthers the partnership’s collective understanding of invasive carp life history, distribution, and movement and can be used to facilitate adaptive management actions (e.g., directing monitoring, sampling, and removal efforts; assessing invasive carp abundance to support modeling efforts; informing deployment of control actions; etc.). An effective data management strategy will streamline the data update process, providing all agencies with timely data and analyses in support of informed decision-making processes.

Interim Summary Report↗

USGS2024

Flooding and dam operations facilitate rapid upstream migrations of native and invasive fish species on a regulated large river

Dams commonly restrict fish movements in large rivers but can also help curtail the spread of invasive species, such as invasive bigheaded carps ( Hypophthalmichthys spp). To determine how dams in the upper Mississippi River (UMR) affect large-scale invasive and native fish migrations, we tracked American paddlefish ( Polyodon spathula ) and bigheaded carp across > 600 river km (rkm) and 16 navigation locks and dams (LD) of the UMR during 2 years with contrasting water levels. In 2022, a low-water year, both native paddlefish and invasive bigheaded carp had low passage rates (4% and 0.6% respectively) through LD15, a movement bottleneck being studied for invasive carp control. In contrast, flooding in 2023 led to open-river conditions across multiple dams simultaneously, allowing 53% of paddlefish and 46% of bigheaded carp detected in Pool 16 to move upstream through LD15. Bigheaded carp passed upstream through LD15 rapidly (μ = 32 rkm per day) a maximum of 381 rkm, whereas paddlefish moved an average of 9 upstream rkm per day (maximum of 337 rkm). Our results can inform managers examining trade-offs between actions that enhance native fish passage or deter movements of invasive species. This understanding is critical because current climate change models project increases in flooding events like that observed during 2023.

Illinois, Iowa, Minnesota, Wisconsin↗

USGS2024

Bayesian multistate models for measuring invasive carp movement and evaluating telemetry array performance

Understanding the movement patterns of an invasive species can be a powerful tool in designing effective management and control strategies. Here, we used a Bayesian multistate model to investigate the movement of two invasive carp species, silver carp ( Hypophthalmichthys molitrix ) and bighead carp ( H. nobilis ), using acoustic telemetry. The invaded portions of the Illinois and Des Plaines Rivers, USA, are a high priority management zone in the broader efforts to combat the spread of invasive carps from reaching the Laurentian Great Lakes. Our main objective was to characterize the rates of upstream and downstream movements by carps between river pools that are maintained by navigation lock and dam structures. However, we also aimed to evaluate the efficacy of the available telemetry infrastructure to monitor carp movements through this system. We found that, on a monthly basis, most individuals of both species remained within their current river pools: averaging 76.2% of silver carp and 75.5% of bighead carp. Conversely, a smaller proportion of silver carp, averaging 14.2%, and bighead carp, averaging 13.9%, moved to downstream river pools. Movements towards upstream pools were the least likely for both species, with silver carp at an average of 6.7% and bighead carp at 7.9%. The highest probabilities for upstream movements were for fish originating from the three most downstream river pools, where most of the population recruitment occurs. However, our evaluation of the telemetry array’s effectiveness indicated low probability to detect fish in this portion of the river. We provide insights to enhance the placement and use of these monitoring tools, aiming to deepen our comprehension of these species’ movement patterns in the system.

Illinois↗

USGS2024

Upstream experience and experimental translocation of invasive bigheaded carps results in increased upstream passage success at a navigation lock in a large river

