Geology topics

Gregory W. Kennedy

Publications and source records attributed to Gregory W. Kennedy.

At least 19 recordsLinked to original sources

USGS2024

Experimental assessment of egg mat gear retention and collection efficacy

Assessment of egg deposition is widely used to provide an index of spawning efforts for lithophilic spawning fishes. However, little is known about the collection efficacy and bias of fish egg collection methods. We conducted a two-phased study consisting of a simulated-river flume study (two-part design), and a field study (an egg drift comparison with capture on mats) to assess egg collection methods and evaluate egg retention and capture on egg mats. Lake whitefish Coregonus clupeaformis egg retention on seeded mats decreased with increasing velocity and walleye Sander vitreus egg retention was variable as velocity increased. Fewer lake whitefish eggs were collected on egg mats when limestone reef rock was present in the flume study during the simulated spawned trials, but the inverse was true for walleye. Similarly, during field collections more lake whitefish eggs were collected in benthic D-shaped frame (D-frame) drift nets set near a known spawning reef compared to egg mats set on the reef, indicating lake whitefish eggs were drifting downstream along the river bottom. In contrast, fewer walleye eggs were observed in D-frame drift nets compared to number of eggs captured on the egg mats. Therefore, egg mats are an informative tool for evaluating walleye egg deposition in an immediate area but may underestimate egg deposition of lake whitefish, especially in lotic systems. Compared to other egg collection methods in the current literature, our study indicates that egg mats are useful for assessing egg deposition by lithophilic spawning fishes, but that the collection and retention efficacy and bias of this gear may vary between species and habitat types.

Journal of Fish and Wildlife Management↗

USGS2023

Repeat bathymetric surveys and model simulation of sedimentation processes near fish spawning placements, Detroit and St. Clair Rivers, Michigan

Nine rock-rubble fish spawning placements, or artificial reef complexes, constructed in the Detroit and St. Clair Rivers between 2004 to 2018 were surveyed periodically with multibeam sonar. These serial bathymetric surveys, conducted in 2015, 2018, 2021, and 2022, identified active sand bedform fields impinging two reef complexes: Fighting Island in the Detroit River and Middle Channel in the St. Clair River delta. The spatial extent over which the bedforms interacted with these reef complexes differed. The Fighting Island reef complex, which was comprised of twelve reef beds oriented across the river channel, experienced partial sedimentation that can be attributed to the streamwise translation and lateral encroachment of a bedform field on several of the eastern reef beds. The Middle Channel reef complex was comprised of nine reef beds also oriented across the river channel. Sedimentation of the Middle Channel reef complex was more comprehensive compared to the Fighting Island reef complex as most of the beds in the Middle Channel reef complex were within a translating bedform field. We simulated the temporal evolution of reef sedimentation at the Middle Channel reef complex using the Wilcock-Kenworthy (WK) two-fraction sediment transport model. In the WK simulation, sand available upstream of the reef migrated into the 36-meter-long gravel reef beds over 10 days of model simulation. The rate of sediment infill predicted by the model was more rapid than the speed of bedform slip face translation measured in the field, approximately 0.3 meters per day. Further, as the supply of sediment from upstream is continuous, once a reef bed fills with sediment it generally remains in place, although some small variations (+/- 0.2 m) in the elevation of the sand overlying the reef beds were observed. Taken together, bathymetric surveys and modeling could be used to identify, monitor, and simulate potential sources of bedload sediment that could impair the longevity of future spawning reef placements. Efforts directed toward enhancement and/or maintenance of reefs impaired by sedimentation could benefit from continued monitoring through periodic high-resolution bathymetric surveys, detailed inspection by diving, and collection of underwater imagery.

