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M. Gabriela Mángano

Publications and source records attributed to M. Gabriela Mángano.

7 recordsLinked to original sources

Trace fossils and sedimentary facies from a Late Cambrian‐Early Ordovician tide‐dominated shelf (Santa Rosita Formation, northwest Argentina): Implications for ichnofacies models of shallow marine successions

The Santa Rosita Formation is one the most widely distributed lower Paleozoic units of northwest Argentina. At the Quebrada del Salto Alto section, east of Purmamarca, Jujuy Province, it is represented by four sedimentary facies: thick‐bedded planar cross‐stratified quartzose sandstones (A), thin‐bedded planar cross‐stratified quartzose sandstones and mudstones (B), wave‐rippled sandstones and bioturbated mudstones (C), and black and greenish gray shales (D). Paleocurrent data, sandstone architecture, and sedimentary structures from faciès A and B indicate bipolar/bimodal paleoflows, suggesting the action of tidal currents. The succession is interpreted as that of a tide‐dominated shelf, with only secondary influence of wave processes. Trace fossils are restricted to facies B and C. The Cruziana ichnocoenosis is preserved on the soles of thin‐bedded planar cross‐stratified quartzose sandstones (faciès B). This ichnocoenosis consists of Conostichus isp., Cruziana omanica, C. semiplicata, C. cf. tortworthi, Cruziana isp. Helminthopsis abeli, Monomorphichnus bilinearis, M. multilineatus, Palaeophycus tubularis, Rusophycus carbonarias, R. latus , and R. isp. The occurrence of Cruziana semiplicata , C. omanica, C. cf. tortworthi , and Rusophycus latus supports a Late Cambrian‐Tremadoc age. Slabbing of Cruziana shows complex interactions between biologic and sedimentologic processes, and suggests a predominance of exhumed traces, washed out and recast by tractive sand deposition. Sandstone soles are densely packed with biogenic structures and exhibit distinctive clusters of Rusophycus isp. that most likely represent trilobite nesting burrows. The Cruziana ichnocoenosis records the resident fauna of a protected, lower intertidal to subtidal interbar setting. The Skolithos ichnocoenosis is represented by high to low density vertical burrows of Skolithos linearis , which extend downwards to the quartzose sandstone soles of faciès B and cross the Cruziana ichnocoenosis. The Skolithos ichnocoenosis represents colonization by suspension‐feeding organisms following a major change in environmental conditions, related to the migration of lower intertidal to subtidal sandwaves. The Planolites ichnocoenosis consists exclusively of Planolites montanus within mudstones overlying wave‐rippled sandstones (facies C). The Planolites ichnocoenosis records opportunistic colonization by inf aunal deposit feeders that mined the organic‐rich fine‐grained sediment during the waning phase of storms that scoured organic detritus from the sea bottom. The section records, from base to top, a Cruziana‐Skolithos ichnofacies zone, a Skolithos ichnofacies zone and an unbioturbated zone typified by the thick‐bedded cross‐stratified quartzose sandstone (fades A). This trend reflects progressively higher energy conditions linked to the establishment of a large sand wave complex. The presence of a mixed Cruziana‐Skolithos ichnofacies in the lower interval reflects changes in substrate and energy levels, rather than water depth. Accordingly, contrasting ichnocoenoses from interbars (Cruziana) and sandwaves (Skolithos) must be considered an example of ichnofacies controlled by local parameters instead of general bathymétrie trends. Conversely, the vertical replacement of the Cruziana ichnofacies by the Skolithos ichnofacies towards the middle interval of the section reflects the environmental changes associated with the transition between the intertidal and subtidal zones. As overall tidal energy increases from supratidal to subtidal settings, the Skolithos ichnofacies tends to occur seaward of the Cruziana ichnofacies in tide‐dominated shallow marine environments. Therefore, onshore‐offshore ichnofacies replacement in tide‐dominated shallow seas is opposite to that in wave‐dominated marine settings.

