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G.F. Sepik

Publications and source records attributed to G.F. Sepik.

At least 19 recordsLinked to original sources

Survival of female American Woodcock breeding in Maine

During 1986-1989, 89 female American woodcock (Scolopax minor) included in this study were radio-marked and survival estimated for the period 1 Aprii-15 June. Eleven woodcock died: five (45%) were killed by mammals, two (18%) by unknown predators, and one (9%) by a raptor; two (18%) died from entanglement in the transmitter harness; and 1 (9%) collided with a vehicle. Survival varied among years from 0.700 (1986) to 0.900 (1989) with a 4-year mean (95% CI) of 0.826. Survival did not differ between age classes (P = 0.900), or among years (P > 0.14), except for higher (P = 0.025) survival (0.875) in 1987 than in 1988 (0.735). A composite survival estimate--based on telemetry studies for the breeding, post-breeding, and winter periods-- was 0.363 for immatures and 0.474 for adults. Mean weights were not different between second year and after second year age classes (P = 0.167), but weight was related to woodcock capture date (P = 0.001). Survival for female woodcock was not related to mean snow depth or to mean, minimum temperature in winter or spring. Habitat use was different between females that died and those that lived, but sample size was small.

Book chapter

Assessing habitat selection in Spring by male American Woodcock in Maine with a geographic information system

Geographic information system (GIS) technology was used to identify habitats available to and used by male American woodcock (Scolopax minor) equipped with radio transmitters--54 in 1987, 51 in 1988, 46 in 1989 at Moosehorn National Wildlife Refuge, Maine. Woodcock were monitored from time of capture (25 March-15 April) to 15 June each year. To determine habitat selection by male woodcock, the following habitat characteristics were measured: land cover, age and stocking density of the forest overstory, soil drainage and texture, aspect, and percent slope. Habitat selection was examined as affected by the covariates weather and age-class of woodcock, and among years for diurnal and crepuscular periods of the breeding period. Multivariate techniques that compare use and availability of habitats were not available, so a statistical model was developed to rate importance of multiple habitat characteristics selected by woodcock. The most critical period for woodcock in terms of survival was from arrival to: mid-April. Second-year and after-second-year woodcock did not select different (P > 0.05) habitat types, but they did select different types among years and within breeding intervals (P < 0.05). In years when weather was moderate, woodcock selected young, dense stands of speckled alder (Alnus rugosa) and hardwoods, interspersed with forest openings. Suitable habitat can be maintained by creating an uneven-aged forest managed in even-aged blocks composed of several hardwood species. Managers can now quantify suitable woodcock habitat in a GIS and plan large-scale forest-harvesting strategies using data on several habitat characteristics (e.g., land cover, stand age, stocking density, soil drainage and texture, and aspect).

Book chapter

Survival of breeding male American woodcock in Maine

The authors investigated the survival of breeding male American woodcock in Washington County, Maine. A total of 150 birds were radio-arked during 1987-1989 and relationships were evaluated between survival, body mass, predation, weather, and habitat use. Survival varied from 0.690 (1989) to 0.924 (1988), with a 3-year mean (95% confidence interval) of 0.789 (0.693-0.885)

Maine

American woodcock

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Book chapter

Critical review of the current knowledge of the biology of the American woodcock and its management on the breeding grounds

We critiqued previous work on the biology and management of the American woodcock (Scolopax minor) on the breeding grounds. We determined that little is known about habitat variables and weather extremes that may limit the population. Most investigators who attempted to define habitat requirements of the woodcock used inadequate sample sizes, limited the duration of their studies, did not account for effects of weather, or failed to adequately measure habitat variables. Furthermore, the effects of hunting on local or regional populations has never been adequately studied. We concluded that obtaining data to understand the biology of the woodcock and the effects of hunting is essential before managers can reverse the long-term decline of woodcock numbers.

Biological Report

Techniques for research into woodcocks: experiences and recommendations

We describe methods for the study of the American woodcock (Scolopax minor) at the Moosehorn National Wildlife Refuge. Described are techniques for the capture of particular age-sex cohorts during different times of the year, the aging and sexing, the evaluation of habitat components, and the estimation of population size. We describe successful and tried but rejected techniques with radio-telemetry for the study of survival and habitat use.