Michigan↗

USGS2021

A structured approach to remediation site assessment: Lessons from 15 years of fish spawning habitat creation in the St. Clair‐Detroit River system

Ideally, restoration re‐establishes natural processes in degraded habitats (e.g., flow and sediment regimes). However, in altered systems where process‐based restoration is not feasible, habitat construction is another approach to mitigate degradation. Because habitat construction does not directly focus on restoring processes that build and maintain desired habitats, projects must be developed and placed within the contemporary regulatory, ecological, and hydrogeomorphic context of a system, to maximize effectiveness. Here, we develop a framework for evaluating the regulatory, ecological, and hydrogeomorphic components using 15 years of fish spawning habitat construction in the St. Clair‐Detroit River System. The process began by identifying regulatory requirements at a coarse resolution to quickly focus on locations where ecological potential and hydrogeomorphic constraints could be assessed at finer resolutions. Next, ecological potential was assessed using a lithophilic fish spawning habitat suitability index. The suitability index identified five sites for habitat construction and Lake sturgeon spawning was documented at each site following construction. However, qualitative monitoring showed fine sediments accumulated at older sites. Thus, geomorphic assessments were incorporated to identify sediment sources and model flow within targeted areas. Since geomorphic assessments required the finest resolution and had the most uncertainty, they were conducted after broad‐scale regulatory considerations and ecological assessments narrowed focus to a few candidate sites. The order of operations identified in this case study evolved from the iterative approach of the restoration team, but in retrospect, it helped develop a framework that directed project development resources to aspects with more uncertainty, where learning is most critical.

Michigan, Ontario↗

USGS2018

Lake Sturgeon, Lake Whitefish, and Walleye egg deposition patterns with response to fish spawning substrate restoration in the St. Clair–Detroit River system

Egg deposition and use of restored spawning substrates by lithophilic fishes (e.g., Lake Sturgeon Acipenser fulvescens , Lake Whitefish Coregonus clupeaformis , and Walleye Sander vitreus ) were assessed throughout the St. Clair–Detroit River system from 2005 to 2016. Bayesian models were used to quantify egg abundance and presence/absence relative to site-specific variables (e.g., depth, velocity, and artificial spawning reef presence) and temperature to evaluate fish use of restored artificial spawning reefs and assess patterns in egg deposition. Lake Whitefish and Walleye egg abundance, probability of detection, and probability of occupancy were assessed with detection-adjusted methods; Lake Sturgeon egg abundance and probability of occurrence were assessed using delta-lognormal methods. The models indicated that the probability of Walleye eggs occupying a site increased with water velocity and that the rate of increase decreased with depth, whereas Lake Whitefish egg occupancy was not correlated with any of the attributes considered. Egg deposition by Lake Whitefish and Walleyes was greater at sites with high water velocities and was lower over artificial spawning reefs. Lake Sturgeon eggs were collected least frequently but were more likely to be collected over artificial spawning reefs and in greater abundances than elsewhere. Detection-adjusted egg abundances were not greater over artificial spawning reefs, indicating that these projects may not directly benefit spawning Walleyes and Lake Whitefish. However, 98% of the Lake Sturgeon eggs observed were collected over artificial spawning reefs, supporting the hypothesis that the reefs provided spawning sites for Lake Sturgeon and could mitigate historic losses of Lake Sturgeon spawning habitat.

Transactions of the American Fisheries Society↗

USGS2017

Egg deposition by lithophilic-spawning fishes in the Detroit and Saint Clair Rivers, 2005–14

A long-term, multiseason, fish egg sampling program conducted annually on the Detroit (2005–14) and Saint Clair (2010–14) Rivers was summarized to identify where productive fish spawning habitat currently exists. Egg mats were placed on the river bottom during the spring and fall at historic spawning areas and candidate fish spawning habitat restoration sites throughout both rivers. Widespread evidence was found of lithophilic spawning by numerous native fish species, including walleye ( Sander vitreus ), lake whitefish ( Coregonus clupeaformis ), lake sturgeon ( Acipenser fulvescens ), suckers (Catostomidae spp.), and trout-perch ( Percopsis omiscomaycus ). Walleye, lake whitefish, and suckers spp. spawned in nearly every region of each river in all years on both reef and nonreef substrates. Lake sturgeon eggs were collected almost exclusively over constructed reefs. Catch-per-unit effort of walleye, lake whitefish, and sucker eggs was much greater in the Detroit River than in the Saint Clair River, while Saint Clair River sites supported the greatest collections of lake sturgeon eggs. Collections during this study of lake sturgeon eggs on man-made spawning reefs suggest that artificial reefs may be an effective tool for restoring fish populations in the Detroit and Saint Clair Rivers; however, the quick response of lake sturgeon to spawn on newly constructed reefs and the fact that walleye, lake whitefish, and sucker eggs were often collected over substrate with little interstitial space to protect eggs from siltation and predators suggests that lack of suitable spawning habitat may continue to limit reproduction of lithophilic-spawning fish species in the Saint Clair-Detroit River System.