Ichnos: An International Journal for Plant and Ani↗

Carboniferous Psammichnites: Systematic re-evaluation, taphonomy and autecology

The ichnogenus Psammichnites Torell 1870 includes a wide variety of predominantly horizontal, sinuous to looped, backfilled traces, characterized by a distinctive median dorsal structure. Though commonly preserved in full relief on upper bedding surfaces, some ichnospecies of Psammichnites may be preserved in negative hyporelief. Psammichnites records the feeding activities of a subsurface animal using a siphon-like device. Several ichnogenera reflect this general behavioral pattern, including Plagiogmus Roedel 1929 and the Carboniferous ichnogenera Olivellites Fenton and Fenton 1937a and Aulichnites Fenton and Fenton 1937b. Based on analysis of specimens from the United States, Spain, and the United Kingdom, three Carboniferous ichnospecies of Psammichnites are reviewed in this paper: P. plummeri (Fenton and Fenton, 1937a), P. grumula (Romano and Meléndez 1979), and P. implexus (Rindsberg 1994). Psammichnites plummeri is the most common Carboniferous ichnospecies and is characterized by a relatively straight, continuous dorsal ridge/groove, fine transverse ridges, larger size range, and non-looping geometric pattern. It represents a grazing trace of deposit feeders. Psammichnites grumula differs from the other ichnospecies of Psammichnites by having median dorsal holes or protruding mounds. The presence of mounds or holes in P. grumula suggests a siphon that was regularly connected to the sediment-water interface. This ichnospecies is interpreted as produced by a deposit feeder using the siphon for respiration or as a device for a chemosymbiotic strategy. Psammichnites implexus is characterized by its consistently smaller size range, subtle backfill structure, and tendency to scribble. Although displaying similarities with Dictyodora scotica , P. implexus is a very shallow-tier, grazing trace. Changes in behavioral pattern, preservational style, and bedform morphology suggest a complex interplay of ecological and taphonomic controls in Carboniferous tidal-flat Psammichnites . A first distributional pattern consists of guided meandering specimens preserved in ripple troughs, probably reflecting food-searching of buried organic matter concentrated in troughs. A second is recorded by concentration of Psammichnites on ripple crests and slopes. In some cases, the course is almost straight to slightly sinuous and closely follows topographic highs, suggesting a direct control of bedform morphology on trace pattern. Occurrences of Carboniferous Psammichnites most likely represent an opportunistic strategy in marginal-marine settings. Analysis of Carboniferous Psammichnites indicates the presence of a siphon-like device in the producer and reestablishes the possibility of a molluscan tracemaker.

Ichnos: An International Journal for Plant and Ani↗

The insect trace fossil Tonganoxichnus from the middle Pennsylvanian of Indiana: Paleobiologic and paleoenvironmental implications

The ichnogenus Tonganoxichnus , produced by one or more monuran insect taxa, is now recorded from the Middle Pennsylvanian Mansfield Formation of Indiana. Tonganoxichnus is a resting trace that has three important implications. First, it represents a recurrent behavioral pattern in Upper Carboniferous to Lower Permian marginal marine environments of North America. Second, it provides finely resolved anatomical information for axial and appendicular body structures and behaviors that are difficult to determine from body‐fossil material alone. Third, integrated sedimentologic and ichnologic observations indicate that the Tonganoxichnus assemblage, inclusive of other ichnotaxa, is common in tidal rhythmites that were developed under freshwater conditions, probably in the innermost part of estuarine systems, close to or at the fluvioestuarine transition.

