Search USGS⌕ Search

Geology topics

H.D. Safford

Publications and source records attributed to H.D. Safford.

5 recordsLinked to original sources

Large-scale causes of variation in the serpentine vegetation of California

Serpentine vegetation in California ranges from forest to shrubland and grassland, harbors many rare and endemic species, and is only moderately altered by invasive exotic species at the present time. To better understand the factors regulating the distribution of common/representative species, endemic/rare species, and the threat of exotics in this important flora, we analyzed broad-scale community patterns and environmental conditions in a geographically stratified set of samples from across the state. We considered three major classes of environmental influences: climate (especially precipitation), soils (especially the Mg2+/Ca2+ ratio), and the indirect influences of climate on soils. We used ordination to identify the major axes of variation in common species abundances, structural equation models to analyze the relationship of community axes and endemic and exotic species richness to the environment, and group analysis techniques to identify consistent groupings of species and characterize their properties. We found that community variation could be explained by a two-axis ordination. One axis ranged from conifer forest to grassland and was strongly related to precipitation. The second axis ranged from chaparral to grassland and had little relationship to current environmental conditions, suggesting a possible role for successional history. Precipitation and elevation were respectively the largest influences on endemic and exotic richness, followed by Mg 2+/Ca2+. The results also support the idea that long-term precipitation patterns have altered the Mg2+/Ca2+ ratio via selective leaching, resulting in indirect influences on endemics (positive) and exotics (negative) but not affecting the abundances of common species. We discuss implications of these findings for the conservation of the California serpentine flora. ?? 2007 Springer Science+Business Media B.V.

Conference Paper↗

A 21st century perspective on postfire seeding

Wildfires elicit a perceived need for emergency action to stabilize denuded landscapes. Aerial seeding of rapidly growing nonnative grasses is used routinely in an attempt to control postfire erosion, despite limited scientific basis for its effectiveness and with little consideration for its unintended ecological impacts. As fire size and magnitude have increased in recent decades, so has the prevalence and cost of postfire seeding and the potential footprint of its unintended impacts. We see a growing consensus in the research community on two important points: this management practice often is not cost-effective and it appears to create more problems than it solves.

Journal of Forestry↗

Invasion in a diversity hotspot: Exotic cover and native richness in the Californian serpentine flora

Exotic species have been observed to be more prevalent in sites where the richness of native species is highest, possibly reflecting variation among sites in resources, propagule supply, heterogeneity, or disturbance. However, such a pattern leaves unclear whether natives at species-rich sites are subject to especially severe impacts from exotics as a result. We considered this question using path models in which relationships between exotic cover and native richness were evaluated in the presence of correlated environmental factors. At 109 sites on serpentine soils across California, USA, exotic cover was positively correlated with total native herbaceous richness and was negatively correlated with the richness of both serpentine-endemic and rare native herbs. However, in path models that accounted for the influences of soil chemistry, disturbance, overstory cover, and regional rainfall and elevation, we found no indication that exotic cover reduced any component of native herb richness. Rather, our results indicated similarities and differences in the conditions favoring exotic, native, endemic, and rare species. Our results suggest that, in spite of some localized impacts, exotic species are not exerting a detectable overall effect on the community richness of the unique native flora of Californian serpentine. ?? 2006 by the Ecological Society of America.

California↗

Regional and local species richness in an insular environment: Serpentine plants in California

We asked how the richness of the specialized (endemic) flora of serpentine rock outcrops in California varies at both the regional and local scales. Our study had two goals: first, to test whether endemic richness is affected by spatial habitat structure (e.g., regional serpentine area, local serpentine outcrop area, regional and local measures of outcrop isolation), and second, to conduct this test in the context of a broader assessment of environmental influences (e.g., climate, soils, vegetation, disturbance) and historical influences (e.g., geologic age, geographic province) on local and regional species richness. We measured endemic and total richness and environmental variables in 109 serpentine sites (1000-m2 paired plots) in 78 serpentine-containing regions of the state. We used structural equation modeling (SEM) to simultaneously relate regional richness to regionalscale predictors, and local richness to both local-scale and regional-scale predictors. Our model for serpentine endemics explained 66% of the variation in local endemic richness based on local environment (vegetation, soils, rock cover) and on regional endemic richness. It explained 73% of the variation in regional endemic richness based on regional environment (climate and productivity), historical factors (geologic age and geographic province), and spatial structure (regional total area of serpentine, the only significant spatial variable in our analysis). We did not find a strong influence of spatial structure on species richness. However, we were able to distinguish local vs. regional influences on species richness to a novel extent, despite the existence of correlations between local and regional conditions. ?? 2006 by the Ecological Society of America.

California↗

Fire suppression impacts on postfire recovery of Sierra Nevada chaparral shrublands

A substantial portion of chaparral shrublands in the southern part of California's Sierra Nevada Mountain Range has never had a recorded fire since record keeping began in 1910. We hypothesised that such long periods without fire are outside the historical range of variability and that when such areas burn, postfire recovery is weaker than in younger stands. We predicted that long fire-free periods will result in loss of shrub species and deterioration of soil seed banks, which, coupled with higher fire intensities from the greater accumulation of dead biomass, will lead to poorer postfire regeneration. The 2002 McNally Fire burned ancient stands that were as much as 150 years old, as well as much younger (mature) stands. Based on shrub skeletons in the burned area as a surrogate for prefire density, we found that ancient stands change in structure, owing primarily to the loss of obligate seeding Ceanothus cuneatus; other species appear to have great longevity. Despite the reduction in C. cuneatus, postfire shrub-seedling recruitment remained strong in these ancient stands, although some seed bank deterioration is suggested by the three-quarters lower seedling recruitment than recorded from mature stands. Total diversity and the abundance of postfire endemic annuals are two other response variables that suggest that these ancient stands are recovering as well as mature stands. The one area of some concern is that non-native species richness and abundance increased in the ancient stands, suggesting that these are more open to alien colonisers. It is concluded that chaparral more than a century old is resilient to such long fire-free periods and fire severity impacts are indistinguishable from those in younger chaparral stands.

International Journal of Wildland Fire↗