Michigan↗

USGS2015

Mudpuppy ( Necturus maculosus maculosus ) spatial distribution, breeding water depth, and use of artificial spawning habitat in the Detroit River

Mudpuppy ( Necturus maculosus maculosus ) populations have been declining in the Great Lakes region of North America. However, during fisheries assessments in the Detroit River, we documented Mudpuppy reproduction when we collected all life stages from egg through adult as by-catch in fisheries assessments. Ten years of fisheries sampling resulted in two occurrences of Mudpuppy egg collection and 411 Mudpuppies ranging in size from 37–392 mm Total Length, collected from water 3.5–15.1 m deep. Different types of fisheries gear collected specific life stages; spawning females used cement structures for egg deposition, larval Mudpuppies found refuge in eggmats, and we caught adults with baited setlines and minnow traps. Based on logistic regression models for setlines and minnow traps, there was a higher probability of catching adult Mudpuppies at lower temperatures and in shallower water with reduced clarity. In addition to documenting the presence of all life stages of this sensitive species in a deep and fast-flowing connecting channel, we were also able to show that standard fisheries research equipment can be used for Mudpuppy research in areas not typically sampled in herpetological studies. Our observations show that typical fisheries assessments and gear can play an important role in data collection for Mudpuppy population and spawning assessments.

Michigan, Ontario↗

USGS2014

A portable freshwater closed-system fish egg incubation system

To identify fish eggs collected in the field to species, a portable closed‐system fish egg incubation system was designed and used to incubate and hatch the eggs in the laboratory. The system is portable, small in scale (2.54 × 1.52 × 2.03 m), and affordable, with the approximate cost of the system being US$8,300 (2012). The main tank is 678 L and holds a battery of up to 21 (egg) incubation jars. The system includes three independent water pumping systems to (1) provide aerated water to hatching jars, (2) filter and sterilize incubation water, and (3) provide temperature‐controlled water in the main tank bath and the incubation jars. The system was successfully used to incubate freshwater fish eggs to raise resulting larvae to the post‐yolk‐sac stage for three seasons (spring 2012, spring 2013, and fall 2013) over two consecutive years, at two different locations, enabling us to identify fish eggs to species by providing identifiable fish larvae from incubated fish eggs.

North American Journal of Aquaculture↗

USGS2014

Habitat used by juvenile lake sturgeon ( Acipenser fulvescens ) in the North Channel of the St. Clair River (Michigan, USA)

Lake sturgeon ( Acipenser fulvescens ) occupy the St. Clair River, part of a channel connecting lakes Huron and Erie in the Laurentian Great Lakes. In the North Channel of the St. Clair River, juvenile lake sturgeon (3–7 years old and 582–793 mm in length) were studied to determine movement patterns and habitat usage. Fourteen juveniles were implanted with ultrasonic transmitters and tracked June–August of 2004, 2005 and 2006. Telemetry data, Geographic Information System software, side-scan sonar, video images of the river bottom, scuba diving, and benthic substrate samples were used to determine the extent and composition of habitats they occupied. Juvenile lake sturgeon habitat selection was strongly related to water depth. No fish were found in <6 m of water and over 97% of the relocations were found at depths greater than 9 m. Available water depths exceeding 18 m only represented 3.5% of the available habitat, however 34.9% of the relocations were found at depths exceeding 18 m. Juvenile lake sturgeon did not use most areas in proportion to their availability. Sturgeon avoided clay ledges and shallow areas with silt or soft clay, which comprised approximately 39% of the benthic habitat in the North Channel. A total of 300 out of 351 documented locations were on sand and gravel habitat types mixed with clay. Lake sturgeon > 700 mm in length selected sand and gravel areas mixed with zebra mussels and areas dominated by zebra mussels, while fish < 700 mm used these habitat types in proportion to their availability.