Ichnos: An International Journal for Plant and Ani↗

The origin and paleoecologic significance of the trace fossil Asteriadtes in the Pennsylvanian of Kansas and Missouri

The trace fossil Asteriacites , recorded in Cambrian to Recent shallow- and deep-marine facies, is traditionally interpreted as the resting trace of asterozoans. Well-preserved specimens of A. lumbricalis are abundant in Pennsylvanian (Upper Carboniferous) shallow- and marginal-marine siliciclastic deposits of eastern Kansas and western Missouri. Detailed morphologic analysis of these specimens suggests that they record the activities of mobile epifaunal ophiuroids. Evidence of a brittle star (ophiuroid) producer rather than sea star (asteroid) is provided by (1) trace-fossil morphologic features reflecting the anatomy of the producer (e.g., well-differentiated central structure, slender vermiform arms) and ophiuroid burrowing technique (e.g., proximal arm expansion, arm branching), and (2) mode of occurrence (e.g., gregarious behavior, horizontal and vertical repetition). Vertical and horizontal repetition produces complex aggregates of A. lumbricalis that are interpreted either as escape structures (fugichnia) or as feeding structures, respectively. Ophiura texturata is proposed as a modern analogue for the A. lumbricalis producer, based on inferred life habit and feeding behavior. Asteriacites lumbricalis is present in two different intertidal trace-fossil assemblages. The first assemblage is characterized by high diversity and records tidal flats developed outside of embayments under normal marine conditions. The second assemblage consists of A. lumbricalis together with a few other ichnotaxa and represents a depauperate association that developed in restricted tidal flats within an embayment or estuarine setting. This challenges the conventional view of Asteriacites as a normal-marine salinity indicator. Some echinoderms, and particularly asterozoans, penetrate and inhabit modern environments of depressed salinity. The presence of Asteriacites in Pennsylvanian marginal-marine facies of Kansas and Missouri provides evidence that ophiuroids had adapted to brackish-water conditions by the late Paleozoic.

Kansas, Missouri↗

Grazing trails formed by soldier fly larvae (Diptera: Stratiomyidae) and their paleoenvironmental and paleoecological implications for the fossil record

Recent trails formed by soldier fly larvae (Díptera: Stratiomyidae) were examined in a shallow pond in the floodplain of a braided river in Jujuy Province, northwestern Argentina. Collected specimens were identified as Stratiomys convexa van der Wulp. Simple, irregularly meandering trails were produced across the surface of a muddy‐silty substrate. Since soldier fly larvae extend their caudal respiratory tube to the water‐air interface, they are restricted to extremely shallow water. The presence of benthic algal remains within the mouthparts of the larvae suggests a feeding habit of algal grazing. If preserved, these trails would be included in the ichno‐genus Helminthopsis, a common element in ancient freshwater ichno‐faunas. Helminthopsis preserved in pond and lacustrine margin deposits younger than Late Jurassic is regarded as one possible trace fossil analogue of the trails documented herein. Additionally, it is suggested that larvae of many aquatic Diptera with similar ecologic restrictions are potential tracemakers of Helminthopsis and other simple trails in these environments, particularly in post‐Paleozoic deposits. Studies of modern shallow aquatic habitats and reex‐amination of the ichnologic record stress the importance of fly larvae as tracemakers in lake margin and pond ecosystems. Ecologic requirements of soldier fly larvae make them inappropriate analogues of Helminthopsis tracemakers in deeper water, lacustrine settings.

Ichnos: An International Journal for Plant and Ani↗

Trace fossils from Jurassic lacustrine turbidites of the Anyao Formation (Central China) and their environmental and evolutionary significance