Fish movements in regulated rivers can be challenging to study because anthropogenic modifications, such as locks and dams, can influence animal behavior. Upper Mississippi River Lock and Dam 19 (LD 19), for example, is an invasive carp movement bottleneck due to an impassable dam. Upstream fish passage at LD19 is restricted to the lock chamber, making it an optimal location to test invasive fish deterrents that could limit further range expansion. Evaluating the effectiveness of experimental deterrents requires baseline knowledge of fish movements and suitable sample sizes of fish encountering the deterrents to ensure adequate statistical power. Some evidence indicates fish with prior upstream experience may return upstream or challenge potential deterrents at a higher rate than fish without such experience. To test how previous upstream experience could increase the rate at which fish moved upstream through a navigation lock chamber, we compared upstream passage through LD 19 using bigheaded carp captured below the dam (downstream-origin) and two groups of bigheaded carp captured upstream from the dam: those that swam downstream on their own volition (upstream-origin fish) and those that were captured upstream and translocated downstream of LD 19 (translocated upstream-origin). Translocated upstream-origin carp demonstrated the highest rate of upstream passage, with 59% of the fish detected downstream from LD 19 passing upstream during our study. In contrast, downstream-origin carp made no upstream passages over 2 years. Fish origin was shown to influence upstream passage success. This may be an important consideration for fish passage studies and deterrent evaluations.

Illinois, Iowa↗

USGS2024

USGS invasive carp database management & integration support

Invasive carp tracking, monitoring, and contracted removal will continue throughout the Upper IWW system as part of an adaptive management effort to mitigate, control, and contain invasive carp. To help facilitate these actions, there is a need to compile and analyze data from the multitude of partner agencies that are collecting invasive carp-related data throughout the Illinois River system. These data are often in disparate formats, and integrating these data into a common format allows both researchers and managers to assess invasive carp monitoring, control, and removal efforts at several scales is necessary. Ensuring the interoperability of these data sets allows for their use in various analyses and modeling efforts. Implementing an interoperable data management framework also provides mechanisms for end users to find and use existing data. Integrating data for use in modeling and analysis furthers the MRWG partnership’s collective understanding of invasive carp life history, distribution, and movement and can be used to facilitate adaptive management actions (e.g., directing monitoring, sampling, and removal efforts; assessing invasive carp abundance to support modeling efforts; informing deployment of control actions). An effective data management strategy will streamline the database update process, providing partners with timely, well-organized, uniformly formatted data and analyses in support of informed decision-making processes.

Illinois↗

USGS2024

USGS invasive carp database management and integration support

Invasive carp tracking, monitoring, and contracted removal will continue throughout the Illinois River and upper Mississippi River as part of an adaptive management effort to mitigate, control, and contain invasive carp. To facilitate these actions, a need to compile and analyze invasive carp-related data from all agencies exists. Invasive carp-related data include all data sources that could inform the MRWG objectives or projects. These data, often in disparate formats, must be integrated into a common format that allows all agencies the opportunity to assess invasive carp monitoring, control, and removal efforts. Ensuring the interoperability of these datasets allows for their use in various analyses and modeling efforts. Implementing an interoperable data management framework provides mechanisms for endusers to find and use integrated data. Integrating data for use in modeling and analysis furthers the partnership’s collective understanding of invasive carp life history, distribution, and movement and can be used to facilitate adaptive management actions (e.g., directing monitoring, sampling, and removal efforts, assessing invasive carp abundance to support modeling efforts, informing deployment of control actions, etc.). An effective data management strategy will streamline the data update process, providing all agencies with timely data and analyses in support of informed decision-making processes.

Interim Summary Report↗

USGS2024

Evaluation of fish passage for assessment of invasive carp deterrents at locks in the upper Mississippi River

Invasive carps are established in the upper, middle, and lower Mississippi River and their expansion upstream threatens a variety of aquatic ecosystem services including fishing and recreational boating. The physical and operational characteristics of Lock and Dam (LD) 19 restrict upstream movement of fishes because the only upstream fish passage route is through the lock chamber. This restriction might be hindering consistent reproduction and recruitment of invasive carps enough to reduce their abundance upstream from LD 19. Locks and Dams 5, 14, and 15 (upriver from LD 19) are infrequently at open-condition and may also be limiting the continued upstream expansion of invasive carps. Upstream passage of fishes at these locations would be limited to the lock chamber for the majority of the year (Wilcox et al. 2004; Bouska 2021).