Michigan↗

USGS2012

Life history characteristics of a recovering lake whitefish Coregonus clupeaformis stock in the Detroit River, North America

The Detroit River is part of a channel connecting Lakes Huron and Erie and was once a prolific spawning area for lake whitefish, Coregonus clupeaformis . Large numbers of lake whitefish migrated into the river to spawn where they were harvested by commercial fisheries and for fish culture operations. Prior to our study, the last lake whitefish was landed from the Detroit River in 1925. Loss of spawning habitat during shipping channel construction and over-fishing, likely reduced lake whitefish spawning runs. Because lake whitefish are recovering in Lake Erie, and spawning in the western basin, we suspected they may also be spawning in the Detroit River. We sampled in the Detroit River for lake whitefish adults and eggs in October–December 2005–07 and for larvae during March–May 2006–08. A total of 15 spawning-ready lake whitefish from 4 to 18 years old, were collected. Viable eggs were collected during mid-November 2006–07; highest egg densities were found mid-river. Sac-fry whitefish larvae were collected in the river and near the river mouth. No whitefish larvae were retained in the river. Because high numbers of larvae were collected from mid- and downstream river sites, reproduction of lake whitefish in the Detroit River could contribute substantially to the Lake Erie lake whitefish metapopulation.

Michigan, Ontario↗

USGS2003

Starvation resistance in lake trout fry

Newly hatched fry were acclimated to 7 or 12°C and either fed daily (controls) or denied food for varying lengths of time and then fed daily until the end of the study (day 91 at 7°C and day 43 at 12°C). Growth was reduced by delays in the onset of feeding of 27 or more days at 7°C and 7 or more days at 12°C. Mortality of fry unfed for more than 34 days at 7°C, or more than 21 days at 12°C, was higher than among controls. Daily mortality increased with the length of the food deprivation period and did not cease immediately when food was made available, but reached zero by the end of the study. Mortality among unfed fry reached 50% in about 59 days at 7°C and 32 days at 12°C. Study results permitted calculation of the "point-of-no-return" (PNR) mortality, which included the mortality that occurred during the period of food deprivation, and also the delayed component of mortality that was directly attributable to starvation and that occurred after food was made available. The PNR for 50% mortality for food-deprived fry occurred after 52 days at 7°C and 24 days at 12°C. Thus, both measures of mortality indicate that lake trout fry would be highly resistant to death by starvation in the thermal habitat they would be expected to occupy in the Great Lakes. We conclude that a more likely adverse effect of reduced food availability would result from a reduction in growth rate that extends the length of time fry remain small and vulnerable to predation by adult alewives and other non-native fishes with which they associate.

Journal of Great Lakes Research↗

USGS1998

Zebra mussels invade Lake Erie muds

Zebra mussels (Dreissena polymorpha) originated in western Russia but have now become widespread in Europe and North America. They are widely known for their conspicuous invasion of rocks and other hard substrates in North American and European watersheds. We have found beds of zebra mussels directly colonizing sand and mud sediments each year across hundreds of square kilometres of North America's Lake Erie. This transformation of sedimentary habitats into mussel beds represents an unforeseen change in the invasive capacity of this species.

Nature↗

USGS1997

Use of remote-sensing techniques to survey the physical habitat of large rivers

Remote-sensing techniques that can be used to quantitatively characterize the physical habitat in large rivers in the United States where traditional survey approaches typically used in small- and medium-sized streams and rivers would be ineffective or impossible to apply. The state-of-the-art remote-sensing technologies that we discuss here include side-scan sonar, RoxAnn, acoustic Doppler current profiler, remotely operated vehicles and camera systems, global positioning systems, and laser level survey systems. The use of these technologies will permit the collection of information needed to create computer visualizations and hard copy maps and generate quantitative databases that can be used in real-time mode in the field to characterize the physical habitat at a study location of interest and to guide the distribution of sampling effort needed to address other habitat-related study objectives. This report augments habitat sampling and characterization guidance provided by Meador et al. (1993) and is intended for use primarily by U.S. Geological Survey National Water Quality Assessment program managers and scientists who are documenting water quality in streams and rivers of the United States.