The Lower Jurassic Anyao Formation crops out near Jiyuan city, western Henan Province, central China. It is part of the infill of the nonmarine early Mesozoic Jiyuan‐Yima Basin. In the Jiyuan section, this unit is about 100 m thick and consists of laterally persistent, thin and thick‐bedded turbidite sandstones and mudstones displaying complete and base‐or top‐absent Bouma sequences, and thick‐bedded massive sandstones. The Anyao Formation records sedimentation within a lacustrine turbidite system developed in a pull‐apart basin. Processes involved include high and low density turbidity currents, sometimes affected by liquefaction or fluidization. Facies analysis suggests that this succession is formed by stacked aggradational turbidite lobes. The absence of thick mudstone packages indicates that background sedimentation was subordinate to high frequency turbidite deposition. The Anyao Formation hosts a moderately diverse ichnofauna preserved as hypichnial casts on the soles of thin‐bedded turbidite sandstones. The ichnofauna consists of Cochlichnus anguineus, Hel‐minthoidichnites tenuis, Helminthopsis abeli, H. hieroglyphica, Mono‐morphichnus lineatus, Paracanthorhaphe togwunia, Tuberculichnus vagans, Vagorichnus anyao, tiny grazing trails, and irregularly branching burrows. Vagorichnus anyao occurs not only as a discrete trace, but also as a compound ichnotaxon intergrading with Gordia marina and Tuberculichnus vagans. Both predepositional and post‐depositional traces are present on the soles of turbidites. This ichnofauna comprises both feeding and grazing traces produced by a deposit‐feeding lacustrine benthic biota. Crawling traces are rare. Although certain ichnofossils (e.g. V. anyao, P. togwunia) show overall similarities with deep‐sea agrichnia, they differ in reflecting remarkably less specialized feeding strategies, displaying overcrossing between specimens (and to a lesser extent, self‐crossing), and in the case of V’ anyao recording post‐turbidite burrowing activity. The development of less specialized strategies than those displayed by deep‐marine ichnofaunas may be related to less stable conditions, typical of lake settings. Oxyenation, energy, sedimentation rate (both event and background), food supply, soft‐sediment deformation and erosion rate have mainly influenced trace‐fossil distribution. Turbidity currents would have ensured oxygen (as well as food) supply to deep lake settings, thus allowing the establishment of a moderately diverse biota. Biogenic structures were mostly confined to the outer, low energy areas. High sedimentation rates and strong erosion precluded preservation of ichnofossils in inner lobe settings. The Anyao ichnofauna is of significance in furthering knowledge of the colonization of deep lakes throughout the stratigraphie record and in identification of additional nonmarine ichnofacies. The ana‐lyzed ichnofauna resembles late Paleozoic lacustrine assemblages described from different localities around the world and is regarded as a Mesozoic example of the Mermia ichnofacies. However, when compared with Paleozoic assemblages, the Anyao ichnofauna shows a clear dominance of burrows over surface trails, deeper burrowing penetration, larger size, and presence of relatively more complex structures. The high burrow/surface trail ratio may be indicative of lower preservation potential in the latter, thus reflecting a tap‐honomic overprint. In contrast to the Paleozoic examples, the establishment of a relatively well‐developed lacustrine infauna may have precluded preservation of surface trails. Burrower activity probably obliterated biogenic structures formed close to the sediment‐water interface.

Ichnos: An International Journal for Plant and Ani↗

Taxonomic reassessment of the ichnogenus Beaconichnus and additional examples from the Carboniferous of Kansas, U.S.A.

The ichnogenus Beaconichnus (Gevers 1973), an arthropod trace fossil, includes very different forms that comprise five ichnospecies, namely B. darwi‐nunt (Gevers 1971), B. gouldi (Gevers 1971), B. ahtarcticum (Gevers 1971), B. giganteum Gevers and Twomey 1982, and B. wrrighti Gevers and Twomey 1982. The original diagnosis of Beaconichnus is rather vague and potentially may accommodate virtually every arthropod trackway described from the fossil record. In view of these problems, the validity of Beaconichnus is reassessed and each of its ichnospecies is reviewed. We conclude that B. darwinum is a junior synonym of Diplopodichnus biformis Brady 1947; B. antarcticum should be regarded as Palmich‐niunt antarcticum; and B. wrighti is a nomen nudum. Additionally, we agree with previous proposals in considering B. gouldi as the senior synonym of B. giganteum , and including it in Diplichnites Dawson 1873. Therefore, we suggest that the ichnogenus Beaconichnus is best disregarded. Additionally, we describe specimens collected from the Late Carboniferous Tonganoxie Sandstone Member (Stranger Formation) of eastern Kansas, ascribed herein to Diplopodichnus biformis and Diplichnites gouldi , which include examples of intergradations between both ichnotaxa, and provide synonymy lists for both ichnospecies.

Kansas↗