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

USGS2024

Evaluation of fish passage for assessment of invasive carp deterrents at locks in the Upper Mississippi River

Invasive carp are established in the upper, middle, and lower Mississippi River and their expansion upstream threatens a variety of aquatic ecosystem services including fishing and recreational boating (U.S. Geological Survey, 2021). The physical and operational characteristics of Lock and Dam (LD) 19 restrict upstream movement of fishes because the only upstream fish passage route is through the lock chamber. This restriction might be hindering consistent reproduction and recruitment of Silver Carp Hypophthalmichthys molitrix and bighead carp H. nobilis (collectively referred to as bigheaded carp) enough to reduce their abundance upstream of LD 19. Locks and Dams 14 and 15 are infrequently in open-river condition (defined as the time when the adjustable spillway gates of the dam are raised out of the water, passing unobstructed water through the gates) and may also be limiting the continued upstream spread of invasive carp populations. Upstream passage of fishes at these locations would be limited to the lock chamber for the majority of the year (Wilcox et al., 2004; Bouska, 2021).

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

USGS2023

Multimodal invasive carp deterrent study at Barkley Lock and Dam—Status update through 2022

Invasive carp ( Hypophthalmichthys nobilis [Bighead Carp], Mylopharyngodon piceus [Black Carp], Ctenopharyngodon idella [Grass Carp], and H. molitrix [Silver Carp]) continue to spread in the United States and deterrents at river navigation locks are one emerging control strategy for slowing the spread. High-head navigation dams on large rivers serve as impediments to the upstream spread of these populations. One possible control technique is using a multimodal deterrent that utilizes a combination of lights, sound, and air bubbles to guide fish away from a location. Laboratory tests and small-scale field deployments of similar multi-stimuli deterrents have demonstrated potential for deterring invasive carp. These earlier studies led to the deployment and initiation of a field study of a multimodal deterrent in 2019 at the downstream lock approach at Barkley Lock and Dam on the Cumberland River near Grand Rivers, Kentucky. We are using two types of telemetry systems to evaluate how the multimodal deterrent affects movement and behavior of Silver Carp, Grass Carp, and four native species: Aplodinotus grunniens (Freshwater Drum), Polyodon spathula (Paddlefish), Ictiobus bubalus (Smallmouth Buffalo), and Acipenser fulvescens (Lake Sturgeon). We are evaluating fish movements in response to the multimodal deterrent by using a study design that cycles between 1 week with the multimodal deterrent on and 1 week with it off. This weekly cycling provides baseline comparisons across variable environmental conditions. When the multimodal deterrent is on, the number of Silver Carp completing upstream passage through Barkley Lock is reduced by about one-half. This study that began in 2019 can provide insights into the effectiveness of a multimodal deterrent for limiting upstream movement of invasive carp or effects on native fishes through strategic navigation locks on large rivers.

Kentucky↗

USGS2023

Lock 19 underwater acoustic deterrent system study—Interim project update, through 2022

Invasive carp ( Hypophthalmichthys nobilis [Richardson, 1845; Bighead Carp], H. molitrix [Valenciennes in Cuvier and Valenciennes, 1844; Silver Carp], Ctenopharyngodon idella [Valenciennes in Cuvier and Valenciennes, 1844; Grass Carp], and Mylopharyngodon piceus [Richardson, 1846; Black Carp]) expansion threatens the Laurentian Great Lakes and other major waterways. Numerous tools and techniques are being tested or developed to curtail the upstream expansion of invasive carps while minimizing the effect to native species. Underwater sound is one technology that has shown promise as a deterrent in the laboratory and ponds and is being considered for use at bottleneck locks and dams to reduce the risk of invasive carps moving to uninvaded areas. To test this technology at a management-relevant scale, an underwater acoustic deterrent system (UADS) was designed and installed at Lock 19 on the Mississippi River in 2021. The experimental UADS operates on a continuous schedule of 80 hours on and 80 hours off to allow for comparisons of fish behavior during varying environmental, operational, and navigation traffic conditions. Two telemetry systems, operating at 69 and 307 kilohertz, were deployed to evaluate how the UADS may affect movement and behavior of invasive carps and native fish species. During 2021–22, Silver Carp were twice as likely to pass over the UADS and into the lock when it was off compared to on; however, Bigmouth Buffalo, a native fish, were 1.2 times more likely to make upstream passage when the UADS was off compared to on.

Illinois, Iowa↗