Report↗

USGS1996

Potential spawning habitat for lake trout on Julian's Reef, Lake Michigan

Julian's Reef is an historical spawning ground for lake trout ( Salvelinus namaycush ) in southwestern Lake Michigan. It is a designated lake trout refuge and is the focus of lake trout restoration efforts in Illinois waters of the lake. We studied the reef to determine its potential as spawning habitat for stocked lake trout. We used side-scan sonar and a remotely operated vehicle equipped with a video camera to survey and map 156 ha of lake bed on the southeast portion of the reef, where an earlier study revealed the presence of loose-rock substrate potentially suitable for use by spawning lake trout. Our survey showed that the substrate on the reef that most closely resembled that described in the literature as suitable for spawning by stocked lake trout in the Great Lakes was rubble patches with interstitial depths greater than 20 cm. These rubble patches occupied about 2 ha of the 13-ha expanse of bedrock and rubble substrate near the reef crest in the surveyed area. We estimated that these rubble patches, if fully used by spawning lake trout, could accommodate egg deposition by at least 1,300–3,300 2.7-kg females.

Journal of Great Lakes Research↗

USGS1995

Availability of lake trout reproductive habitat in the Great Lakes

A decades-long program to reestablish self-sustaining stocks of lake trout ( Salvelinus namaycush ) in the four lower Great Lakes produced excellent fisheries supported by stocked fish. These fish spawned widely and small numbers of their offspring were collected intermittently from Lakes Michigan, Huron, and Ontario, but no self-sustaining stocks were established. Irt this paper we address habitat sufficiency as a factor in the failure of stocked lake trout to established self-sustaining populations in the four lower Great Lakes. We present the previously unpublished results of lake trout spawning habitat surveys conducted at seven sites in the Great Lakes since 1987 and we compare them with the published results of similar surveys conducted at 24 other sites in the four lower lakes since 1981. Our evaluation indicates all but two of these sites can support the production of viable fry from spawnings by the shallow-water strains of lake trout that are stocked in the Great Lakes. However, some of the best spawning, egg, and fry habitat in the lower Great Lakes seems to be at deeper offshore sites that may be unattractive to these shallow-water strains. Thus, we suggest also stocking the lower four lakes with strains from Lake Superior that might more fully exploit the best spawning habitat at these deeper, offshore sites.

Journal of Great Lakes Research↗

USGS1995

Survival of lake trout eggs on reputed spawning grounds in Lakes Huron and Superior: In situ incubation, 1987-1988

Lake trout reproduce widely in Lake Superior but little in Lake Huron. We examined whether survival of lake trout eggs and fry in either lake was reduced by physical disturbances and swim-up mortality. Eggs were collected from feral lake trout in Lake Superior and placed in 108 plastic incubators. A total of 48 incubators was set at Partridge Island Reef in southern Lake Superior, 48 were set at Port Austin Reef in southern Lake Huron, and 12 were held as controls inflowing well water at a laboratory. Survival-to-hatching of these eggs at Partridge Island Reef (18%) was significantly different from that at Port Austin Reef (43%) and significantly different in the laboratory (88%) from that at either reef (P < 0.05). During egg-fry incubation from 28 October 1987 to 5 May 1988, 11–18 cm of sediment accumulated in sediment traps placed on the reefs but < 1 cm of sediment was present on each reef in May 1988. Analysis showed that 44% of the eggs at Port Austin Reef and 28% of those at Partridge Island Reef were buried and killed by sediments. During the first week after deployment, mean wave energy was 90% higher at Partridge Island Reef and significantly different from that at Port Austin Reef. Wave energy may be a habitat condition that makes Partridge Island Reef less suitable than Port Austin Reef for incubation of lake trout eggs. Fry from eggs incubated at all three sites experienced no swim-up mortality. We conclude that in 1987–88 habitat conditions required for survival of lake trout eggs were more suitable at Port Austin Reef than at Partridge Island Reef.