Biological Report

Behavior of radio-marked breeding American woodcocks

During spring 1986-89, we equipped 175 male and 89 female American woodcocks (Scolopax minor) with radio transmitters. Radio-marking had little effect on behavior; within 1 day of marking, 37 of 64 (58%) displaying males were dominant and within 7 days, 138 of 157 (88%) were dominant. All females marked before nesting proceeded to nest, and marked females with broods remained with broods after release. Dominance of males declined from 73% in April to 69% in May and to 26% in June as breeding activity waned. In all years, after-second-year (ASY) males were dominant more often than second-year (SY) males (67.5% vs. 58.9%). ;Most males displayed at more than one (range = 2-12) site. Distances that males moved between the primary singing ground and subsequent singing grounds averaged 775 m in all years and ranged from 618 m (1986) to 966 m (1988). Females visited males at singing grounds throughout the breeding cycle; some females visited more than one site. During prenesting, females remained in daytime covers during the crepuscular period (55%), flew to different feeding covers (22%), visited singing grounds (14%), or flew from daytime covers to unknown locations (9%). During nesting, females left nests during the crepuscular period (72%) and moved to singing grounds (5%), to feeding areas (59%), and to undetermined locations (7%). Females with broods remained with their broods during the crepuscular period (62%), flew to feeding areas (30%), visited singing grounds (1%), or flew to undetermined locations (6%). The woodcock mating system is similar to a resource-based polygyny. Males compete for singing grounds near high-quality nesting areas. The fitness of males is expressed by dominance at more than one singing ground. Woodcocks are not monogamous; females may visit more than three different males during a single courtship period but do not visit males regularly. Most females leave nests during the crepuscular period to feed in a different cover.

Biological Report

Renesting by American woodcocks ( Scolopax minor ) in Maine

The American Woodcock ( Scolopax minor ) is one of the earliest ground-nesting birds in the northeastern United States. In Maine, nesting begins in early April when temperatures can drop below freezing and significant snowfall can accumulate. Nests are usually in open woods, where eggs are laid on the ground in a shallow depression (Pettingill 1936, Mendall and Aldous 1943, Sheldon 1967). Peak hatching occurs in early May (Dwyer et al. 1982), when temperatures are cool and precipitation is common. Woodcock chicks are dependent on the female for most of their food for at least seven days after hatching (Gregg 1984). During cool, wet weather, chicks require constant brooding by females; prolonged periods of inclement weather may lead to substantial mortality of chicks (Dwyer et al. 1988).

The Auk

An improved method to monitor nest attentiveness using radio-telemetry

An improved method of automatically monitoring nest attentiveness was designed and tested using radio-equipped American Woodcock (Scolopax minor). Shielded coaxial cable (RG-58) was extended from a receiver and placed 30 cm above the nest, with a 3.8 cm section of the inner wire exposed. Presence, absence, and activity of birds within 10.1 ? 5.2 m (SD) of the nest were clearly indicated on a Rustrak recorder while extraneous signal interference was minimized.

Journal of Field Ornithology

Demographic Characteristics of a Maine Woodcock Population and Effects of Habitat Management

A population of American woodcock (Scolopax minor) was studied on a 3,401-ha area of the Moosehorn National Wildlife Refuge in northeastern Maine from 1976 through 1985. During 1976-83, from 4 to 64 clearcuts were created each year, opening up large contiguous blocks of forest. A combination of mist nets, ground traps, nightlighting techniques, and trained dogs were used to capture and band 1,884 birds during the first 5 years. Capture and recapture data (totaling 3,009 observations) were used with both demographically closed and open population models to estimate population size and, for open population models, summer survival. Flying young, especially young males, represented the greatest proportion of all captures; analysis showed that young males were more prone to capture than young females. Male courtship began about 24 March each year, usually when there was still snow in wooded areas. Males ~2 years old dominated singing grounds during April each year, but this situation changed and first-year males dominated singing grounds in May. Singing males shifted from older established singing grounds to new clearcuts soon after we initiated forest management. Many males were subdominant at singing grounds despite an abundance of unoccupied openings. Three hundred adult females were captured and, except for 1978, the majority were ~2 years old. The year in which female homing rate was lowest(1979) was preceded by the year with the largest number of l-year-old brood female captures and a summer drought. Summer survival of young was lowest in 1978 and was attributed to summer drought. The year 1979 had an abnormally cool and wet spring, and was the poorest for production of young. Capture ratios of young-to-adult females obtained by nightlighting could be used to predict production on our study area. Closed population model estimates did not seem to fit either young or adult data sets well. Instead, a partially open capture-recapture model that allowed death but no immigration seemed to fit best. Only the number of males in the population changed significantly during the study. An increase from 88 males in 1976 to 156 in 1980 was attributed to habitat management. Singingmale surveys on our area detected little change in the number of singing males, but our independent population estimates from mark-recapture data showed a larger total male population by 1980. Annual density estimates for all age and sex classes ranged from 19 to 25 birds/l00 ha. A hypothesis on the breeding system of the American woodcock is presented as well as a discussion of management implications, including the importance of creating high-quality habitat on private lands.

Fish and Wildlife Research