Journal of Great Lakes Research↗

USGS1995

An evaluation of lake trout reproductive habitat on Clay Banks Reef, northwestern Lake Michigan

The extinction of the native populations of lake trout ( Salvelinus namaycush ) in Lake Michigan in about 1956 has been followed by a decades-long attempt to reestablish self-sustaining populations of this valuable species in habitats it formerly occupied throughout the lake. One of the most recent management strategies designed to facilitate recovery was to make a primary management objective the establishment of sanctuaries where stocked lake trout could be protected and self-sustaining populations reestablished. In the present study we employed habitat survey and mapping techniques, field and laboratory bioassays, egg traps, sediment traps, and gill nets to examine the potential for successful natural reproduction by stocked lake trout on Clay Banks Reef in the Door-Kewaunee sanctuary in Wisconsin waters of Lake Michigan. Our study revealed (1) there was suitable habitat on the reef to support the production of viable fry, (2) spawner abundance on the reef was the highest recorded in the great lakes, and (3) eggs taken from spawners on the reef and held on the reef in plexiglas incubators hatched and produced fry that survived through swim-up. We conclude that Clay Banks Reef has the potential to support successful natural reproduction by stocked lake trout.

Journal of Great Lakes Research↗

USGS1993

Distribution, abundance, and resting microhabitat of burbot on Julian's Reef, southwestern Lake Michigan

We used a remotely operated submersible vehicle equipped with a color video camera to videotape the lake bed and document the distribution and abundance of burbot Lota lota on a 156‐hectare portion of Julianˈs Reef in southwestern Lake Michigan. The substrates and bathymetry of the study area had been mapped recently by side‐scan sonar. Burbot density determined from videotapes covering 6,900 m 2 of lake bed at depths of 23–41 m averaged 139 individuals/ hectare (range, 0–571/hectare). This density was substantially higher than the highest burbot density (59–95/hectare) reported in the literature. Burbot were present on the lake bed at depths of 23–36 m, but were most abundant near the crest of the reef at 23–28 m, where the water temperature was 8–13°C, their preferred summer temperature range. Substrates in that temperature range on the reef were bedrock, bedrock ridges, and bedrock and rubble. Burbot were most abundant on the bedrock and rubble. Small fish and macroinvertebrates typically eaten by burbot elsewhere in western Lake Michigan were distributed on the reef according to their summer preferred temperatures and were not seen in abundance where burbot density was highest. We saw no lake trout Salvelinus namaycush on Julianˈs Reef, although large numbers of juvenile lake trout have been stocked there annually and temperatures on the reef were in the preferred summer temperature range for lake trout.

Michigan, Wisconsin↗

USGS1992

Lake trout spawning habitat in the Six Fathom Bank-Yankee Reef lake trout sanctuary, Lake Huron

Attempts to reestablish self-sustaining stocks of lake trout ( Salvelinus namaycush ) in the lower four Great Lakes, where the species was extinguished in the 1950s and 1960s, have been largely unsuccessful. To avoid many of the problems believed to be contributing to this failure, the fishery management community recently established several sanctuaries in the offshore waters of the Great Lakes where the development and protection of self-sustaining stocks of lake trout would be a primary management objective. One of these, the Six Fathom Bank-Yankee Reef sanctuary, was created in the south-central portion of Lake Huron. This sanctuary covers 168,000 ha and includes the shallower portions of the Six Fathom and Ipperwash scarps, which are major bathymetric features in the southern half of the lake. Historical accounts describe Six Fathom Bank as the most important lake trout spawning ground in the lake. Here we present the results of lake bed surveys conducted in the sanctuary with side-scan sonar, underwater videocamera systems, and a small research submarine. Our observations of the lake bed are consistent with what is known of the bedrock stratigraphy, glacial history, and karst geomorphology of the Lake Huron basin. Most of the loose rock we found seemed to be derived from local carbonate bedrock formations, although non-carbonate rock probably from Precambrian sources to the north was also present in some areas. Much of the bedrock and loose rock displayed karst solution features described for the Bruce Peninsula on the Ontario shoreline. Our surveys revealed substantial areas of lake bed at water depths of 20–36 m that resembled suitable spawning and fry production habitat for the shallow-water strains of lake trout that are the focus of the rehabilitation effort. Low mid-lake nutrient levels documented recently by others and the extremely high abundance of Mysis relicta (an important item in the diet of young lake trout) that we documented on Yankee Reef also contributed to our evaluation of the sanctuary as a site with high potential to support a self-sustaining population of lake trout.

Journal of Great Lakes Research↗